| 1 | Bielefeld University | 14 | 10.1K | 415 | 9 |
| 2 | University of New South Wales | 61 | 8.6K | 415 | 75 |
| 3 | University of Connecticut | 23 | 6.8K | 798 | 49 |
| 4 | University of Aberdeen | 18 | 6.7K | 582 | 29 |
| 5 | University of Melbourne | 64 | 6.4K | 559 | 176 |
| 6 | University of Otago | 26 | 5.7K | 370 | 17 |
| 7 | University of Oxford | 85 | 5.0K | 526 | 418 |
| 8 | University of Exeter | 59 | 5.0K | 116 | 51 |
| 9 | United States Geological Survey | 102 | 4.7K | 184 | 131 |
| 10 | Joseph Fourier University | 7 | 4.5K | 530 | 33 |
| 11 | Natural History Museum | 52 | 4.4K | 111 | 45 |
| 12 | University of Cambridge | 49 | 3.8K | 855 | 496 |
| 13 | Harvard University | 33 | 3.5K | 2461 | 1163 |
| 14 | College of William & Mary | 6 | 3.4K | 464 | 21 |
| 15 | Virginia Institute of Marine Science | 4 | 3.4K | 152 | 10 |
| 16 | Wageningen University & Research | 34 | 3.3K | 413 | 218 |
| 17 | University of St Andrews | 72 | 3.3K | 63 | 73 |
| 18 | Iowa State University | 20 | 3.2K | 766 | 164 |
| 19 | Patuxent Wildlife Research Center | 40 | 3.2K | 27 | 14 |
| 20 | Tsinghua University | 10 | 3.1K | 2334 | 317 |
| 21 | University of Helsinki | 40 | 3.0K | 629 | 301 |
| 22 | University of Queensland | 48 | 3.0K | 717 | 417 |
| 23 | National Tsing Hua University | 9 | 2.9K | 680 | 79 |
| 24 | University of Hong Kong | 10 | 2.9K | 2253 | 216 |
| 25 | University of Gothenburg | 13 | 2.8K | 1390 | 239 |
| 26 | University of Michigan | 16 | 2.7K | 3439 | 971 |
| 27 | University of California, Berkeley | 46 | 2.7K | 759 | 681 |
| 28 | University of Montpellier | 53 | 2.5K | 190 | 134 |
| 29 | Imperial College London | 47 | 2.5K | 926 | 704 |
| 30 | Muséum national d'histoire naturelle | 25 | 2.5K | 127 | 38 |
| 31 | University of Sheffield | 54 | 2.5K | 328 | 293 |
| 32 | Chinese Academy of Sciences | 47 | 2.5K | 1923 | 1043 |
| 33 | University of Jena | 18 | 2.4K | 458 | 61 |
| 34 | Centre for Ecology & Hydrology | 42 | 2.4K | 42 | 54 |
| 35 | McGill University | 14 | 2.3K | 2476 | 598 |
| 36 | University of Santiago de Compostela | 6 | 2.3K | 814 | 46 |
| 37 | Manaaki Whenua – Landcare Research | 14 | 2.3K | 80 | 27 |
| 38 | Macquarie University | 23 | 2.3K | 345 | 92 |
| 39 | Smithsonian Tropical Research Institute | 34 | 2.2K | 36 | 34 |
| 40 | University College London | 38 | 2.2K | 1079 | 736 |
| 41 | Duke University | 27 | 2.1K | 1388 | 769 |
| 42 | Uppsala University | 14 | 2.0K | 1600 | 433 |
| 43 | University of Oslo | 19 | 2.0K | 1155 | 340 |
| 44 | University of Florida | 47 | 2.0K | 874 | 697 |
| 45 | University of Wisconsin–Madison | 29 | 1.9K | 1584 | 954 |
| 46 | University of Turin | 4 | 1.9K | 3190 | 242 |
| 47 | City University of New York | 6 | 1.9K | 1876 | 129 |
| 48 | Graduate Center, CUNY | 5 | 1.9K | 799 | 41 |
| 49 | Albertina | 1 | 1.9K | 194 | 1 |
| 50 | Norwegian University of Science and Technology | 31 | 1.9K | 306 | 162 |
| 51 | North Carolina State University | 26 | 1.9K | 685 | 316 |
| 52 | Claude Bernard University Lyon 1 | 24 | 1.9K | 589 | 216 |
| 53 | University of Tasmania | 34 | 1.9K | 175 | 104 |
| 54 | Columbia University | 10 | 1.9K | 3640 | 969 |
| 55 | Max Planck Institute for Ornithology | 20 | 1.8K | 21 | 9 |
| 56 | University of Lausanne | 21 | 1.8K | 510 | 227 |
| 57 | Umeå University | 13 | 1.8K | 806 | 215 |
| 58 | University of Jyväskylä | 13 | 1.8K | 388 | 58 |
| 59 | Centre d'Écologie Fonctionnelle et Évolutive | 23 | 1.7K | 24 | 23 |
| 60 | Zoological Society of London | 28 | 1.7K | 26 | 22 |
| 61 | University of Copenhagen | 26 | 1.7K | 1198 | 686 |
| 62 | Yale University | 18 | 1.7K | 1772 | 898 |
| 63 | University of East Anglia | 23 | 1.7K | 298 | 162 |
| 64 | Arizona State University | 9 | 1.6K | 2176 | 420 |
| 65 | National Marine Fisheries Service | 33 | 1.6K | 72 | 62 |
| 66 | Western Sydney University | 6 | 1.6K | 1028 | 69 |
| 67 | University of Glasgow | 20 | 1.6K | 978 | 442 |
| 68 | University of Minnesota | 33 | 1.5K | 1458 | 1125 |
| 69 | University of California, Davis | 27 | 1.5K | 1275 | 914 |
| 70 | Liverpool John Moores University | 10 | 1.5K | 343 | 43 |
| 71 | University of Zurich | 29 | 1.5K | 685 | 521 |
| 71 | Colorado State University | 37 | 1.5K | 327 | 333 |
| 73 | Australian National University | 34 | 1.5K | 496 | 423 |
| 74 | University of Washington | 28 | 1.5K | 1916 | 1367 |
| 75 | Smithsonian Conservation Biology Institute | 18 | 1.5K | 18 | 8 |
| 76 | Helmholtz Centre for Environmental Research | 30 | 1.4K | 67 | 63 |
| 77 | University of Wyoming | 17 | 1.4K | 259 | 96 |
| 77 | CSIRO Oceans and Atmosphere | 14 | 1.4K | 58 | 40 |
| 79 | German Centre for Integrative Biodiversity Research | 35 | 1.4K | 19 | 25 |
| 80 | Sustainability Institute | 23 | 1.4K | 22 | 18 |
| 81 | City, University of London | 3 | 1.4K | 1580 | 87 |
| 82 | University of Konstanz | 24 | 1.4K | 174 | 123 |
| 83 | Australian Research Council | 5 | 1.4K | 217 | 42 |
| 84 | Queen's University Belfast | 7 | 1.4K | 1386 | 184 |
| 85 | University of Maryland | 29 | 1.4K | 1307 | 1018 |
| 86 | Ghent University | 27 | 1.4K | 965 | 703 |
| 87 | Deakin University | 22 | 1.4K | 410 | 178 |
| 88 | Korea University of Science and Technology | 34 | 1.3K | 967 | 459 |
| 89 | Swansea University | 22 | 1.3K | 235 | 107 |
| 90 | Cornell University | 42 | 1.3K | 830 | 1094 |
| 91 | Washington State University | 11 | 1.3K | 1174 | 353 |
| 92 | University of Hawaii at Manoa | 11 | 1.3K | 362 | 92 |
| 93 | University of Western Australia | 19 | 1.3K | 1025 | 524 |
| 94 | University of Leeds | 27 | 1.3K | 759 | 570 |
| 95 | University of Idaho | 17 | 1.3K | 227 | 101 |
| 96 | Ludwig Maximilian University of Munich | 17 | 1.2K | 1571 | 724 |
| 97 | Technical University of Denmark | 16 | 1.2K | 806 | 422 |
| 98 | University of Toronto | 28 | 1.2K | 2275 | 1687 |
| 98 | University of California, Santa Barbara | 15 | 1.2K | 730 | 433 |
| 100 | University of Edinburgh | 28 | 1.2K | 958 | 806 |
| 101 | University of Tartu | 10 | 1.2K | 481 | 94 |
| 102 | Stockholm University | 17 | 1.2K | 734 | 405 |
| 102 | Université de Montréal | 18 | 1.2K | 1216 | 576 |
| 104 | Swedish University of Agricultural Sciences | 35 | 1.2K | 229 | 251 |
| 105 | University of British Columbia | 34 | 1.2K | 1251 | 1295 |
| 106 | Université du Québec à Montréal | 11 | 1.2K | 405 | 77 |
| 107 | University of Alberta | 30 | 1.2K | 1144 | 910 |
| 108 | Université de Lyon | 17 | 1.2K | 352 | 155 |
| 109 | Czech Academy of Sciences | 20 | 1.2K | 846 | 346 |
| 109 | National Zoological Park | 21 | 1.2K | 31 | 22 |
| 111 | University of York | 24 | 1.2K | 369 | 307 |
| 112 | National Center for Ecological Analysis and Synthesis | 6 | 1.1K | 31 | 22 |
| 113 | Stony Brook University | 18 | 1.1K | 619 | 423 |
| 114 | University of California, Los Angeles | 29 | 1.1K | 1280 | 1289 |
| 115 | Centre for Environment, Fisheries and Aquaculture Science | 12 | 1.1K | 43 | 33 |
| 116 | University of Arizona | 20 | 1.1K | 1338 | 896 |
| 117 | Goethe University Frankfurt | 5 | 1.1K | 1878 | 291 |
| 117 | University of Notre Dame | 6 | 1.1K | 1469 | 266 |
| 119 | Great Lakes Institute of Management | 10 | 1.1K | 118 | 34 |
| 120 | Institute for Marine and Antarctic Studies | 19 | 1.1K | 43 | 28 |
| 121 | University of Southampton | 22 | 1.1K | 846 | 619 |
| 122 | University of Tennessee | 9 | 1.0K | 2390 | 674 |
| 123 | University of Windsor | 9 | 1.0K | 406 | 59 |
| 124 | University College Cork | 3 | 1.0K | 1652 | 177 |
