| # | Institution | Articles | Citations | R ARank | R CRank |
|---|
| 1 | National Center for Scientific Research | 27 | 15,025 | 2.0K | 112 |
| 2 | French National Institute of Health and Medical Research | 9 | 13,222 | 1.0K | 22 |
| 3 | University of California, Santa Cruz | 4 | 12,970 | 367 | 7 |
| 4 | University College Dublin | 2 | 12,967 | 2.9K | 7 |
| 5 | University of California, San Diego | 41 | 8,523 | 372 | 105 |
| 6 | ETH Zürich | 61 | 8,416 | 198 | 66 |
| 7 | University of California | 64 | 7,676 | 2.3K | 983 |
| 8 | Keio University | 7 | 7,408 | 1.2K | 10 |
| 9 | Tel Aviv University | 25 | 3,787 | 807 | 145 |
| 10 | Weizmann Institute of Science | 33 | 3,427 | 154 | 111 |
| 11 | Princeton University | 21 | 3,254 | 660 | 293 |
| 12 | University of Toronto | 38 | 3,104 | 1.4K | 718 |
| 13 | University of Edinburgh | 21 | 2,895 | 1.1K | 317 |
| 14 | Uppsala University | 15 | 2,698 | 1.2K | 280 |
| 15 | European Bioinformatics Institute | 15 | 2,542 | 26 | 17 |
| 16 | Technical University of Munich | 10 | 2,417 | 1.6K | 259 |
| 17 | Massachusetts Institute of Technology | 34 | 2,390 | 648 | 589 |
| 18 | Helmholtz Zentrum München | 5 | 2,358 | 335 | 32 |
| 19 | Stanford University | 19 | 2,225 | 1.6K | 836 |
| 20 | Heidelberg University | 31 | 2,124 | 546 | 351 |
| 21 | Howard Hughes Medical Institute | 18 | 2,090 | 241 | 207 |
| 22 | University of Basel | 16 | 2,053 | 481 | 167 |
| 23 | University Hospital Heidelberg | 10 | 2,040 | 267 | 41 |
| 24 | University of California, San Francisco | 17 | 2,033 | 778 | 360 |
| 25 | California Institute of Technology | 12 | 2,005 | 835 | 355 |
| 26 | Chalmers University of Technology | 14 | 1,954 | 531 | 128 |
| 27 | KAIST | 4 | 1,897 | 1.4K | 148 |
| 28 | Imperial College London | 18 | 1,822 | 1.6K | 753 |
| 29 | Humboldt University of Berlin | 21 | 1,806 | 400 | 166 |
| 30 | Harvard University | 20 | 1,718 | 1.1K | 772 |
| 31 | University of Cambridge | 21 | 1,630 | 1.6K | 913 |
| 32 | University of California, Los Angeles | 16 | 1,524 | 830 | 477 |
| 33 | University of Groningen | 12 | 1,454 | 1.4K | 529 |
| 34 | Wellcome Genome Campus | 5 | 1,438 | 157 | 60 |
| 35 | Northwestern University | 4 | 1,428 | 3.8K | 755 |
| 36 | Newcastle University | 4 | 1,359 | 2.6K | 331 |
| 37 | University of Gothenburg | 10 | 1,357 | 1.3K | 444 |
| 38 | Memorial Sloan Kettering Cancer Center | 13 | 1,331 | 646 | 347 |
| 39 | University of Washington | 12 | 1,330 | 3.1K | 1.4K |
| 40 | (former) Institut National de Recherche pour l'Agriculture, l'Alimentation et l'Environnement | 7 | 1,315 | 885 | 307 |
| 41 | University of Virginia | 6 | 1,301 | 2.4K | 510 |
| 42 | Columbia University | 14 | 1,288 | 2.1K | 1.1K |
| 43 | University of Copenhagen | 13 | 1,247 | 1.8K | 809 |
| 44 | SRI International | 1 | 1,234 | 968 | 61 |
| 45 | University of Texas at Austin | 9 | 1,228 | 3.4K | 1.1K |
| 46 | Yale University | 12 | 1,189 | 1.5K | 813 |
| 47 | Utrecht University | 10 | 1,171 | 1.7K | 725 |
| 48 | Chinese Academy of Sciences | 10 | 1,126 | 3.6K | 1.5K |
| 49 | University of Ottawa | 3 | 1,110 | 3.8K | 405 |
| 50 | Duke University | 14 | 1,086 | 1.1K | 686 |
| 51 | University of Michigan | 8 | 1,079 | 4.3K | 1.7K |
| 52 | University of Zurich | 16 | 1,071 | 943 | 640 |
| 53 | Tokyo Institute of Technology | 2 | 992 | 2.7K | 397 |
| 54 | Assistance Publique – Hôpitaux de Paris | 1 | 991 | 1.7K | 180 |
| 55 | European Molecular Biology Laboratory | 11 | 918 | 28 | 37 |
| 56 | Ludwig Maximilian University of Munich | 10 | 917 | 1.6K | 743 |
| 57 | Delft University of Technology | 5 | 899 | 1.7K | 469 |
| 58 | Lunenfeld-Tanenbaum Research Institute | 9 | 898 | 102 | 87 |
| 59 | Weill Cornell Medicine | 4 | 871 | 1.8K | 502 |
| 60 | University of Warwick | 6 | 839 | 1.8K | 470 |
| 61 | Rice University | 9 | 833 | 682 | 362 |
| 62 | Université de Montréal | 7 | 828 | 2.1K | 676 |
| 62 | University of Wisconsin–Madison | 6 | 828 | 4.7K | 1.8K |
| 64 | University of California, Berkeley | 7 | 821 | 1.4K | 705 |
| 65 | Tsinghua University | 2 | 809 | 4.1K | 819 |
| 66 | Boston University | 6 | 775 | 1.9K | 690 |
| 67 | University of Virginia Health System | 1 | 759 | 1.1K | 125 |
| 68 | Charité | 9 | 756 | 490 | 201 |
| 69 | University of Tokyo | 6 | 753 | 4.4K | 1.5K |
| 70 | Institució Catalana de Recerca i Estudis Avançats | 8 | 725 | 34 | 31 |
| 71 | AlbaNova | 3 | 716 | 118 | 29 |
| 72 | Purdue University | 2 | 709 | 5.7K | 1.2K |
| 73 | Vrije Universiteit Amsterdam | 5 | 688 | 1.9K | 694 |
| 74 | Lawrence Berkeley National Laboratory | 4 | 661 | 839 | 408 |
| 75 | Technical University of Denmark | 8 | 657 | 1.1K | 604 |
| 76 | University of Freiburg | 8 | 656 | 1.1K | 549 |
| 77 | Wyss Institute for Biologically Inspired Engineering | 9 | 654 | 22 | 33 |
| 78 | National Institutes of Health | 13 | 648 | 1.6K | 1.4K |
| 79 | McGill University | 4 | 644 | 4.9K | 1.5K |
| 80 | University of Leeds | 2 | 642 | 5.2K | 943 |
| 81 | National Research Council Canada | 3 | 640 | 1.0K | 278 |
| 82 | University of Oxford | 11 | 639 | 2.8K | 1.9K |
| 83 | University of Liverpool | 3 | 635 | 3.7K | 808 |
| 84 | Peking University | 7 | 627 | 2.2K | 826 |
| 85 | Cancer Research UK | 3 | 618 | 287 | 134 |
| 86 | Centre for Genomic Regulation | 4 | 608 | 5 | 5 |
| 87 | Pacific Northwest National Laboratory | 4 | 600 | 630 | 277 |
| 88 | Institut National de Recherche en Informatique et en Automatique | 2 | 596 | 537 | 81 |
| 89 | University of British Columbia | 4 | 595 | 5.7K | 2.0K |
| 90 | Université catholique de Louvain | 2 | 590 | 3.2K | 671 |
| 91 | Université de Lyon | 3 | 587 | 1.2K | 205 |
| 92 | University Health Network | 6 | 575 | 860 | 376 |
| 92 | École Normale Supérieure de Lyon | 3 | 575 | 500 | 138 |
| 94 | University of Helsinki | 8 | 571 | 2.1K | 1.1K |
| 95 | Ontario Cancer Institute | 3 | 563 | 188 | 89 |
| 96 | Claude Bernard University Lyon 1 | 2 | 562 | 1.9K | 304 |
| 97 | University of Cologne | 9 | 558 | 1.2K | 685 |
| 98 | Ghent University | 6 | 557 | 2.7K | 1.1K |
| 99 | Virginia Tech | 7 | 550 | 1.9K | 869 |
| 100 | New York University | 8 | 541 | 1.6K | 917 |
| 101 | Cornell University | 7 | 538 | 2.9K | 1.8K |
| 102 | The Rockefeller University | 5 | 535 | 552 | 384 |
| 103 | Hebrew University of Jerusalem | 4 | 532 | 2.5K | 795 |
| 104 | University of Lausanne | 11 | 529 | 571 | 546 |
| 105 | University of Tübingen | 6 | 526 | 1.8K | 818 |
| 105 | Johns Hopkins University | 7 | 526 | 2.8K | 1.6K |
| 105 | Greehey Children's Cancer Research Institute | 1 | 526 | 155 | 22 |
| 108 | Pennsylvania State University | 4 | 521 | 5.2K | 1.8K |
| 109 | John Innes Centre | 1 | 520 | 438 | 142 |
| 110 | University Medical Center Utrecht | 7 | 513 | 1.0K | 631 |
| 111 | National University of Singapore | 5 | 504 | 3.6K | 1.4K |
| 112 | Sahlgrenska University Hospital | 4 | 498 | 1.1K | 418 |
| 113 | German Cancer Research Center | 6 | 492 | 282 | 201 |
| 114 | University of Notre Dame | 1 | 489 | 3.9K | 542 |
| 114 | University of Notre Dame Australia | 1 | 489 | 1.4K | 146 |
| 116 | Technical University of Berlin | 6 | 480 | 1.1K | 445 |
| 117 | Technion – Israel Institute of Technology | 5 | 477 | 1.6K | 567 |
| 118 | Royal Institute of Technology | 3 | 468 | 1.8K | 495 |
| 118 | University of Southern Denmark | 1 | 468 | 3.2K | 460 |
| 120 | Novo Nordisk Foundation Center for Protein Research | 2 | 449 | 17 | 11 |
| 121 | Simon Fraser University | 2 | 443 | 3.1K | 616 |
| 122 | University of Texas MD Anderson Cancer Center | 5 | 441 | 1.1K | 611 |
| 123 | Joint BioEnergy Institute | 1 | 440 | 88 | 28 |
| 124 | National Institute of Allergy and Infectious Diseases | 4 | 425 | 564 | 338 |
| 125 | Baylor College of Medicine | 3 | 418 | 2.8K | 1.1K |
| 126 | Biozentrum University of Basel | 6 | 412 | 20 | 32 |
| 127 | Pompeu Fabra University | 6 | 409 | 314 | 198 |