| 126 | Princeton University | 12 | 1.0K | 1122 | 675 |
| 127 | Museu de Ciências Naturais da Fundação Zoobotânica do Rio Grande do Sul | 11 | 1.0K | 7 | 3 |
| 128 | University of Canberra | 13 | 1.0K | 110 | 26 |
| 129 | Oeko Institut | 12 | 1.0K | 23 | 5 |
| 130 | University of Bern | 18 | 1.0K | 1016 | 616 |
| 131 | Stanford University | 11 | 1.0K | 3604 | 1820 |
| 132 | Kansas State University | 4 | 993 | 1742 | 265 |
| 133 | Smithsonian Institution | 17 | 981 | 165 | 92 |
| 133 | Agroscope | 4 | 981 | 257 | 49 |
| 135 | CSIRO Land and Water | 9 | 971 | 128 | 80 |
| 136 | University of Bristol | 27 | 967 | 802 | 826 |
| 137 | University of Sydney | 14 | 963 | 2639 | 1278 |
| 138 | Chalmers University of Technology | 1 | 962 | 3248 | 315 |
| 138 | Sahlgrenska University Hospital | 1 | 962 | 2722 | 356 |
| 140 | University of Georgia | 16 | 951 | 1276 | 699 |
| 140 | Curtin University | 13 | 951 | 805 | 320 |
| 142 | National Centre for Earth Observation | 8 | 948 | 31 | 13 |
| 143 | Environmental Change Institute | 11 | 943 | 67 | 50 |
| 144 | University of Liverpool | 15 | 941 | 1285 | 646 |
| 145 | British Trust for Ornithology | 17 | 920 | 13 | 16 |
| 146 | Ohio State University | 15 | 916 | 2711 | 1451 |
| 147 | University of Freiburg | 15 | 914 | 759 | 475 |
| 147 | Aarhus University | 24 | 914 | 806 | 723 |
| 149 | Centre for Geogenetics | 3 | 889 | 37 | 14 |
| 150 | University of Antwerp | 15 | 881 | 673 | 396 |
| 151 | Washington University in St. Louis | 7 | 878 | 3107 | 1199 |
| 152 | University of Cape Town | 18 | 873 | 639 | 416 |
| 153 | MRC Centre for Global Infectious Disease Analysis | 3 | 863 | 74 | 15 |
| 154 | Linnaeus University | 28 | 856 | 154 | 161 |
| 155 | University of Duisburg-Essen | 6 | 855 | 1416 | 268 |
| 156 | Grenoble University - Grenoble Alpes University | 18 | 849 | 361 | 171 |
| 157 | James Cook University | 19 | 836 | 296 | 221 |
| 158 | Cooperative Research Centre | 6 | 825 | 398 | 141 |
| 159 | Vrije Universiteit Amsterdam | 9 | 817 | 2104 | 1027 |
| 160 | University of California, Riverside | 7 | 816 | 1293 | 470 |
| 161 | North Carolina Museum of Natural Sciences | 10 | 815 | 8 | 3 |
| 162 | Max Planck Society | 17 | 793 | 520 | 509 |
| 163 | University of Kentucky | 8 | 789 | 2365 | 768 |
| 164 | University of Illinois at Urbana–Champaign | 8 | 784 | 2940 | 1262 |
| 165 | University of Göttingen | 24 | 779 | 556 | 594 |
| 165 | University of Chinese Academy of Sciences | 15 | 779 | 1517 | 703 |
| 167 | Sapienza University of Rome | 15 | 774 | 2054 | 1069 |
| 168 | University of Montana | 11 | 773 | 204 | 135 |
| 169 | Eötvös Loránd University | 7 | 765 | 791 | 142 |
| 170 | Chatham University | 6 | 764 | 18 | 1 |
| 171 | Dakota State University | 5 | 752 | 159 | 16 |
| 172 | Queen Mary University of London | 9 | 751 | 1120 | 434 |
| 172 | Royal Botanic Gardens | 8 | 751 | 162 | 69 |
| 174 | North Dakota State University | 4 | 750 | 926 | 120 |
| 175 | King's Buildings | 8 | 740 | 282 | 144 |
| 176 | University of Vienna | 11 | 732 | 2015 | 938 |
| 177 | Trent University | 10 | 729 | 177 | 96 |
| 178 | University of Tromsø – The Arctic University of Norway | 18 | 721 | 272 | 248 |
| 178 | University of Basel | 10 | 721 | 1170 | 602 |
| 180 | James Hutton Institute | 17 | 720 | 50 | 68 |
| 181 | Queensland University of Technology | 13 | 718 | 755 | 417 |
| 182 | Norwich Research Park | 8 | 714 | 242 | 156 |
| 183 | Leibniz Association | 21 | 713 | 252 | 247 |
| 184 | Kunming Institute of Zoology | 5 | 706 | 167 | 41 |
| 185 | Monash University | 17 | 704 | 1976 | 1438 |
| 186 | Utrecht University | 13 | 698 | 1681 | 1215 |
| 187 | Radboud University Nijmegen | 11 | 697 | 1693 | 1009 |
| 188 | University of South Bohemia in České Budějovice | 9 | 695 | 225 | 77 |
| 189 | University of Bath | 7 | 693 | 1213 | 415 |
| 190 | Engineer Research and Development Center | 2 | 692 | 371 | 30 |
| 191 | Leipzig University | 19 | 685 | 636 | 468 |
| 192 | Microsoft | 6 | 683 | 333 | 149 |
| 193 | University of New England | 7 | 679 | 565 | 136 |
| 194 | University of Kansas | 9 | 678 | 1828 | 756 |
| 195 | American Museum of Natural History | 9 | 673 | 145 | 75 |
| 195 | Finnish Environment Institute | 9 | 673 | 96 | 66 |
| 197 | University of Colorado Boulder | 9 | 671 | 675 | 395 |
| 198 | Tampere University of Technology | 2 | 664 | 813 | 86 |
| 199 | Finnish Museum of Natural History | 9 | 657 | 49 | 20 |
| 200 | Massey University | 13 | 653 | 432 | 288 |
| 200 | Smithsonian Environmental Research Center | 14 | 653 | 122 | 107 |
| 202 | Bangor University | 11 | 652 | 271 | 172 |
| 203 | Luther University | 14 | 648 | 137 | 95 |
| 204 | University of Toledo | 3 | 647 | 1494 | 163 |
| 205 | Central Michigan University | 1 | 639 | 1638 | 34 |
| 205 | Shimane University | 1 | 639 | 1784 | 56 |
| 205 | IFREMER | 19 | 639 | 152 | 162 |
| 205 | United States Army | 1 | 639 | 1803 | 116 |
| 209 | Oregon State University | 22 | 638 | 581 | 682 |
| 210 | University of North Carolina | 8 | 633 | 3290 | 1447 |
| 211 | Autonomous University of Barcelona | 12 | 629 | 1344 | 723 |
| 211 | University of South Florida | 8 | 629 | 1907 | 722 |
| 213 | Dalhousie University | 18 | 628 | 702 | 661 |
| 214 | Climate Change Research Centre | 2 | 625 | 108 | 44 |
| 215 | University of Auckland | 23 | 621 | 662 | 861 |
| 216 | University of Adelaide | 13 | 612 | 1308 | 926 |
| 217 | National Park Service | 11 | 610 | 91 | 48 |
| 217 | Butterfly Conservation | 11 | 610 | 6 | 12 |
| 219 | University of Guelph | 12 | 605 | 820 | 569 |
| 220 | Stellenbosch University | 12 | 603 | 750 | 331 |
| 221 | Florida Museum of Natural History | 8 | 602 | 92 | 37 |
| 222 | University of Kent | 15 | 595 | 290 | 214 |
| 223 | Pennsylvania State University | 26 | 593 | 1364 | 1780 |
| 224 | NOAA | 21 | 590 | 128 | 163 |
| 225 | Victoria University of Wellington | 11 | 587 | 354 | 179 |
| 226 | Western Washington University | 1 | 583 | 1183 | 49 |
| 227 | Simon Fraser University | 11 | 582 | 823 | 491 |
| 228 | Lund University | 15 | 579 | 1668 | 1389 |
| 229 | Concordia University | 4 | 573 | 1641 | 295 |
| 230 | École Normale Supérieure | 7 | 570 | 686 | 332 |
| 231 | Louisiana State University | 7 | 566 | 2493 | 987 |
| 232 | The Nature Conservancy | 10 | 564 | 107 | 77 |
| 233 | University of Chicago | 8 | 562 | 3197 | 1672 |
| 234 | Université de Sherbrooke | 12 | 559 | 522 | 320 |
| 235 | Xi'an Jiaotong-Liverpool University | 2 | 554 | 546 | 23 |
| 236 | University of New Brunswick | 2 | 552 | 1557 | 173 |
| 236 | Canadian Rivers Institute | 2 | 552 | 74 | 11 |
| 238 | Charles University | 10 | 551 | 1604 | 687 |
| 239 | Purdue University | 16 | 550 | 1875 | 1589 |
| 240 | Daniel B. Warnell School of Forestry and Natural Resources | 9 | 549 | 81 | 37 |
| 241 | Forest Research | 10 | 547 | 172 | 97 |
| 242 | University of Groningen | 17 | 545 | 1300 | 1341 |
| 243 | Newcastle University | 11 | 543 | 1455 | 940 |
| 244 | Royal Society for the Protection of Birds | 6 | 541 | 36 | 19 |
| 245 | Laval University | 9 | 536 | 1707 | 896 |
| 246 | City College of New York | 4 | 535 | 559 | 144 |
| 247 | University of Reading | 16 | 534 | 549 | 613 |
| 248 | Marine Biological Laboratory | 2 | 533 | 469 | 147 |
| 248 | Cardiff University | 9 | 533 | 1503 | 865 |
| 250 | Santa Fe Institute | 6 | 529 | 124 | 90 |
| 251 | Utah State University | 13 | 519 | 432 | 367 |