| 127 | Hyogo College of Medicine | 1 | 409 | 1.2K | 130 |
| 127 | National Institute for Basic Biology, Japan | 1 | 409 | 352 | 97 |
| 130 | Vanderbilt University Medical Center | 3 | 405 | 1.6K | 556 |
| 131 | Whitehead Institute | 3 | 404 | 151 | 133 |
| 131 | Max Planck Institute of Biochemistry | 3 | 404 | 52 | 36 |
| 131 | Università degli Studi di Milano Bicocca | 1 | 404 | 2.0K | 182 |
| 134 | University of Pittsburgh | 4 | 399 | 4.5K | 2.0K |
| 135 | Moscow State University | 2 | 393 | 2.8K | 554 |
| 135 | University of California, Santa Barbara | 2 | 393 | 1.5K | 462 |
| 137 | Thomas Jefferson University | 3 | 390 | 1.8K | 640 |
| 138 | Carnegie Mellon University | 4 | 383 | 1.9K | 1.0K |
| 138 | Karolinska Institute | 8 | 383 | 1.9K | 1.5K |
| 140 | University of Texas Southwestern Medical Center | 6 | 377 | 1.5K | 945 |
| 141 | Nanyang Technological University | 3 | 376 | 2.8K | 1.0K |
| 142 | École polytechnique fédérale de Lausanne | 6 | 374 | 861 | 602 |
| 142 | University of Waterloo | 1 | 374 | 5.6K | 1.2K |
| 144 | Wellcome Sanger Institute | 4 | 367 | 206 | 161 |
| 145 | National Heart, Lung, and Blood Institute | 2 | 364 | 820 | 329 |
| 146 | University of Luxembourg | 6 | 361 | 202 | 123 |
| 147 | Oak Ridge National Laboratory | 3 | 353 | 1.4K | 703 |
| 148 | University of Naples Federico II | 2 | 349 | 3.7K | 1.0K |
| 149 | Laval University | 6 | 345 | 1.8K | 1.1K |
| 150 | Pasteur Institute | 5 | 342 | 742 | 522 |
| 151 | Hospices Civils de Lyon | 1 | 340 | 838 | 140 |
| 152 | King's College London | 4 | 335 | 2.9K | 1.4K |
| 153 | Genome Institute of Singapore | 4 | 326 | 83 | 67 |
| 154 | Sage Bionetworks | 3 | 325 | 28 | 27 |
| 155 | Michigan Medicine | 2 | 322 | 2.3K | 879 |
| 156 | University College London | 4 | 319 | 4.7K | 2.4K |
| 157 | Duke University Hospital | 4 | 318 | 2.1K | 1.3K |
| 158 | North Carolina State University | 4 | 312 | 2.9K | 1.5K |
| 159 | Swedish University of Agricultural Sciences | 2 | 295 | 1.4K | 615 |
| 159 | Aix-Marseille University | 3 | 295 | 1.6K | 517 |
| 161 | University of New South Wales | 2 | 293 | 6.4K | 2.2K |
| 162 | University of Chicago | 4 | 288 | 3.6K | 2.1K |
| 162 | University of Minnesota | 6 | 288 | 4.7K | 3.2K |
| 164 | ComUE Paris-Saclay University | 3 | 284 | 1.6K | 469 |
| 165 | University of Paris-Sud | 1 | 277 | 2.1K | 562 |
| 166 | Leiden University | 4 | 274 | 2.0K | 1.1K |
| 166 | University of Birmingham | 5 | 274 | 3.3K | 2.0K |
| 168 | Radboud University Nijmegen | 3 | 272 | 2.5K | 1.2K |
| 168 | Ottawa Hospital Research Institute | 1 | 272 | 934 | 274 |
| 170 | University of Colorado | 4 | 270 | 3.0K | 1.8K |
| 171 | University of Iceland | 1 | 264 | 1.8K | 340 |
| 172 | PSL University | 3 | 263 | 635 | 233 |
| 173 | University of Toulouse | 5 | 262 | 1.0K | 595 |
| 173 | Northeastern University | 2 | 262 | 3.6K | 1.2K |
| 175 | WEHI | 3 | 261 | 245 | 179 |
| 176 | Okinawa Institute of Science and Technology | 2 | 257 | 227 | 51 |
| 177 | University of Rochester Medical Center | 2 | 254 | 1.5K | 632 |
| 178 | University Hospital of Basel | 3 | 253 | 797 | 396 |
| 179 | University of California, Davis | 4 | 246 | 1.9K | 1.3K |
| 180 | University of Veterinary Medicine Vienna | 4 | 243 | 192 | 114 |
| 181 | University of Utah | 3 | 235 | 4.1K | 2.1K |
| 182 | Oxford Brookes University | 1 | 232 | 1.6K | 241 |
| 182 | University of Delaware | 2 | 232 | 3.1K | 1.2K |
| 184 | University of Nebraska–Lincoln | 2 | 231 | 2.3K | 824 |
| 185 | University of Oulu | 1 | 230 | 3.4K | 935 |
| 185 | Finnish Institute for Health and Welfare | 1 | 230 | 735 | 201 |
| 187 | University of Graz | 2 | 229 | 1.7K | 636 |
| 188 | Rutgers University | 3 | 228 | 4.5K | 2.3K |
| 189 | Seoul National University | 2 | 222 | 4.6K | 1.9K |
| 189 | University of Vienna | 3 | 222 | 3.3K | 1.7K |
| 191 | Janssen Pharmaceuticals | 1 | 220 | 375 | 94 |
| 192 | Vanderbilt University | 2 | 216 | 3.9K | 1.7K |
| 193 | Australian Antarctic Division | 1 | 215 | 177 | 61 |
| 193 | CSIRO Oceans and Atmosphere | 1 | 215 | 281 | 126 |
| 195 | Yong Loo Lin School of Medicine | 3 | 214 | 797 | 426 |
| 196 | Freie Universität Berlin | 3 | 212 | 2.4K | 1.2K |
| 197 | Helsinki University of Technology | 2 | 209 | 839 | 437 |
| 198 | University of Turku | 2 | 208 | 2.9K | 1.1K |
| 199 | École Normale Supérieure | 2 | 207 | 1.0K | 473 |
| 200 | Russian Academy of Sciences | 1 | 205 | 5.8K | 1.7K |
| 201 | Bar-Ilan University | 2 | 200 | 2.2K | 729 |
| 202 | Trinity College Dublin | 1 | 197 | 3.4K | 917 |
| 202 | Trinity College | 1 | 197 | 2.7K | 676 |
| 204 | University of Exeter | 3 | 196 | 2.4K | 1.3K |
| 205 | Texas Children's Hospital | 1 | 194 | 919 | 292 |
| 206 | Lawrence Livermore National Laboratory | 2 | 191 | 1.1K | 640 |
| 207 | University of Calgary | 2 | 189 | 5.5K | 2.4K |
| 208 | Broad Institute | 2 | 188 | 136 | 107 |
| 209 | McGovern Institute for Brain Research | 1 | 187 | 159 | 84 |
| 210 | Helsinki University Central Hospital | 2 | 186 | 1.3K | 699 |
| 210 | Hosei University | 1 | 186 | 739 | 78 |
| 210 | Loyola University Chicago | 1 | 186 | 2.2K | 505 |
| 210 | University of Texas at Dallas | 2 | 186 | 1.7K | 792 |
| 214 | Massachusetts General Hospital | 4 | 184 | 2.7K | 2.1K |
| 215 | Emory University | 3 | 180 | 3.2K | 2.0K |
| 215 | United States National Library of Medicine | 3 | 180 | 209 | 203 |
| 217 | California State University | 1 | 178 | 5.5K | 1.3K |
| 217 | University of Connecticut Health Center | 1 | 178 | 1.6K | 626 |
| 217 | Université PSL Paris | 1 | 178 | 4 | 2 |
| 220 | University of Massachusetts | 3 | 173 | 3.1K | 1.9K |
| 221 | University of Geneva | 2 | 172 | 2.9K | 1.5K |
| 222 | Wageningen University & Research | 4 | 170 | 1.6K | 1.4K |
| 222 | Hadassah Medical Center | 3 | 170 | 444 | 276 |
| 222 | Academia Sinica | 3 | 170 | 2.1K | 1.2K |
| 225 | University of Hong Kong | 1 | 169 | 6.7K | 2.5K |
| 225 | McMaster University | 1 | 169 | 6.7K | 2.5K |
| 225 | Fred Hutchinson Cancer Research Center | 6 | 169 | 509 | 637 |
| 228 | California NanoSystems Institute | 1 | 168 | 169 | 91 |
| 229 | Kyoto University | 2 | 167 | 5.6K | 2.7K |
| 230 | University of Pennsylvania | 6 | 165 | 4.0K | 3.7K |
| 231 | Erasmus MC | 3 | 162 | 1.7K | 1.2K |
| 232 | Stowers Institute for Medical Research | 2 | 160 | 114 | 103 |
| 233 | Novo Nordisk | 1 | 158 | 603 | 253 |
| 234 | Katholieke Universiteit Leuven | 3 | 157 | 4.4K | 3.0K |
| 234 | Christian Albrechts Universität zu Kiel | 1 | 157 | 728 | 188 |
| 236 | Public Health Research Institute | 3 | 154 | 68 | 89 |
| 236 | National University Health System | 1 | 154 | 863 | 231 |
| 238 | Johns Hopkins School of Medicine | 1 | 153 | 2.8K | 1.3K |
| 238 | Structural Genomics Consortium | 1 | 153 | 85 | 47 |
| 240 | Earlham Institute | 1 | 152 | 65 | 37 |
| 241 | University of Manchester | 5 | 151 | 4.0K | 3.4K |
| 242 | Sheba Medical Center | 2 | 150 | 977 | 542 |
| 243 | Karlsruhe Institute of Technology | 2 | 149 | 2.6K | 1.4K |
| 244 | University of Amsterdam | 1 | 147 | 6.8K | 3.0K |
| 244 | University of Western Australia | 1 | 147 | 5.4K | 2.3K |
| 246 | National Institute of Environmental Health Sciences | 1 | 146 | 1.1K | 577 |
| 247 | Tokyo Metropolitan University | 1 | 145 | 1.9K | 488 |
| 247 | Agricultural Research Service | 1 | 145 | 1.2K | 681 |
| 247 | University of Tennessee | 2 | 145 | 4.6K | 2.3K |
| 250 | University of Barcelona | 4 | 143 | 3.1K | 2.3K |
| 251 | Colorado School of Mines | 1 | 139 | 1.3K | 502 |
| 251 | Helmholtz Centre for Infection Research | 1 | 139 | 274 | 143 |
| 251 | Joint Genome Institute | 1 | 139 | 109 | 77 |
| 251 | Agency for Science, Technology and Research | 1 | 139 | 634 | 266 |
| 255 | Butler University | 1 | 138 | 314 | 41 |
| 256 | Florida Institute for Human and Machine Cognition | 1 | 137 | 17 | 6 |
| 257 | Osaka University | 3 | 135 | 3.4K | 2.1K |