| 252 | Michigan State University | 28 | 508 | 882 | 1544 |
| 253 | University of Würzburg | 17 | 507 | 713 | 754 |
| 253 | University of Barcelona | 11 | 507 | 1899 | 1179 |
| 255 | University of Oulu | 5 | 505 | 1594 | 526 |
| 256 | Tel Aviv University | 17 | 504 | 1510 | 1396 |
| 256 | University of Costa Rica | 4 | 504 | 493 | 68 |
| 258 | University of Rhode Island | 7 | 499 | 609 | 290 |
| 259 | ETH Zürich | 16 | 498 | 1489 | 1563 |
| 260 | Université libre de Bruxelles | 10 | 497 | 962 | 640 |
| 261 | University of Massachusetts Boston | 7 | 491 | 188 | 77 |
| 262 | Australian Institute of Marine Science | 13 | 490 | 53 | 69 |
| 263 | University of Salford | 4 | 489 | 872 | 157 |
| 264 | University of Bergen | 12 | 487 | 977 | 806 |
| 264 | University of Akron | 1 | 487 | 2102 | 206 |
| 266 | University of Nebraska–Lincoln | 12 | 482 | 866 | 693 |
| 267 | Durham University | 8 | 481 | 1185 | 585 |
| 268 | Florida International University | 10 | 472 | 641 | 408 |
| 268 | Fisheries and Oceans Canada | 12 | 472 | 163 | 150 |
| 270 | National Biodiversity Institute | 2 | 471 | 133 | 26 |
| 270 | University West | 14 | 471 | 139 | 108 |
| 272 | University of New Hampshire | 8 | 464 | 494 | 318 |
| 272 | Université du Québec à Rimouski | 6 | 464 | 146 | 40 |
| 272 | United States Forest Service | 19 | 464 | 289 | 332 |
| 275 | University of Natural Resources and Life Sciences, Vienna | 5 | 463 | 623 | 233 |
| 276 | University of the French Antilles | 6 | 462 | 86 | 22 |
| 277 | University of Stirling | 12 | 461 | 299 | 295 |
| 278 | Max Planck Institute for Demographic Research | 7 | 458 | 37 | 22 |
| 279 | Australian Antarctic Division | 7 | 453 | 62 | 44 |
| 280 | Baylor University | 4 | 448 | 1303 | 337 |
| 281 | Humboldt University of Berlin | 17 | 445 | 1050 | 1035 |
| 281 | Universidade Federal de Santa Maria | 1 | 445 | 2959 | 168 |
| 281 | Universidad Santa María | 1 | 445 | 456 | 16 |
| 284 | University of Vermont | 13 | 444 | 596 | 647 |
| 285 | University of California, Santa Cruz | 18 | 442 | 285 | 501 |
| 286 | Nicholas School of the Environment | 7 | 440 | 163 | 137 |
| 286 | Environment Agency | 2 | 440 | 329 | 88 |
| 288 | Trout Unlimited | 1 | 439 | 23 | 4 |
| 289 | University of Canterbury | 17 | 438 | 279 | 367 |
| 289 | University of São Paulo | 15 | 438 | 3392 | 2169 |
| 291 | Colorado Parks and Wildlife | 7 | 435 | 28 | 17 |
| 292 | University of Massachusetts Amherst | 19 | 432 | 155 | 323 |
| 293 | Murdoch University | 6 | 428 | 648 | 281 |
| 294 | National Oceanography Centre | 8 | 426 | 137 | 104 |
| 295 | University of Technology Sydney | 9 | 425 | 961 | 565 |
| 296 | Technion – Israel Institute of Technology | 2 | 420 | 3214 | 734 |
| 296 | University of KwaZulu-Natal | 10 | 420 | 877 | 436 |
| 298 | University of Manitoba | 5 | 417 | 2551 | 974 |
| 299 | Norwegian University of Life Sciences | 11 | 415 | 282 | 277 |
| 300 | University of Potsdam | 11 | 410 | 435 | 343 |
| 301 | University of Trento | 5 | 409 | 1026 | 338 |
| 302 | Flinders University | 8 | 406 | 989 | 547 |
| 302 | University of Utah | 8 | 406 | 2742 | 1713 |
| 302 | University of Maryland, Baltimore County | 1 | 406 | 1196 | 128 |
| 305 | University of California, Merced | 6 | 405 | 255 | 145 |
| 305 | Fauna and Flora International | 4 | 405 | 21 | 8 |
| 307 | Syracuse University | 3 | 404 | 2031 | 510 |
| 308 | Max Planck Institute for Biogeochemistry | 6 | 401 | 67 | 61 |
| 309 | Northeastern University | 5 | 399 | 2402 | 932 |
| 310 | Universidade Federal de Goiás | 7 | 397 | 762 | 144 |
| 311 | Pompeu Fabra University | 7 | 394 | 543 | 315 |
| 312 | University of Fribourg | 10 | 392 | 591 | 481 |
| 313 | University of Iceland | 3 | 391 | 1112 | 285 |
| 313 | Hostelling International USA | 7 | 391 | 30 | 25 |
| 315 | Virginia Tech | 14 | 390 | 1290 | 1236 |
| 316 | Zhejiang University | 5 | 389 | 3887 | 1604 |
| 317 | Max Planck Institute for Evolutionary Anthropology | 7 | 386 | 104 | 93 |
| 317 | Centre for Environmental Policy | 3 | 386 | 113 | 42 |
| 319 | Trinity College Dublin | 10 | 384 | 678 | 617 |
| 320 | University of Évora | 8 | 383 | 175 | 69 |
| 320 | Broadway | 8 | 383 | 706 | 605 |
| 322 | University of the Azores | 5 | 379 | 127 | 43 |
| 323 | East China Normal University | 3 | 377 | 1864 | 442 |
| 324 | University of Amsterdam | 15 | 376 | 1782 | 2017 |
| 325 | Peking University | 8 | 374 | 2727 | 1502 |
| 326 | The Australia Institute | 9 | 373 | 126 | 138 |
| 327 | University of Newcastle | 5 | 372 | 2223 | 1006 |
| 328 | Wildlife Conservation Society | 7 | 369 | 130 | 104 |
| 328 | Université de Moncton | 6 | 369 | 103 | 28 |
| 328 | Université de Yaoundé I | 5 | 369 | 265 | 62 |
| 331 | Swiss Ornithological Institute Sempach | 13 | 368 | 4 | 3 |
| 332 | University of Maine | 7 | 367 | 523 | 330 |
| 333 | Technische Universität Ilmenau | 3 | 365 | 451 | 76 |
| 334 | Aalto University | 3 | 362 | 1478 | 387 |
| 334 | Marine Biological Association of the United Kingdom | 5 | 362 | 78 | 54 |
| 336 | Institute for Biodiversity and Ecosystem Dynamics | 14 | 361 | 56 | 106 |
| 337 | Texas A&M University | 9 | 360 | 3304 | 2219 |
| 337 | École polytechnique fédérale de Lausanne | 11 | 360 | 785 | 813 |
| 337 | Sorbonne University | 18 | 360 | 1276 | 1375 |
| 340 | Arthur Rylah Institute for Environmental Research | 8 | 357 | 20 | 22 |
| 341 | Carleton University | 12 | 356 | 537 | 499 |
| 342 | University of Poitiers | 5 | 354 | 724 | 242 |
| 343 | Sorbonne | 6 | 350 | 1125 | 517 |
| 344 | Royal Botanic Garden Edinburgh | 4 | 348 | 81 | 43 |
| 345 | Leuphana University of Lüneburg | 3 | 345 | 353 | 100 |
| 345 | University of Saskatchewan | 5 | 345 | 2090 | 932 |
| 347 | Woods Hole Oceanographic Institution | 13 | 344 | 159 | 241 |
| 347 | University of the Sunshine Coast | 3 | 344 | 604 | 110 |
| 349 | Aix-Marseille University | 8 | 342 | 1857 | 1158 |
| 350 | Université catholique de Louvain | 6 | 339 | 1763 | 1185 |
| 351 | University of Regensburg | 9 | 338 | 845 | 706 |
| 352 | Dartmouth College | 4 | 337 | 1259 | 611 |
| 353 | Federal University of Rio Grande do Norte | 3 | 336 | 1689 | 194 |
| 353 | Pacific Biological Station | 5 | 336 | 70 | 42 |
| 355 | École Normale Supérieure de Lyon | 3 | 334 | 769 | 299 |
| 356 | University of Münster | 5 | 329 | 2091 | 1079 |
| 357 | University of Strasbourg | 8 | 326 | 1292 | 876 |
| 358 | University of London | 7 | 324 | 2396 | 1596 |
| 359 | University of French Guiana | 3 | 320 | 48 | 6 |
| 360 | United States Department of Agriculture | 4 | 318 | 1502 | 903 |
| 360 | Fera Science | 3 | 318 | 74 | 35 |
| 360 | Australian Museum | 5 | 318 | 119 | 69 |
| 363 | University of Southern Denmark | 10 | 317 | 955 | 902 |
| 363 | Royal Holloway, University of London | 6 | 317 | 515 | 352 |
| 363 | Region of Southern Denmark | 10 | 317 | 176 | 169 |
| 366 | Griffith University | 8 | 315 | 1399 | 987 |
| 367 | Boston University | 10 | 313 | 2066 | 1981 |
| 368 | Centre for the Study of Existential Risk | 2 | 312 | 9 | 3 |
| 368 | The James Cancer Hospital | 2 | 312 | 722 | 244 |
| 370 | Environment and Climate Change Canada | 11 | 311 | 214 | 237 |
| 371 | University of Essex | 4 | 306 | 1073 | 413 |
| 372 | International Union for Conservation of Nature | 2 | 304 | 93 | 36 |
| 373 | Microbial Ecology | 4 | 302 | 607 | 368 |
| 374 | University of Tübingen | 4 | 301 | 2817 | 1340 |
| 375 | King Juan Carlos University | 6 | 300 | 587 | 214 |