| 258 | National Institute of Advanced Industrial Science and Technology | 1 | 134 | 1.7K | 626 |
| 258 | Benaroya Research Institute | 1 | 134 | 102 | 68 |
| 258 | Merck & Co. | 1 | 134 | 655 | 450 |
| 261 | Merrimack Pharmaceuticals (United States) | 1 | 132 | 14 | 9 |
| 262 | Boğaziçi University | 1 | 131 | 1.4K | 353 |
| 262 | National Hellenic Research Foundation | 1 | 131 | 383 | 133 |
| 262 | Scuola Normale Superiore di Pisa | 1 | 131 | 608 | 176 |
| 265 | Swiss Institute of Bioinformatics | 5 | 129 | 45 | 99 |
| 266 | University of Maryland | 3 | 128 | 5.7K | 4.1K |
| 267 | New York Genome Center | 1 | 125 | 60 | 42 |
| 268 | National University of Ireland | 1 | 122 | 2.4K | 917 |
| 269 | University of North Carolina at Chapel Hill | 4 | 121 | 3.0K | 2.8K |
| 269 | Michigan State University | 1 | 121 | 8.0K | 3.6K |
| 271 | Washington University School of Medicine | 3 | 119 | 2.4K | 2.0K |
| 271 | University of Iowa | 1 | 119 | 6.5K | 3.0K |
| 273 | London Research Institute | 2 | 117 | 104 | 137 |
| 274 | Sun Yat-sen University | 1 | 116 | 5.3K | 2.2K |
| 275 | Nazarbayev University | 1 | 115 | 667 | 130 |
| 276 | Columbia University Irving Medical Center | 2 | 114 | 1.8K | 1.1K |
| 277 | Academic Medical Center | 1 | 113 | 1.6K | 938 |
| 278 | Hungarian Academy of Sciences | 2 | 112 | 2.2K | 1.3K |
| 278 | National Institute on Alcohol Abuse and Alcoholism | 1 | 112 | 438 | 312 |
| 280 | University of Medicine and Dentistry of New Jersey | 1 | 111 | 848 | 458 |
| 281 | Leonard M. Miller School of Medicine | 2 | 110 | 2.1K | 1.4K |
| 282 | University of Nottingham | 2 | 108 | 5.1K | 3.2K |
| 282 | Argonne National Laboratory | 1 | 108 | 1.6K | 919 |
| 282 | Universitat Politècnica de Catalunya | 2 | 108 | 1.5K | 872 |
| 285 | Nara Institute of Science and Technology | 2 | 105 | 600 | 350 |
| 286 | University of Extremadura | 1 | 104 | 2.1K | 658 |
| 286 | Umeå University | 1 | 104 | 3.9K | 1.9K |
| 288 | Ministry of Health | 1 | 103 | 3.3K | 1.1K |
| 288 | Babraham Institute | 2 | 103 | 132 | 153 |
| 290 | Ruhr University Bochum | 2 | 102 | 2.9K | 1.8K |
| 290 | University of Alberta | 2 | 102 | 7.2K | 4.4K |
| 290 | Cooperative Institute for Research in Environmental Sciences | 1 | 102 | 288 | 207 |
| 290 | Canadian Institute for Advanced Research | 1 | 102 | 123 | 123 |
| 290 | National Institute on Aging | 1 | 102 | 959 | 682 |
| 295 | Goethe University Frankfurt | 3 | 98 | 2.0K | 1.7K |
| 295 | Capital Normal University | 1 | 98 | 1.0K | 349 |
| 297 | Tokyo Medical University | 1 | 97 | 1.2K | 468 |
| 298 | Üsküdar University | 1 | 96 | 169 | 33 |
| 298 | Antalya Bilim University | 1 | 96 | 80 | 27 |
| 298 | Sabancı University | 1 | 96 | 781 | 312 |
| 301 | University of Jena | 1 | 94 | 3.0K | 1.5K |
| 301 | TU Dresden | 2 | 94 | 2.8K | 1.9K |
| 301 | AMOLF | 1 | 94 | 115 | 82 |
| 304 | Biohub | 1 | 91 | 50 | 32 |
| 304 | Ludwig Cancer Research | 1 | 91 | 654 | 463 |
| 306 | St. Michael's Hospital | 1 | 90 | 1.5K | 809 |
| 306 | Princess Margaret Hospital | 1 | 90 | 871 | 470 |
| 306 | Medical University of South Carolina | 2 | 90 | 2.5K | 1.8K |
| 309 | RWTH Aachen University | 2 | 89 | 3.2K | 2.2K |
| 310 | Monash University | 3 | 86 | 5.1K | 4.2K |
| 311 | University of Erlangen-Nuremberg | 2 | 83 | 1.6K | 1.3K |
| 311 | University of Bath | 3 | 83 | 2.0K | 1.9K |
| 311 | Korea University of Science and Technology | 1 | 83 | 4.3K | 1.7K |
| 314 | Queen's University Belfast | 1 | 82 | 3.9K | 1.9K |
| 315 | University of Illinois at Urbana–Champaign | 1 | 81 | 6.2K | 3.5K |
| 315 | Université de Paris | 2 | 81 | 1.1K | 599 |
| 317 | Lund University | 1 | 80 | 6.9K | 3.9K |
| 317 | Children's Hospital Boston | 1 | 80 | 781 | 655 |
| 317 | Harvard Stem Cell Institute | 1 | 80 | 172 | 158 |
| 320 | University of Sydney | 1 | 79 | 9.3K | 5.2K |
| 320 | La Trobe University | 1 | 79 | 3.9K | 2.0K |
| 322 | UC San Diego School of Medicine | 1 | 78 | 653 | 454 |
| 323 | Technische Universität Darmstadt | 2 | 77 | 1.7K | 1.3K |
| 323 | Norwegian University of Science and Technology | 2 | 77 | 3.2K | 2.4K |
| 323 | University of Miami | 1 | 77 | 5.3K | 2.9K |
| 323 | Tor Vergata University of Rome | 2 | 77 | 1.5K | 1.1K |
| 323 | University of Texas at El Paso | 1 | 77 | 1.8K | 721 |
| 328 | University of Strasbourg | 1 | 76 | 2.3K | 1.2K |
| 328 | Grenoble University - Grenoble Alpes University | 1 | 76 | 1.3K | 517 |
| 328 | Université de Sherbrooke | 4 | 76 | 1.2K | 1.4K |
| 328 | University of Regina | 2 | 76 | 888 | 595 |
| 328 | Institute For Medical Research, Israel-Canada | 1 | 76 | 26 | 25 |
| 333 | Scripps Research | 2 | 74 | 1.1K | 1.1K |
| 334 | Salk Institute for Biological Studies | 3 | 72 | 196 | 350 |
| 334 | Beatson West of Scotland Cancer Centre | 1 | 72 | 227 | 166 |
| 336 | National Polytechnic Institute of Toulouse | 1 | 69 | 58 | 32 |
| 337 | Royal Netherlands Academy of Arts and Sciences | 2 | 68 | 75 | 142 |
| 337 | Austrian Academy of Sciences | 3 | 68 | 503 | 673 |
| 337 | Utah State University | 1 | 68 | 2.9K | 1.6K |
| 337 | Rose–Hulman Institute of Technology | 1 | 68 | 124 | 54 |
| 337 | Virginia Commonwealth University | 1 | 68 | 4.6K | 2.6K |
| 337 | Jena University Hospital | 2 | 68 | 479 | 406 |
| 343 | Microsoft Research | 1 | 67 | 589 | 446 |
| 344 | Albert Einstein College of Medicine | 1 | 66 | 2.8K | 1.9K |
| 344 | Humanitas University | 1 | 66 | 432 | 211 |
| 346 | University of Massachusetts Boston | 2 | 65 | 635 | 582 |
| 346 | Pohang University of Science and Technology | 1 | 65 | 1.4K | 894 |
| 348 | Durham University | 1 | 64 | 4.0K | 2.2K |
| 348 | University of São Paulo | 1 | 64 | 9.4K | 4.8K |
| 348 | Paris Descartes University | 1 | 64 | 2.4K | 1.6K |
| 348 | Wilhelmina Children's Hospital | 1 | 64 | 364 | 278 |
| 352 | Collège de France | 1 | 63 | 608 | 467 |
| 352 | Sorbonne University | 1 | 63 | 2.3K | 960 |
| 352 | Sorbonne Universités | 1 | 63 | 1.3K | 848 |
| 355 | Sony Computer Science Laboratories | 1 | 62 | 21 | 16 |
| 356 | University of Hamburg | 1 | 61 | 4.2K | 2.5K |
| 356 | University of Bristol | 1 | 61 | 6.4K | 4.0K |
| 356 | Ontario Institute for Cancer Research | 2 | 61 | 98 | 163 |
| 359 | Eindhoven University of Technology | 1 | 60 | 2.7K | 1.8K |
| 359 | Eötvös Loránd University | 1 | 60 | 2.1K | 1.1K |
| 359 | Saarland University | 1 | 60 | 2.8K | 1.7K |
| 362 | Southern University of Science and Technology | 4 | 59 | 572 | 627 |
| 363 | Niels Bohr Institute | 1 | 58 | 300 | 231 |
| 363 | Ryerson University | 1 | 58 | 2.2K | 1.1K |
| 363 | Kavli Institute of Nanoscience | 2 | 58 | 111 | 142 |
| 363 | Sanford Burnham Prebys Medical Discovery Institute | 1 | 58 | 420 | 447 |
| 367 | Huntsman Cancer Institute | 2 | 57 | 339 | 369 |
| 368 | Linköping University | 2 | 56 | 2.8K | 2.6K |
| 368 | University of Côte d'Azur | 2 | 56 | 392 | 359 |
| 370 | Western Washington University | 1 | 55 | 957 | 526 |
| 370 | Huazhong University of Science and Technology | 1 | 55 | 4.6K | 3.0K |
| 370 | CHDI Foundation | 1 | 55 | 17 | 20 |
| 373 | Medical University of Vienna | 2 | 54 | 2.2K | 2.0K |
| 373 | RIKEN | 1 | 54 | 1.5K | 987 |
| 373 | University of Veterinary Medicine Hanover, Foundation | 1 | 54 | 489 | 339 |
| 376 | Bielefeld University | 1 | 53 | 2.4K | 1.5K |
| 376 | University of Bonn | 1 | 53 | 4.5K | 3.0K |
| 376 | University of Rostock | 1 | 53 | 2.4K | 1.5K |
| 376 | University of Münster | 1 | 53 | 4.0K | 2.7K |
| 376 | University of Turin | 1 | 53 | 5.5K | 3.5K |
| 376 | University of Konstanz | 1 | 53 | 2.2K | 1.5K |
| 376 | Ankara University | 1 | 53 | 3.0K | 1.4K |
| 376 | Maastricht University | 1 | 53 | 3.6K | 2.5K |
| 376 | Karolinska University Hospital | 1 | 53 | 2.4K | 1.8K |
| 376 | University of Eastern Finland | 1 | 53 | 2.1K | 1.2K |
| 376 | University of Lorraine | 1 | 53 | 2.2K | 1.1K |
| 376 | Biomax Informatics AG | 1 | 53 | 11 | 21 |