| 376 | Brown University | 1 | 299 | 6066 | 1667 |
| 376 | Khoury College of Computer Sciences | 1 | 299 | 346 | 26 |
| 378 | Commonwealth Scientific and Industrial Research Organisation | 12 | 296 | 69 | 186 |
| 379 | Federal Government of Belgium | 5 | 291 | 58 | 36 |
| 380 | University of Minho | 1 | 290 | 3722 | 546 |
| 381 | Palacký University Olomouc | 5 | 287 | 646 | 294 |
| 382 | National Autonomous University of Mexico | 5 | 286 | 3111 | 1142 |
| 383 | Nelson Mandela University | 6 | 285 | 293 | 85 |
| 384 | French Institute of Pondicherry | 1 | 284 | 52 | 5 |
| 385 | Leiden University | 8 | 283 | 2305 | 1880 |
| 386 | University of New Mexico | 6 | 281 | 1946 | 1240 |
| 387 | La Trobe University | 5 | 278 | 1749 | 901 |
| 388 | University of Milan | 4 | 276 | 4663 | 2333 |
| 388 | University of Burgundy | 4 | 276 | 1159 | 494 |
| 390 | University of Missouri | 7 | 272 | 3133 | 2116 |
| 390 | Campus university | 2 | 272 | 183 | 36 |
| 392 | Bournemouth University | 4 | 271 | 500 | 184 |
| 392 | Florida Fish and Wildlife Conservation Commission | 7 | 271 | 57 | 37 |
| 394 | Center for International Forestry Research | 2 | 270 | 127 | 62 |
| 395 | Universidade do Porto | 10 | 269 | 628 | 611 |
| 396 | New York University | 1 | 268 | 6249 | 1469 |
| 396 | University of Plymouth | 8 | 268 | 542 | 513 |
| 398 | Charles Darwin University | 5 | 267 | 341 | 176 |
| 398 | Sea Mammal Research Unit | 12 | 267 | 22 | 42 |
| 400 | Loyola University | 1 | 265 | 3662 | 686 |
| 401 | RMIT University | 6 | 263 | 1192 | 730 |
| 402 | Hochschule Koblenz | 2 | 257 | 23 | 2 |
| 403 | New York Botanical Garden | 2 | 253 | 147 | 57 |
| 403 | Missouri Botanical Garden | 2 | 253 | 150 | 71 |
| 405 | University of Birmingham | 7 | 252 | 3000 | 2212 |
| 405 | NatureServe | 1 | 252 | 39 | 16 |
| 405 | Brookfield Zoo | 3 | 252 | 45 | 30 |
| 405 | Royal Holloway University of London | 5 | 252 | 171 | 160 |
| 409 | Edmund Mach Foundation | 6 | 251 | 116 | 108 |
| 409 | National Scientific and Technical Research Council | 6 | 251 | 1053 | 374 |
| 411 | Minnesota Department of Natural Resources | 4 | 250 | 51 | 18 |
| 412 | Research Center for Eco-Environmental Sciences | 3 | 249 | 487 | 278 |
| 413 | Max Planck Institute of Animal Behavior | 8 | 248 | 13 | 8 |
| 414 | Swiss Institute of Bioinformatics | 10 | 247 | 98 | 170 |
| 415 | Vrije Universiteit Brussel | 2 | 246 | 2868 | 926 |
| 416 | Pontificia Universidad Católica del Ecuador | 3 | 245 | 233 | 31 |
| 416 | Lancaster Environment Centre | 14 | 245 | 90 | 209 |
| 418 | University of Neuchâtel | 4 | 244 | 487 | 288 |
| 418 | Kunming Institute of Botany | 2 | 244 | 387 | 143 |
| 420 | Allan Wilson Centre | 2 | 242 | 56 | 32 |
| 421 | Western University | 7 | 239 | 2738 | 2147 |
| 421 | New Mexico State University | 4 | 239 | 877 | 412 |
| 423 | Santa Clara University | 1 | 237 | 1005 | 138 |
| 424 | World Agroforestry Centre | 3 | 233 | 116 | 89 |
| 424 | Museo delle Scienze | 5 | 233 | 19 | 14 |
| 426 | Plymouth Marine Laboratory | 4 | 232 | 139 | 141 |
| 426 | British Antarctic Survey | 3 | 232 | 305 | 196 |
| 428 | University of California | 10 | 231 | 2618 | 2889 |
| 428 | McMaster University | 3 | 231 | 4623 | 2178 |
| 428 | University of Calgary | 6 | 231 | 3317 | 2249 |
| 431 | University of North Carolina at Chapel Hill | 4 | 230 | 3308 | 2065 |
| 432 | Australian Nuclear Science and Technology Organisation | 1 | 229 | 604 | 145 |
| 433 | Technical University of Munich | 11 | 228 | 1928 | 2002 |
| 433 | University of Granada | 3 | 228 | 3222 | 1176 |
| 435 | Wenzhou University | 1 | 227 | 1267 | 200 |
| 436 | California Institute of Technology | 5 | 223 | 1592 | 1334 |
| 436 | University of South Africa | 2 | 223 | 1387 | 194 |
| 438 | Xishuangbanna Tropical Botanical Garden | 5 | 222 | 155 | 109 |
| 439 | University of Erlangen-Nuremberg | 4 | 221 | 2457 | 1398 |
| 439 | AGH University of Science and Technology | 1 | 221 | 1593 | 282 |
| 439 | University of Florence | 6 | 221 | 2368 | 1640 |
| 439 | University of Crete | 1 | 221 | 2675 | 580 |
| 443 | Paris-Saclay University | 8 | 220 | 1367 | 861 |
| 444 | National University of Singapore | 8 | 219 | 3185 | 2776 |
| 444 | University of Liège | 6 | 219 | 1661 | 1322 |
| 444 | Universidad Metropolitana | 8 | 219 | 280 | 155 |
| 447 | Ulm University | 6 | 217 | 1500 | 1162 |
| 447 | Mount Holyoke College | 3 | 217 | 153 | 71 |
| 449 | Kenyon College | 2 | 216 | 136 | 53 |
| 449 | Montana State University | 12 | 216 | 309 | 559 |
| 451 | University of Suffolk | 4 | 215 | 35 | 10 |
| 452 | University of Antofagasta | 1 | 213 | 434 | 54 |
| 453 | Wisconsin Department of Natural Resources | 2 | 212 | 104 | 42 |
| 454 | University of Buenos Aires | 2 | 211 | 4118 | 1064 |
| 454 | University of Twente | 8 | 211 | 1002 | 1046 |
| 454 | Adler Planetarium | 1 | 211 | 23 | 14 |
| 457 | National Physical Laboratory | 4 | 210 | 621 | 370 |
| 458 | University of Manchester | 5 | 209 | 4321 | 2866 |
| 458 | Senckenberg Biodiversity and Climate Research Centre | 6 | 209 | 40 | 48 |
| 458 | University of Hull | 4 | 209 | 1204 | 650 |
| 461 | Central University of Ecuador | 1 | 208 | 406 | 19 |
| 461 | Ashoka Trust for Research in Ecology and the Environment | 1 | 208 | 147 | 29 |
| 463 | Lincoln University | 5 | 207 | 336 | 247 |
| 463 | Natural England | 4 | 207 | 43 | 39 |
| 465 | Eawag | 5 | 206 | 162 | 178 |
| 465 | University of North Texas | 2 | 206 | 3014 | 935 |
| 465 | Lakehead University | 1 | 206 | 1417 | 238 |
| 468 | Sun Yat-sen University | 12 | 205 | 2148 | 2073 |
| 468 | Rocky Mountain Research Station | 6 | 205 | 132 | 129 |
| 470 | Massachusetts Institute of Technology | 5 | 204 | 3071 | 2628 |
| 470 | University of Insubria | 1 | 204 | 1880 | 421 |
| 470 | National Institute of Amazonian Research | 4 | 204 | 267 | 117 |
| 470 | New York State Museum | 2 | 204 | 51 | 18 |
| 474 | University of Texas at Austin | 8 | 202 | 5237 | 4545 |
| 474 | James Madison University | 1 | 202 | 1206 | 157 |
| 474 | University of Miami | 4 | 202 | 3505 | 2153 |
| 474 | Sciences Po | 1 | 202 | 979 | 144 |
| 474 | Madison University | 1 | 202 | 221 | 22 |
| 479 | Ikerbasque | 4 | 201 | 469 | 301 |
| 480 | University of Bologna | 4 | 200 | 4143 | 2333 |
| 480 | Ocean University of China | 1 | 200 | 2215 | 569 |
| 482 | Laboratory of Tree-Ring Research | 1 | 199 | 110 | 57 |
| 483 | Nine Network of Public Media | 5 | 197 | 1412 | 969 |
| 483 | Museu Paraense Emílio Goeldi | 3 | 197 | 163 | 66 |
| 485 | Macaulay Institute | 1 | 196 | 291 | 135 |
| 485 | Forest Research Institute | 3 | 196 | 532 | 245 |
| 485 | Scotland's Rural College | 1 | 196 | 489 | 160 |
| 488 | Pace University | 2 | 194 | 336 | 91 |
| 488 | University of Alcalá | 6 | 194 | 770 | 571 |
| 490 | Kearney | 6 | 193 | 58 | 68 |
| 491 | University of Padua | 4 | 192 | 4096 | 2451 |
| 492 | Polish Academy of Sciences | 6 | 190 | 2471 | 1744 |
| 493 | Technical University of Braunschweig | 4 | 189 | 1105 | 639 |
| 494 | Lancaster University | 11 | 188 | 562 | 1038 |
| 495 | Museum of Vertebrate Zoology | 2 | 186 | 122 | 79 |
| 495 | Universidad del Rosario | 4 | 186 | 235 | 70 |
| 497 | West Virginia University | 4 | 184 | 2297 | 1260 |
| 497 | University of California, San Diego | 13 | 184 | 2409 | 3415 |
| 499 | Delft University of Technology | 4 | 183 | 2479 | 1550 |
| 500 | University of Nottingham | 5 | 182 | 3786 | 2757 |
| 500 | Universidad de los Andes, Chile | 4 | 182 | 989 | 352 |