| 376 | University of Maryland Biotechnology Institute | 1 | 53 | 266 | 302 |
| 376 | Quadram Institute | 1 | 53 | 218 | 178 |
| 376 | Hospital Universitario Virgen del Rocío | 1 | 53 | 677 | 416 |
| 376 | Laboratoire d'Ingénierie des Systèmes Biologiques et des Procédés | 1 | 53 | 37 | 36 |
| 392 | University of Pécs | 1 | 52 | 1.5K | 852 |
| 393 | Seattle Children's | 1 | 51 | 914 | 698 |
| 394 | Royal College of Surgeons in Ireland | 1 | 49 | 1.0K | 758 |
| 394 | Royal College of Surgeons of England | 1 | 49 | 543 | 380 |
| 396 | University of Bern | 2 | 48 | 3.7K | 3.5K |
| 396 | Friedrich Miescher Institute for Biomedical Research | 1 | 48 | 201 | 248 |
| 398 | University of South Florida | 2 | 46 | 4.2K | 3.8K |
| 398 | Leica Microsystems | 1 | 46 | 4 | 6 |
| 398 | University of Science and Technology of China | 1 | 46 | 3.2K | 2.1K |
| 398 | MRC Human Genetics Unit | 1 | 46 | 60 | 69 |
| 402 | University of Trento | 1 | 44 | 2.4K | 1.6K |
| 402 | Indiana University Bloomington | 1 | 44 | 1.7K | 1.2K |
| 402 | Indiana University | 1 | 44 | 6.5K | 4.4K |
| 405 | Czech Academy of Sciences | 1 | 42 | 3.4K | 2.4K |
| 405 | Shanghai Institute of Materia Medica | 1 | 42 | 482 | 424 |
| 407 | University Medical Center Freiburg | 1 | 41 | 915 | 836 |
| 407 | Leipzig University | 1 | 41 | 3.9K | 2.8K |
| 407 | University of Buenos Aires | 2 | 41 | 3.2K | 2.4K |
| 407 | Stony Brook University | 1 | 41 | 4.3K | 3.3K |
| 407 | Budapest University of Technology and Economics | 1 | 41 | 1.7K | 1.0K |
| 407 | University of North Carolina | 2 | 41 | 4.9K | 4.8K |
| 407 | Cancer Research Center | 1 | 41 | 265 | 264 |
| 414 | National Institute of Standards and Technology | 1 | 40 | 1.9K | 1.5K |
| 414 | SLAC National Accelerator Laboratory | 1 | 40 | 409 | 339 |
| 416 | University of Bologna | 1 | 39 | 6.8K | 4.8K |
| 416 | Atatürk University | 1 | 39 | 1.9K | 1.1K |
| 416 | University of Essex | 1 | 39 | 2.4K | 1.7K |
| 419 | University of Salzburg | 1 | 38 | 1.3K | 1.0K |
| 419 | Addenbrooke's Hospital | 1 | 38 | 1.7K | 1.4K |
| 419 | University of Saskatchewan | 1 | 38 | 4.8K | 3.5K |
| 422 | Dartmouth College | 1 | 37 | 2.6K | 2.1K |
| 423 | Colorado State University | 1 | 36 | 4.7K | 3.7K |
| 423 | University of Houston | 2 | 36 | 3.2K | 3.2K |
| 425 | Stellenbosch University | 1 | 35 | 3.5K | 2.5K |
| 426 | University of Colorado Boulder | 1 | 34 | 3.2K | 2.7K |
| 427 | Tampere University of Technology | 1 | 33 | 1.2K | 1.1K |
| 427 | University of Tampere | 1 | 33 | 2.9K | 2.2K |
| 427 | Tampere University Hospital | 1 | 33 | 1.1K | 1.0K |
| 430 | University of Aberdeen | 1 | 32 | 4.4K | 3.5K |
| 430 | Scottish Universities Physics Alliance | 1 | 32 | 261 | 248 |
| 432 | Universitätsmedizin Göttingen | 1 | 31 | 753 | 634 |
| 432 | Berlin Institute of Health at Charité - Universitätsmedizin Berlin | 1 | 31 | 738 | 581 |
| 434 | University of California, Irvine | 1 | 30 | 3.0K | 2.8K |
| 435 | Pfizer | 1 | 29 | 951 | 987 |
| 435 | Research Institute of Molecular Pathology | 1 | 29 | 113 | 168 |
| 437 | Hong Kong Baptist University | 1 | 27 | 1.9K | 1.7K |
| 437 | City University of Hong Kong | 1 | 27 | 3.4K | 2.9K |
| 437 | Hong Kong University of Science and Technology | 1 | 27 | 2.5K | 2.3K |
| 437 | University Medical Center Groningen | 1 | 27 | 2.2K | 2.0K |
| 437 | Ginkgo Bioworks | 2 | 27 | 2 | 8 |
| 442 | Janelia Research Campus | 1 | 25 | 116 | 169 |
| 442 | New Jersey Institute of Technology | 1 | 25 | 1.4K | 1.3K |
| 442 | Wistar Institute | 1 | 25 | 426 | 525 |
| 442 | Fluidigm | 1 | 25 | 23 | 41 |
| 442 | United States Army Corps of Engineers | 1 | 25 | 169 | 167 |
| 447 | Arizona State University | 1 | 24 | 6.6K | 5.6K |
| 448 | University of Missouri | 1 | 22 | 6.6K | 5.7K |
| 448 | Koç University | 1 | 22 | 1.3K | 1.2K |
| 448 | Institut Pasteur de Lille | 1 | 22 | 458 | 597 |
| 448 | University of Lille | 1 | 22 | 1.2K | 1.2K |