| 502 | Federal University of Rio Grande do Sul | 6 | 181 | 2389 | 1255 |
| 502 | Universidade Estadual Paulista | 5 | 181 | 2522 | 1098 |
| 504 | Google | 4 | 180 | 328 | 255 |
| 504 | Northern Arizona University | 7 | 180 | 322 | 351 |
| 506 | University of the Aegean | 2 | 179 | 736 | 215 |
| 506 | Government of Western Australia | 2 | 179 | 44 | 22 |
| 509 | University of Victoria | 7 | 177 | 1048 | 1122 |
| 509 | Scottish Association for Marine Science | 5 | 177 | 73 | 94 |
| 511 | University of Nevada, Reno | 5 | 176 | 1205 | 934 |
| 512 | Rhodes University | 3 | 175 | 691 | 312 |
| 513 | Park University | 2 | 174 | 65 | 18 |
| 514 | Manchester Metropolitan University | 6 | 171 | 624 | 570 |
| 515 | Leibniz Centre for Tropical Marine Research | 2 | 170 | 96 | 37 |
| 515 | Virginia Commonwealth University | 2 | 170 | 3900 | 1751 |
| 517 | Paris Diderot University | 7 | 168 | 1575 | 1664 |
| 518 | University of Arkansas | 2 | 167 | 4605 | 1854 |
| 518 | Hopkins Marine Station | 3 | 167 | 117 | 107 |
| 518 | The Open University | 4 | 167 | 1653 | 931 |
| 521 | University of Salento | 1 | 164 | 1914 | 501 |
| 521 | Howard Hughes Medical Institute | 1 | 164 | 1503 | 912 |
| 521 | National University of Córdoba | 2 | 164 | 1697 | 435 |
| 524 | Masaryk University | 5 | 163 | 1088 | 653 |
| 524 | University of Massachusetts | 8 | 163 | 2199 | 2672 |
| 524 | Charles Sturt University | 2 | 163 | 1216 | 411 |
| 527 | University of Portsmouth | 1 | 162 | 2576 | 586 |
| 528 | University of Cologne | 3 | 161 | 3206 | 1755 |
| 528 | Graduate University for Advanced Studies | 3 | 161 | 349 | 214 |
| 530 | National Institute of Polar Research | 2 | 160 | 193 | 83 |
| 531 | Toulouse III University - Paul-Sabatier | 6 | 159 | 139 | 267 |
| 532 | University of Valencia | 5 | 158 | 2494 | 1667 |
| 533 | University of Lincoln | 6 | 157 | 191 | 223 |
| 534 | Oklahoma State University–Stillwater | 4 | 156 | 145 | 88 |
| 534 | Whitman College | 1 | 156 | 229 | 45 |
| 536 | Katholieke Universiteit Leuven | 10 | 155 | 2692 | 3447 |
| 536 | University of Regina | 2 | 155 | 1085 | 382 |
| 536 | Centre Régional d’Innovation et de Transferts Technologiques des Industries du Bois | 3 | 155 | 53 | 31 |
| 539 | University of Innsbruck | 2 | 154 | 2618 | 1228 |
| 539 | University of Picardie Jules Verne | 6 | 154 | 196 | 223 |
| 539 | Bucknell University | 3 | 154 | 305 | 149 |
| 542 | Indiana University | 4 | 153 | 5709 | 3709 |
| 542 | European Commission | 4 | 153 | 867 | 696 |
| 544 | University of Sussex | 4 | 152 | 1941 | 1477 |
| 545 | European Bird Census Council | 1 | 151 | 13 | 10 |
| 546 | Goddard Space Flight Center | 6 | 149 | 385 | 429 |
| 547 | San Francisco State University | 1 | 148 | 1764 | 459 |
| 548 | Swiss Tropical and Public Health Institute | 1 | 147 | 588 | 244 |
| 548 | Rocky Mountain Biological Laboratory | 3 | 147 | 54 | 44 |
| 548 | Robertson Centre for Biostatistics | 1 | 147 | 175 | 73 |
| 551 | University of Bayreuth | 4 | 146 | 1047 | 762 |
| 551 | University of Campinas | 4 | 146 | 3402 | 1791 |
| 553 | Technical University of Madrid | 5 | 145 | 1233 | 866 |
| 553 | University of Ottawa | 6 | 145 | 2841 | 2595 |
| 553 | Norwegian Institute of Marine Research | 8 | 145 | 146 | 215 |
| 553 | Royal Ontario Museum | 2 | 145 | 150 | 72 |
| 557 | University of Agder | 4 | 144 | 346 | 201 |
| 557 | College of Charleston | 3 | 144 | 337 | 180 |
| 559 | Marian University | 2 | 143 | 46 | 4 |
| 559 | Norwegian Research Centre | 4 | 143 | 80 | 48 |
| 561 | Francis Marion University | 1 | 142 | 93 | 10 |
| 561 | Universidade Federal do Pará | 1 | 142 | 2848 | 355 |
| 561 | Columbus State University | 1 | 142 | 197 | 21 |
| 561 | Smithsonian Migratory Bird Center | 4 | 142 | 34 | 38 |
| 561 | Autonomous University of San Luis Potosí | 1 | 142 | 1176 | 172 |
| 566 | University College Dublin | 8 | 141 | 1238 | 1735 |
| 567 | University of Warwick | 5 | 140 | 2432 | 2056 |
| 567 | Boise State University | 5 | 140 | 324 | 294 |
| 567 | Seattle Children's | 3 | 140 | 783 | 536 |
| 570 | Field Museum of Natural History | 8 | 138 | 117 | 163 |
| 570 | Royal Tyrrell Museum | 1 | 138 | 35 | 16 |
| 572 | Southern Cross University | 4 | 137 | 415 | 330 |
| 572 | Marine Institute of Memorial University of Newfoundland | 7 | 137 | 111 | 155 |
| 572 | China National GeneBank | 2 | 137 | 50 | 39 |
| 572 | Gembloux Agro-Bio Tech | 3 | 137 | 249 | 177 |
| 576 | Freie Universität Berlin | 6 | 136 | 1629 | 1641 |
| 576 | King Abdullah University of Science and Technology | 7 | 136 | 536 | 633 |
| 578 | Hebrew University of Jerusalem | 5 | 135 | 3336 | 2847 |
| 578 | University of Gloucestershire | 3 | 135 | 183 | 110 |
| 578 | China Institute | 2 | 135 | 2955 | 1320 |
| 581 | Jagiellonian University | 6 | 133 | 1868 | 1347 |
| 581 | Ministry of Natural Resources and Forestry | 4 | 133 | 47 | 25 |
| 583 | United States Environmental Protection Agency | 3 | 132 | 956 | 796 |
| 584 | Jet Propulsion Laboratory | 3 | 131 | 719 | 565 |
| 584 | University of Naples Federico II | 6 | 131 | 3054 | 2879 |
| 586 | University of Kisangani | 2 | 130 | 36 | 11 |
| 586 | Czech University of Life Sciences Prague | 7 | 130 | 199 | 194 |
| 588 | Fraunhofer Society | 1 | 129 | 1664 | 590 |
| 588 | Algoma University | 1 | 129 | 91 | 32 |
| 588 | Agricultural Research Service | 2 | 129 | 1746 | 1160 |
| 588 | Hill International | 1 | 129 | 195 | 79 |
| 588 | Institut national de l'information géographique et forestière | 2 | 129 | 36 | 28 |
| 588 | Australian Museums and Galleries Association | 5 | 129 | 14 | 37 |
| 594 | Georgetown University | 6 | 128 | 1795 | 1873 |
| 594 | South Australian Research and Development Institute | 3 | 128 | 119 | 92 |
| 596 | San José State University | 4 | 127 | 507 | 368 |
| 596 | Edith Cowan University | 2 | 127 | 1341 | 663 |
| 598 | Rutgers University | 4 | 126 | 4512 | 3409 |
| 599 | Natural Resources Conservation Service | 1 | 125 | 204 | 71 |
| 600 | Karlsruhe Institute of Technology | 2 | 124 | 3115 | 1687 |
| 600 | University Toulouse I - Capitole | 9 | 124 | 1102 | 1574 |
| 600 | Animal and Plant Health Agency | 3 | 124 | 110 | 88 |
| 600 | Agroecology | 4 | 124 | 492 | 394 |
| 604 | University of Tokyo | 12 | 122 | 3395 | 4016 |
| 604 | Ben-Gurion University of the Negev | 6 | 122 | 1994 | 2017 |
| 606 | Emory University | 3 | 121 | 4419 | 3108 |
| 606 | Mississippi State University | 7 | 121 | 787 | 1000 |
| 608 | United Nations | 2 | 120 | 829 | 368 |
| 608 | Silliman University | 1 | 120 | 39 | 17 |
| 608 | Joint Institute for Marine and Atmospheric Research | 1 | 120 | 63 | 41 |
| 608 | South African Institute for Aquatic Biodiversity | 1 | 120 | 127 | 38 |
| 608 | Department of Environment Water and Natural Resources | 1 | 120 | 18 | 3 |
| 613 | University of Koblenz - Landau | 3 | 119 | 172 | 152 |
| 613 | Institute for Environmental Sciences | 3 | 119 | 234 | 214 |
| 615 | National Center for Scientific Research | 3 | 118 | 1430 | 1125 |
| 615 | University of Puerto Rico, Río Piedras Campus | 1 | 118 | 94 | 30 |
| 617 | National Science Foundation | 2 | 117 | 503 | 330 |
| 617 | University of the Witwatersrand | 3 | 117 | 2849 | 1603 |
| 619 | University of Halle-Wittenberg | 1 | 116 | 2240 | 837 |
| 619 | Scripps Institution of Oceanography | 10 | 116 | 294 | 586 |
| 619 | Northern Illinois University | 1 | 116 | 2508 | 733 |
| 619 | WISERD | 1 | 116 | 211 | 60 |