| 448 | Garvan Institute of Medical Research | 1 | 22 | 480 | 618 |
| 448 | University of Pittsburgh Medical Center | 1 | 22 | 2.0K | 1.9K |
| 448 | University of Washington Bothell | 1 | 22 | 101 | 149 |
| 448 | MRC London Institute of Medical Sciences | 2 | 22 | 74 | 203 |
| 448 | Nottingham University Hospitals NHS Trust | 1 | 22 | 697 | 651 |
| 448 | Data61 | 1 | 22 | 79 | 111 |
| 458 | Ulm University | 1 | 21 | 2.0K | 2.0K |
| 458 | St. Jude Children's Research Hospital | 1 | 21 | 1.1K | 1.3K |
| 458 | VTT Technical Research Centre of Finland | 1 | 21 | 1.1K | 1.2K |
| 458 | EMBL Australia | 1 | 21 | 16 | 28 |
| 458 | University of Maryland, Baltimore County | 1 | 21 | 1.1K | 1.3K |
| 463 | University of Regensburg | 1 | 20 | 2.8K | 2.8K |
| 463 | New Mexico State University | 1 | 20 | 2.2K | 2.0K |
| 465 | Uppsala University Hospital | 1 | 19 | 776 | 977 |
| 465 | University of Surrey | 1 | 19 | 3.7K | 3.6K |
| 465 | Rothamsted Research | 1 | 19 | 646 | 757 |
| 465 | Institut Laue–Langevin | 1 | 19 | 335 | 370 |
| 469 | Ruijin Hospital | 1 | 18 | 1.1K | 1.1K |
| 469 | Mathematical Institute, University of Oxford | 1 | 18 | 571 | 582 |
| 471 | Vaccine Research Center | 1 | 17 | 61 | 100 |
| 471 | Montpellier University Hospital | 1 | 17 | 340 | 418 |
| 471 | Duke University Human Vaccine Institute | 1 | 17 | 30 | 45 |
| 474 | University of Oslo | 1 | 16 | 6.3K | 6.1K |
| 475 | Institute for Molecular Medicine Finland | 1 | 15 | 43 | 86 |
| 475 | European Molecular Biology Organization | 7 | 15 | 3 | 9 |
| 477 | University of Sheffield | 1 | 14 | 6.6K | 6.6K |
| 477 | Okayama University | 1 | 14 | 2.8K | 2.7K |
| 477 | Cincinnati Children's Hospital Medical Center | 1 | 14 | 1.7K | 2.1K |
| 480 | University of St Andrews | 1 | 13 | 2.2K | 2.5K |
| 480 | AstraZeneca | 1 | 13 | 921 | 1.2K |
| 480 | University of Osijek | 1 | 13 | 527 | 533 |
| 480 | Krembil Research Institute | 1 | 13 | 224 | 365 |
| 480 | Emmanuel College | 1 | 13 | 201 | 207 |
| 480 | Yale Cancer Center | 1 | 13 | 225 | 320 |
| 486 | University of Melbourne | 1 | 12 | 9.1K | 8.9K |
| 486 | University of Alabama at Birmingham | 1 | 12 | 4.8K | 5.1K |
| 486 | Helen Wills Neuroscience Institute | 1 | 12 | 137 | 237 |
| 489 | University of Bordeaux | 1 | 11 | 2.6K | 3.0K |
| 489 | University of Düsseldorf | 1 | 11 | 1.9K | 2.5K |
| 489 | University of Florida | 1 | 11 | 9.6K | 9.4K |
| 489 | University of Georgia | 1 | 11 | 6.1K | 6.3K |
| 489 | Idorsia (Switzerland) | 1 | 11 | 3 | 7 |
| 494 | Yonsei University | 1 | 10 | 5.3K | 5.5K |
| 494 | Autonomous University of Barcelona | 1 | 10 | 5.0K | 5.3K |
| 494 | Ben-Gurion University of the Negev | 1 | 10 | 4.6K | 5.2K |
| 494 | Ritsumeikan University | 1 | 10 | 1.5K | 1.6K |
| 498 | University of Bergen | 1 | 9 | 4.1K | 4.9K |
| 498 | University of Wrocław | 1 | 9 | 1.9K | 1.9K |
| 498 | NUI Galway | 1 | 9 | 1.4K | 2.0K |
| 498 | University of Valladolid | 1 | 9 | 2.2K | 2.4K |
| 498 | Charles Darwin University | 1 | 9 | 914 | 1.4K |
| 498 | United States Naval Research Laboratory | 1 | 9 | 1.1K | 1.4K |
| 498 | Norwegian University of Life Sciences | 1 | 9 | 1.2K | 1.7K |
| 498 | Menzies School of Health Research | 1 | 9 | 345 | 526 |
| 506 | Merck KGaA | 1 | 8 | 364 | 594 |
| 506 | Oslo University Hospital | 1 | 8 | 1.8K | 2.2K |
| 508 | Chung-Ang University | 1 | 7 | 2.2K | 2.9K |
| 508 | Oxford University Hospitals NHS Trust | 1 | 7 | 709 | 899 |
| 510 | University of Minho | 1 | 6 | 2.8K | 3.8K |
| 511 | University of Southern California | 1 | 5 | 7.9K | 9.2K |
| 512 | Brandenburg University of Technology | 1 | 3 | 573 | 1.1K |
| 513 | Nanjing Tech University | 1 | 2 | 1.2K | 1.7K |
| 514 | Food and Drug Administration | 1 | 1 | 907 | 1.7K |
| 514 | Kyushu University | 1 | 1 | 5.6K | 7.4K |
| 514 | Grenoble Institute of Technology | 1 | 1 | 769 | 1.4K |
| 517 | Texas Tech University Health Sciences Center | 1 | 0 | 1.5K | 2.7K |