| 623 | University of Oklahoma | 8 | 115 | 1704 | 2422 |
| 623 | Conservation International | 2 | 115 | 143 | 127 |
| 625 | Brigham Young University | 4 | 114 | 1543 | 1355 |
| 626 | Boston College | 1 | 112 | 2698 | 1016 |
| 628 | Technische Universität Darmstadt | 3 | 111 | 1763 | 1137 |
| 628 | United States Fish and Wildlife Service | 11 | 111 | 142 | 253 |
| 628 | Nanjing Agricultural University | 3 | 111 | 1116 | 782 |
| 628 | Te Pūnaha Matatini | 2 | 111 | 24 | 8 |
| 632 | University of Turku | 7 | 108 | 1471 | 1922 |
| 632 | University of Vigo | 6 | 108 | 771 | 849 |
| 632 | OpenAI | 1 | 108 | 8 | 16 |
| 635 | University of Waikato | 2 | 107 | 1428 | 781 |
| 635 | Queensland Brain Institute | 2 | 107 | 281 | 232 |
| 635 | University of Virginia | 4 | 107 | 3867 | 3271 |
| 638 | Institute for Advanced Study | 6 | 106 | 1040 | 1280 |
| 638 | Southern Research | 4 | 106 | 271 | 323 |
| 638 | Università degli Studi del Molise | 1 | 106 | 219 | 81 |
| 641 | University of Bordeaux | 5 | 105 | 2310 | 2227 |
| 641 | Aston University | 1 | 105 | 2405 | 925 |
| 641 | University of Texas at Arlington | 2 | 105 | 1842 | 950 |
| 641 | Universidad Nacional del Comahue | 4 | 105 | 203 | 173 |
| 641 | Azim Premji University | 1 | 105 | 104 | 19 |
| 641 | Wellcome Trust | 2 | 105 | 1043 | 824 |
| 647 | University of Colorado Denver | 5 | 104 | 938 | 1403 |
| 647 | Canadian Wildlife Service | 3 | 104 | 107 | 117 |
| 649 | York University | 7 | 103 | 1315 | 1898 |
| 649 | National Evolutionary Synthesis Center | 2 | 103 | 34 | 42 |
| 651 | Micropolis | 1 | 102 | 6 | 4 |
| 652 | Wellcome Sanger Institute | 1 | 101 | 639 | 401 |
| 652 | Universitat Politècnica de Catalunya | 2 | 101 | 2026 | 1116 |
| 652 | ComUE Paris-Saclay University | 3 | 101 | 1927 | 1701 |
| 655 | China Agricultural University | 4 | 100 | 1342 | 1143 |
| 655 | Naturalis Biodiversity Center | 9 | 100 | 81 | 121 |
| 655 | University of North Carolina at Charlotte | 1 | 100 | 772 | 223 |
| 655 | University of Savoy | 2 | 100 | 572 | 327 |
| 655 | National Museums of Kenya | 1 | 100 | 214 | 102 |
| 655 | Zoo Atlanta | 1 | 100 | 44 | 22 |
| 655 | The Dian Fossey Gorilla Fund International | 1 | 100 | 22 | 11 |
| 662 | University of Lisbon | 8 | 99 | 1805 | 2422 |
| 662 | Mammal Research Institute of the Polish Academy of Sciences | 4 | 99 | 125 | 122 |
| 664 | German Primate Center | 4 | 98 | 99 | 145 |
| 664 | University of the Basque Country | 4 | 98 | 2077 | 1673 |
| 664 | University of Wollongong | 3 | 98 | 2381 | 1829 |
| 667 | Temple University | 2 | 97 | 4057 | 2362 |
| 667 | Nord University | 2 | 97 | 426 | 215 |
| 669 | University of Hawaii | 3 | 96 | 1757 | 1645 |
| 669 | Clemson University | 4 | 96 | 1861 | 1668 |
| 671 | The Rockefeller University | 2 | 95 | 1121 | 950 |
| 671 | PSL University | 14 | 95 | 299 | 781 |
| 671 | Texas State University | 4 | 95 | 755 | 630 |
| 671 | University of San Francisco | 1 | 95 | 965 | 283 |
| 675 | University of Hohenheim | 3 | 94 | 922 | 821 |
| 675 | Jacob Blaustein Institutes for Desert Research | 4 | 94 | 157 | 221 |
| 675 | Marine and Freshwater Research Institute | 1 | 94 | 398 | 180 |
| 675 | Inter-American Tropical Tuna Commission | 1 | 94 | 43 | 33 |
| 679 | University of Leicester | 6 | 92 | 1659 | 2187 |
| 680 | University of Paris 1 Pantheon-Sorbonne | 4 | 91 | 2611 | 2432 |
| 680 | NOVA University Lisbon | 3 | 91 | 1716 | 1285 |
| 682 | United States Air Force Academy | 1 | 90 | 3346 | 1062 |
| 682 | Wright-Patterson Air Force Base | 1 | 90 | 1339 | 635 |
| 682 | Montpellier SupAgro | 2 | 90 | 347 | 256 |
| 685 | University of Giessen | 5 | 88 | 1450 | 1760 |
| 685 | Universiti Brunei Darussalam | 1 | 88 | 842 | 226 |
| 685 | Mendel University in Brno | 2 | 88 | 377 | 193 |
| 685 | Mato Grosso State University | 1 | 88 | 566 | 65 |
| 685 | University of Côte d'Azur | 1 | 88 | 2924 | 1208 |
| 685 | Universidad Autónoma del Beni | 1 | 88 | 18 | 11 |
| 685 | National Agency for National Parks | 3 | 88 | 10 | 15 |
| 685 | Royal Museum for Central Africa | 1 | 88 | 244 | 138 |
| 685 | Universidad Nacional de San Antonio Abad del Cusco | 2 | 88 | 77 | 20 |
| 694 | Southern Illinois University Carbondale | 4 | 87 | 1754 | 1742 |
| 694 | University of Puerto Rico | 3 | 87 | 1045 | 912 |
| 694 | Emerging Pathogens Institute | 5 | 87 | 84 | 171 |
| 697 | Bavarian Forest National Park | 7 | 86 | 17 | 47 |
| 697 | Botanischer Garten und Botanisches Museum Berlin | 1 | 86 | 62 | 31 |
| 699 | Parks Canada | 1 | 85 | 152 | 56 |
| 699 | National Fund for Scientific Research | 1 | 85 | 374 | 216 |
| 699 | African Institute for Mathematical Sciences | 4 | 85 | 33 | 45 |
| 699 | Wellcome Centre for Human Genetics | 1 | 85 | 510 | 390 |
| 699 | Ministry for Primary Industries | 1 | 85 | 98 | 40 |
| 704 | University of Málaga | 2 | 84 | 2309 | 1192 |
| 704 | Kenya Wildlife Service | 2 | 84 | 31 | 28 |
| 704 | Middlesex University | 2 | 84 | 878 | 507 |
| 707 | Rothamsted Research | 4 | 83 | 375 | 500 |
| 707 | Great Lakes Science Center | 4 | 83 | 47 | 58 |
| 709 | Nicolaus Copernicus University in Toruń | 4 | 82 | 981 | 718 |
| 709 | Museo Nacional de Ciencias Naturales | 5 | 82 | 198 | 264 |
| 709 | The Institute of Conservation | 5 | 82 | 48 | 79 |
| 709 | South African Environmental Observation Network | 2 | 82 | 57 | 43 |
| 709 | University of the West of England | 3 | 82 | 914 | 795 |
| 714 | Johns Hopkins University | 3 | 81 | 6010 | 4871 |
| 714 | Schiller International University | 2 | 81 | 255 | 273 |
| 714 | Bjerknes Centre for Climate Research | 2 | 81 | 145 | 119 |
| 714 | Institute for Genetics, Environment and Plant Protection | 1 | 81 | 7 | 7 |
| 714 | International Institute for Applied Systems Analysis | 7 | 81 | 142 | 274 |
| 719 | Adam Mickiewicz University in Poznań | 4 | 80 | 1173 | 902 |
| 719 | Polytechnic University of Milan | 2 | 80 | 2931 | 1819 |
| 719 | University of Porto | 3 | 80 | 2442 | 1954 |
| 723 | National Taiwan University | 8 | 79 | 2548 | 3409 |
| 723 | École polytechnique | 3 | 79 | 1644 | 1486 |
| 723 | University of Haifa | 5 | 79 | 1257 | 1494 |
| 726 | University of Tampere | 2 | 78 | 2674 | 1652 |
| 726 | Christopher Newport University | 2 | 78 | 107 | 75 |
| 726 | Tanzania Wildlife Research Institute | 2 | 78 | 40 | 38 |
| 726 | Instituto Multidisciplinario de Biología Vegetal | 1 | 78 | 240 | 116 |
| 726 | Royal Belgian Institute of Natural Sciences | 1 | 78 | 331 | 165 |
| 731 | Federal University of Rio de Janeiro | 2 | 77 | 5178 | 2508 |
| 731 | Royal Saskatchewan Museum | 1 | 77 | 26 | 8 |
| 733 | Kazan Federal University | 2 | 76 | 1181 | 527 |
| 733 | Graz University of Technology | 1 | 76 | 1968 | 1018 |
| 733 | Memorial University of Newfoundland | 4 | 76 | 1573 | 1667 |
| 736 | Heriot-Watt University | 1 | 75 | 2643 | 1218 |
| 736 | Swedish Environmental Protection Agency | 1 | 75 | 34 | 32 |
| 738 | University of Lleida | 2 | 74 | 450 | 397 |
| 738 | Marie Curie | 2 | 74 | 2896 | 2302 |
| 738 | New England Aquarium | 2 | 74 | 55 | 54 |
| 738 | Swiss National Park | 1 | 74 | 18 | 5 |
| 742 | Florida State University | 3 | 73 | 3144 | 2711 |
| 742 | Universidad de Antioquia | 3 | 73 | 1175 | 730 |
| 742 | Bio21 Institute | 1 | 73 | 325 | 230 |
| 742 | National Wetlands Research Center | 3 | 73 | 33 | 68 |
| 746 | Engineering Institute | 2 | 72 | 1894 | 1051 |
| 746 | Université de Perpignan | 1 | 72 | 533 | 306 |
| 748 | Durrell Wildlife Conservation Trust | 3 | 71 | 23 | 38 |
| 748 | The Climate Group | 1 | 71 | 119 | 87 |
| 750 | National University of Colombia | 2 | 70 | 2237 | 940 |
| 750 | German Center for Lung Research | 1 | 70 | 514 | 272 |
| 750 | Federal University of Lavras | 3 | 70 | 813 | 506 |
| 750 | Evolution Institute | 3 | 70 | 34 | 38 |
| 750 | Universidad de Castilla la Mancha | 2 | 70 | 860 | 585 |
| 755 | Oxford Brookes University | 4 | 69 | 601 | 820 |
| 755 | Museums Victoria | 2 | 69 | 103 | 78 |
| 755 | Linköping University | 5 | 69 | 1772 | 2517 |
| 755 | Clermont Auvergne University | 1 | 69 | 2413 | 1077 |
| 755 | Information Technology University | 3 | 69 | 97 | 95 |
| 760 | Tianjin Normal University | 1 | 68 | 849 | 294 |
| 761 | National Ecological Observatory Network | 8 | 66 | 5 | 43 |
| 761 | South China Normal University | 1 | 66 | 1950 | 909 |
| 761 | San Diego Natural History Museum | 1 | 66 | 54 | 26 |
| 761 | Cancer Research Center | 2 | 66 | 2536 | 2038 |
| 765 | Aberystwyth University | 3 | 65 | 707 | 753 |
| 765 | Beijing Normal University | 4 | 65 | 1876 | 1807 |
| 767 | TU Dresden | 8 | 64 | 1617 | 2783 |
| 767 | Medical University of Vienna | 1 | 64 | 3064 | 1897 |
| 767 | Tulane University | 3 | 64 | 2206 | 2185 |
| 767 | Naval Facilities Engineering Command | 1 | 64 | 27 | 20 |
| 767 | Her Majesty's Coastguard | 1 | 64 | 130 | 34 |
| 767 | United States Coast Guard | 1 | 64 | 20 | 13 |
| 767 | Escuela Politécnica Nacional | 1 | 64 | 360 | 128 |
| 774 | Alaska Department of Fish and Game | 4 | 63 | 63 | 105 |
| 775 | Pontifical Catholic University of Chile | 2 | 62 | 2875 | 1862 |
| 775 | University of Siena | 3 | 62 | 1771 | 1852 |
| 775 | Southwest Research Institute | 2 | 62 | 543 | 475 |
| 778 | University of Concepción | 2 | 61 | 1555 | 973 |
| 778 | University of Rennes 1 | 4 | 61 | 1289 | 1659 |
| 778 | Helmholtz-Zentrum Dresden-Rossendorf | 2 | 61 | 597 | 455 |
| 778 | University of Ferrara | 1 | 61 | 3200 | 1813 |
| 782 | Cornell Lab of Ornithology | 2 | 60 | 36 | 50 |
| 782 | State University of New York at Purchase | 2 | 60 | 214 | 215 |
| 782 | National Institute for Environmental Studies | 6 | 60 | 358 | 636 |
| 785 | AgResearch | 2 | 59 | 519 | 558 |
| 785 | University of Oldenburg | 4 | 59 | 690 | 971 |
| 785 | University of Puget Sound | 1 | 59 | 291 | 133 |
| 785 | University of Colorado | 5 | 59 | 3858 | 4921 |
| 785 | Public Health Institute Ostrava | 1 | 59 | 7067 | 3792 |
| 790 | Lawrence Berkeley National Laboratory | 3 | 58 | 1639 | 1874 |
| 790 | TUM School of Life Sciences | 5 | 58 | 92 | 205 |
| 790 | PRBO Conservation Science | 1 | 58 | 39 | 41 |
| 790 | autonomous university | 2 | 58 | 511 | 415 |
| 794 | Bushfire and Natural Hazards CRC | 1 | 57 | 47 | 41 |
| 794 | Industrial University of Santander | 1 | 57 | 887 | 253 |
| 794 | Ohio Department of Natural Resources | 1 | 57 | 44 | 26 |
| 797 | National Institutes of Health | 5 | 56 | 2840 | 3652 |
| 797 | Auckland University of Technology | 1 | 56 | 1853 | 881 |
| 797 | State University of New York | 3 | 56 | 4999 | 4742 |
| 797 | Plant & Food Research | 1 | 56 | 549 | 409 |
| 797 | Idaho State University | 4 | 56 | 313 | 557 |
| 797 | Seattle City Light | 1 | 56 | 9 | 1 |
| 803 | Kyoto University | 6 | 55 | 4014 | 4532 |
| 804 | University of Hawaiʻi at Hilo | 3 | 54 | 116 | 250 |
| 804 | The School for Field Studies | 1 | 54 | 61 | 35 |
| 804 | University of Belize | 1 | 54 | 15 | 7 |
| 807 | Federal University of Minas Gerais | 1 | 53 | 5955 | 2551 |
| 807 | 23andMe | 2 | 53 | 39 | 64 |
| 807 | University of Pretoria | 4 | 53 | 1917 | 2151 |
| 807 | Information Technologies Institute | 1 | 53 | 185 | 88 |
| 807 | Informatics Philippines | 1 | 53 | 778 | 444 |
| 807 | United States Army Corps of Engineers | 1 | 53 | 424 | 228 |
| 813 | Georgia Institute of Technology | 2 | 52 | 3989 | 3431 |
| 813 | University of Waterloo | 2 | 52 | 4569 | 3695 |
| 813 | Okinawa Institute of Science and Technology | 7 | 52 | 118 | 341 |
| 813 | Santos | 2 | 52 | 1359 | 929 |
| 813 | Natural Environment Research Council | 1 | 52 | 393 | 348 |
| 818 | National University of Malaysia | 1 | 51 | 3727 | 1653 |
| 818 | University of Trieste | 2 | 51 | 2201 | 1869 |
| 818 | Universidad Veracruzana | 2 | 51 | 841 | 395 |
| 821 | University of Marburg | 5 | 50 | 1412 | 2304 |
| 821 | University of Jaén | 1 | 50 | 1793 | 912 |
| 821 | National University of Misiones | 1 | 50 | 210 | 76 |
| 821 | University of Abertay Dundee | 3 | 50 | 136 | 252 |
| 821 | Quaid-i-Azam University | 1 | 50 | 1886 | 1111 |
| 821 | Sichuan University | 3 | 50 | 4191 | 3571 |
| 821 | University of Massachusetts Dartmouth | 1 | 50 | 746 | 464 |
| 821 | Universidade Federal da Integração Latino-Americana | 1 | 50 | 255 | 23 |
| 821 | Instituto de Biología Subtropical | 1 | 50 | 84 | 32 |
| 821 | The Peregrine Fund | 3 | 50 | 21 | 39 |
| 821 | University of Texas Rio Grande Valley | 2 | 50 | 1067 | 860 |
| 821 | Western Norway University of Applied Sciences | 2 | 50 | 447 | 333 |
| 833 | National Institute of Technology | 1 | 49 | 3210 | 1376 |
| 833 | Fisheries Agency | 1 | 49 | 40 | 34 |
| 833 | Global Wildlife Conservation | 1 | 49 | 31 | 27 |
| 833 | Big Data Institute | 3 | 49 | 123 | 188 |
| 833 | York College | 1 | 49 | 556 | 268 |
| 838 | University of La Laguna | 2 | 48 | 1549 | 1190 |
| 838 | University of Prince Edward Island | 1 | 48 | 708 | 397 |
| 838 | Office of Naval Research | 1 | 48 | 239 | 170 |
| 838 | Environmental Defense Fund | 1 | 48 | 160 | 120 |
| 838 | Monterey Bay Aquarium Research Institute | 1 | 48 | 188 | 175 |
| 838 | Atlantic Veterinary College | 1 | 48 | 251 | 176 |
| 838 | Rosenstiel School of Marine and Atmospheric Science | 1 | 48 | 421 | 368 |
| 838 | Julius Kühn-Institut | 1 | 48 | 331 | 233 |
| 838 | Laboratoire Dynamiques Sociales et Recomposition des Espaces | 1 | 48 | 3 | 4 |
| 838 | Bat Conservation Trust | 1 | 48 | 11 | 14 |
| 838 | Ocean Tracking Network | 1 | 48 | 16 | 14 |
| 838 | Atlas of Living Australia | 1 | 48 | 9 | 13 |
| 850 | University of Salzburg | 2 | 47 | 1178 | 1128 |
| 850 | University of Northern British Columbia | 2 | 47 | 445 | 489 |
| 850 | Hungarian University of Agriculture and Life Sciences | 1 | 47 | 865 | 456 |
| 850 | National Polytechnic Institute | 1 | 47 | 3199 | 1533 |
| 850 | Avignon University | 3 | 47 | 195 | 311 |
| 850 | Marine Scotland | 3 | 47 | 117 | 191 |
| 850 | Qingdao Agricultural University | 2 | 47 | 626 | 502 |
| 850 | Lincoln Park Zoo | 3 | 47 | 34 | 60 |
| 850 | Spanish National Research Council | 2 | 47 | 2376 | 2189 |
| 850 | Universidad de Córdoba | 2 | 47 | 1203 | 847 |
| 860 | University of San Diego | 1 | 46 | 813 | 457 |
| 860 | Radford University | 1 | 46 | 369 | 186 |
| 860 | China University of Geosciences | 1 | 46 | 2176 | 1342 |
| 860 | University of Seville | 2 | 46 | 3720 | 2628 |
| 860 | Bogor Agricultural University | 1 | 46 | 643 | 303 |
| 860 | Indian Statistical Institute | 1 | 46 | 1246 | 649 |
| 860 | California Department of Fish and Wildlife | 1 | 46 | 161 | 125 |
| 860 | College of Staten Island | 1 | 46 | 642 | 399 |
| 860 | Indian Statistical Institute Bangalore | 1 | 46 | 8 | 3 |
| 869 | University of Salamanca | 2 | 45 | 2268 | 1689 |
| 869 | Free University of Bozen-Bolzano | 2 | 45 | 520 | 519 |
| 869 | University of Bari | 1 | 45 | 4032 | 2748 |
| 869 | Universitat de Lleida | 1 | 45 | 740 | 446 |
| 873 | College of Science and Technology | 1 | 44 | 2302 | 1123 |
| 873 | Geoscience Australia | 2 | 44 | 168 | 189 |
| 873 | Oeschger Centre for Climate Change Research | 1 | 44 | 226 | 187 |
| 873 | Chapman University | 1 | 44 | 959 | 569 |
| 877 | Austral University of Chile | 2 | 43 | 891 | 739 |
| 877 | Minnesota Zoo | 1 | 43 | 15 | 12 |
| 877 | Morton Arboretum | 3 | 43 | 44 | 74 |
| 877 | Federal University of Uberlândia | 1 | 43 | 2695 | 957 |
| 877 | Japan Science and Technology Agency | 2 | 43 | 1349 | 1444 |
| 882 | Auburn University | 4 | 42 | 2138 | 2790 |
| 882 | Tokyo Tokyo | 6 | 42 | 254 | 418 |
| 884 | Hokkaido University | 2 | 41 | 4773 | 3842 |
| 884 | George Mason University | 2 | 41 | 2848 | 2536 |
| 884 | Hatfield Marine Science Center | 2 | 41 | 98 | 153 |
| 884 | Sant'Anna School of Advanced Studies | 1 | 41 | 1033 | 769 |
| 888 | University of Pittsburgh | 3 | 40 | 5984 | 6160 |
| 888 | California Academy of Sciences | 2 | 40 | 177 | 192 |
| 888 | Japan Agency for Marine-Earth Science and Technology | 3 | 40 | 403 | 465 |
| 888 | Flanders Marine Institute | 3 | 40 | 11 | 26 |
| 888 | Moss Landing Marine Laboratories | 2 | 40 | 106 | 145 |
| 888 | Garvan Institute of Medical Research | 1 | 40 | 660 | 606 |
| 888 | University of South Alabama | 2 | 40 | 1154 | 1251 |
| 888 | Dauphin Island Sea Lab | 2 | 40 | 77 | 92 |
| 897 | Indian Institute of Science | 2 | 39 | 2518 | 2124 |
| 897 | Northwest University | 2 | 39 | 1595 | 1267 |
| 897 | Higher University of San Andrés | 1 | 39 | 297 | 191 |
| 897 | Capital Normal University | 2 | 39 | 914 | 769 |
| 897 | Bigelow Laboratory for Ocean Sciences | 2 | 39 | 104 | 155 |
| 897 | State Key Laboratory of Remote Sensing Science | 3 | 39 | 127 | 195 |
| 897 | Poznań University of Life Sciences | 2 | 39 | 502 | 469 |
| 904 | University of Lyon | 1 | 38 | 1492 | 1118 |
| 904 | Russian Academy of Sciences | 1 | 38 | 7428 | 4247 |
| 904 | University of Łódź | 1 | 38 | 1563 | 839 |
| 904 | University of Coimbra | 1 | 38 | 4721 | 2755 |
| 904 | Rzeszów University | 1 | 38 | 180 | 82 |
| 904 | SLAC National Accelerator Laboratory | 1 | 38 | 533 | 438 |
| 904 | Åbo Akademi University | 1 | 38 | 1952 | 1385 |
| 904 | Fort Lewis College | 1 | 38 | 142 | 98 |
| 904 | University of Palermo | 1 | 38 | 4074 | 2820 |
| 904 | Life Sciences Institute | 1 | 38 | 842 | 737 |
| 904 | Wildlife Alliance | 1 | 38 | 57 | 16 |
| 904 | Siedlce University of Natural Sciences and Humanities | 1 | 38 | 232 | 61 |
| 904 | State University of Santa Cruz | 1 | 38 | 935 | 322 |
| 904 | Douglas | 3 | 38 | 773 | 1260 |
| 904 | Museum and Institute of Zoology | 1 | 38 | 171 | 97 |
| 904 | Mathematical Institute, University of Oxford | 4 | 38 | 763 | 967 |
| 904 | Northland College | 1 | 38 | 34 | 34 |
| 904 | Federal Institute For Materials Research and Testing | 1 | 38 | 744 | 592 |
| 922 | National Centre for Biological Sciences | 1 | 37 | 354 | 299 |
| 922 | University of Oregon | 3 | 37 | 2066 | 2611 |
| 922 | Maastricht University | 1 | 37 | 4822 | 3551 |
| 922 | University of Strathclyde | 3 | 37 | 2455 | 3053 |
| 922 | SASTRA University | 1 | 37 | 526 | 376 |
| 922 | Swedish Museum of Natural History | 3 | 37 | 233 | 334 |
| 928 | Florida Atlantic University | 2 | 36 | 1871 | 1822 |
| 928 | National and Kapodistrian University of Athens | 1 | 36 | 6481 | 4363 |
| 928 | University of International Business and Economics | 1 | 36 | 1152 | 644 |
| 928 | Athens University of Economics and Business | 1 | 36 | 630 | 449 |
| 928 | Government of Andalusia | 1 | 36 | 454 | 371 |
| 928 | Dutch Butterfly Conservation | 1 | 36 | 15 | 18 |
| 928 | Species360 | 1 | 36 | 10 | 12 |
| 928 | Atlantic University | 2 | 36 | 326 | 391 |
| 928 | Universidade Federal do Amazonas | 2 | 36 | 915 | 504 |
| 928 | Harbor Branch Oceanographic Institute | 1 | 36 | 184 | 172 |
| 928 | Prevention Institute | 3 | 36 | 204 | 446 |
| 928 | Temple College | 1 | 36 | 52 | 31 |
| 940 | Laboratory of Molecular Biology | 1 | 35 | 1849 | 1536 |
| 940 | National Chung Hsing University | 4 | 35 | 1129 | 1716 |
| 940 | University of Extremadura | 1 | 35 | 2414 | 1486 |
| 940 | Brock University | 1 | 35 | 2007 | 1367 |
| 940 | Northwest A&F University | 1 | 35 | 1925 | 1383 |
| 940 | East Malling Research Station | 1 | 35 | 102 | 107 |
| 940 | Natural Resources Wales | 1 | 35 | 39 | 36 |
| 940 | Mauritian Wildlife Foundation | 2 | 35 | 13 | 26 |
| 940 | Senckenberg Natural History Collections Dresden | 1 | 35 | 75 | 49 |
| 949 | Technical University of Berlin | 3 | 34 | 2086 | 2631 |
| 949 | University at Buffalo | 4 | 34 | 3190 | 4504 |
| 949 | Senckenberg Nature Research Society | 3 | 34 | 33 | 103 |
| 949 | University of Fukui | 1 | 34 | 1522 | 928 |
| 949 | University of Wisconsin–Milwaukee | 1 | 34 | 3228 | 2336 |
| 949 | Teagasc | 2 | 34 | 439 | 596 |
| 949 | Ocean Networks Canada | 1 | 34 | 25 | 26 |
| 949 | Gulf of Maine Research Institute | 1 | 34 | 39 | 43 |
| 957 | University of Aveiro | 1 | 33 | 3378 | 2306 |
| 957 | Max Planck Institute for Evolutionary Biology | 4 | 33 | 64 | 169 |
| 957 | Earlham Institute | 2 | 33 | 80 | 128 |
| 957 | Wyss Institute for Biologically Inspired Engineering | 1 | 33 | 263 | 309 |
| 957 | Institut Pasteur de la Guyane | 1 | 33 | 85 | 74 |
| 957 | Tall Timbers Research Station and Land Conservancy | 1 | 33 | 61 | 50 |
| 957 | Data61 | 3 | 33 | 152 | 273 |
| 964 | Stazione Zoologica Anton Dohrn | 3 | 32 | 244 | 440 |
| 964 | University of Bremen | 1 | 32 | 3201 | 2361 |
| 964 | International Institute of Tropical Agriculture | 1 | 32 | 463 | 385 |
| 964 | University of Colorado Museum of Natural History | 1 | 32 | 25 | 30 |
| 964 | Jambi University | 1 | 32 | 1048 | 78 |
| 964 | Telethon Institute for Child Health Research | 1 | 32 | 666 | 587 |
| 964 | McDonald Institute for Archaeological Research | 1 | 32 | 94 | 86 |
| 964 | Perth Children's Hospital | 1 | 32 | 443 | 354 |
| 973 | Taras Shevchenko National University of Kyiv | 1 | 31 | 1628 | 758 |
| 973 | Institute for Advanced Study Berlin | 1 | 31 | 59 | 81 |
| 973 | Born Free Foundation | 1 | 31 | 8 | 7 |
| 973 | Instituto Antártico Argentino | 1 | 31 | 108 | 105 |
| 973 | University of Delaware | 1 | 31 | 4703 | 3653 |
| 973 | London School of Hygiene & Tropical Medicine | 1 | 31 | 2238 | 1862 |
| 973 | Delaware State University | 1 | 31 | 306 | 246 |
| 973 | Sheba Medical Center | 1 | 31 | 1997 | 1651 |
| 973 | Agrocampus Ouest | 2 | 31 | 252 | 385 |
| 983 | University of Bonn | 4 | 30 | 2752 | 3998 |
| 983 | Drexel University | 3 | 30 | 2376 | 3376 |
| 983 | Bar-Ilan University | 2 | 30 | 2646 | 2629 |
| 983 | Tokyo University of Marine Science and Technology | 2 | 30 | 375 | 373 |
| 983 | BirdLife Australia | 1 | 30 | 19 | 17 |
| 983 | The Institute for Bird Populations | 1 | 30 | 21 | 23 |
| 983 | Old Road Campus | 2 | 30 | 653 | 926 |
| 983 | Universitat Jaume I | 2 | 30 | 973 | 1070 |
| 991 | University of Rostock | 1 | 29 | 2975 | 2313 |
| 991 | Brooklyn College | 2 | 29 | 714 | 875 |
| 991 | University of Genoa | 2 | 29 | 3560 | 3591 |
| 991 | University of Southern Mississippi | 1 | 29 | 1728 | 1303 |
| 991 | Ball State University | 1 | 29 | 1500 | 1003 |
| 991 | Pacific Northwest National Laboratory | 2 | 29 | 1181 | 1388 |
| 991 | Northumbria University | 1 | 29 | 2727 | 1923 |
| 991 | University of Worcester | 1 | 29 | 407 | 306 |
| 999 | Hungarian Academy of Sciences | 3 | 28 | 1867 | 2718 |
| 999 | Australian Defence Force Academy | 1 | 28 | 366 | 380 |