| # | Institution | Articles | Citations | R ARank | R CRank |
|---|
| 1 | University of Tokyo | 334 | 15,096 | 114 | 83 |
| 2 | Kyoto University | 262 | 13,361 | 109 | 70 |
| 3 | Osaka University | 244 | 10,656 | 90 | 61 |
| 4 | University of Tsukuba | 83 | 5,445 | 119 | 31 |
| 5 | Nara Institute of Science and Technology | 71 | 4,050 | 15 | 12 |
| 6 | Kyushu University | 112 | 3,954 | 173 | 128 |
| 7 | National Institute of Genetics | 71 | 3,895 | 13 | 17 |
| 8 | Nagoya University | 95 | 3,800 | 159 | 91 |
| 9 | Kumamoto University | 69 | 3,733 | 42 | 17 |
| 10 | Kobe University | 66 | 3,547 | 63 | 25 |
| 11 | Tohoku University | 75 | 3,128 | 314 | 184 |
| 12 | Tokyo Institute of Technology | 66 | 2,981 | 277 | 127 |
| 13 | Japan Science and Technology Agency | 45 | 2,927 | 32 | 37 |
| 14 | Hokkaido University | 63 | 2,732 | 288 | 149 |
| 15 | National Institute for Basic Biology, Japan | 39 | 2,708 | 17 | 25 |
| 16 | Tokyo Medical and Dental University | 49 | 2,382 | 100 | 70 |
| 17 | RIKEN | 54 | 1,934 | 61 | 74 |
| 18 | Hiroshima University | 64 | 1,902 | 140 | 103 |
| 19 | Hyogo College of Medicine | 10 | 1,235 | 239 | 34 |
| 20 | Yokohama City University | 29 | 1,217 | 49 | 34 |
| 21 | Waseda University | 20 | 1,199 | 347 | 84 |
| 22 | University of York | 2 | 921 | 2.9K | 371 |
| 23 | Osaka Medical College | 9 | 804 | 204 | 26 |
| 24 | Tokyo University of Science | 35 | 749 | 177 | 150 |
| 25 | Howard Hughes Medical Institute | 10 | 724 | 379 | 373 |
| 26 | Chiba University | 16 | 701 | 555 | 277 |
| 27 | Keio University | 14 | 672 | 657 | 309 |
| 28 | Kanazawa University | 22 | 670 | 317 | 224 |
| 29 | Niigata University | 17 | 665 | 283 | 145 |
| 30 | Teikyo University | 6 | 652 | 230 | 50 |
| 31 | Ehime University | 12 | 644 | 272 | 96 |
| 32 | Okayama University | 12 | 642 | 566 | 233 |
| 33 | National Institutes of Health | 14 | 583 | 1.6K | 1.5K |
| 34 | Osaka City University | 12 | 548 | 307 | 130 |
| 35 | Nagoya City University | 16 | 536 | 102 | 84 |
| 36 | Kansai Medical University | 10 | 495 | 207 | 102 |
| 37 | Tokyo University of Agriculture | 8 | 490 | 157 | 54 |
| 38 | Tokyo University of Pharmacy and Life Sciences | 13 | 487 | 98 | 72 |
| 39 | Harvard University | 6 | 484 | 2.9K | 1.8K |
| 40 | University of Central Florida | 3 | 477 | 2.0K | 393 |
| 41 | National Center for Scientific Research | 13 | 476 | 3.2K | 2.3K |
| 42 | Tokai University | 14 | 469 | 205 | 93 |
| 43 | Shizuoka University | 6 | 457 | 561 | 140 |
| 44 | Fujita Health University | 8 | 455 | 171 | 74 |
| 45 | Kurume University | 4 | 449 | 224 | 63 |
| 46 | University of Occupational and Environmental Health Japan | 10 | 436 | 229 | 124 |
| 47 | Robert Wood Johnson Medical School | 9 | 422 | 200 | 188 |
| 47 | Kanagawa Dental College | 4 | 422 | 73 | 13 |
| 49 | National Institute of Information and Communications Technology | 12 | 403 | 76 | 62 |
| 50 | Scripps Research | 2 | 388 | 1.1K | 513 |
| 51 | Kindai University | 18 | 386 | 209 | 205 |
| 52 | University of Chicago | 3 | 381 | 4.2K | 1.7K |
| 53 | University of Cambridge | 3 | 374 | 5.9K | 2.5K |
| 54 | University of Hyogo | 10 | 370 | 250 | 119 |
| 55 | Japanese Foundation For Cancer Research | 7 | 360 | 86 | 85 |
| 56 | University of Toyama | 15 | 359 | 334 | 331 |
| 57 | University of California | 10 | 354 | 8.1K | 6.2K |
| 58 | RIKEN Brain Science Institute | 12 | 346 | 43 | 96 |
| 59 | California Institute of Technology | 1 | 328 | 3.0K | 1.1K |
| 60 | National Institute of Neurosciences & Hospital | 11 | 323 | 78 | 119 |
| 61 | Cancer Research UK | 5 | 320 | 200 | 193 |
| 62 | St. Vincent's Institute of Medical Research | 1 | 319 | 133 | 47 |
| 63 | Hosei University | 10 | 316 | 110 | 47 |
| 64 | Juntendo University | 5 | 310 | 479 | 182 |
| 65 | Gifu University | 6 | 308 | 640 | 248 |
| 66 | Tokyo Metropolitan University | 14 | 306 | 326 | 272 |
| 67 | Northwestern University | 6 | 304 | 3.0K | 2.2K |
| 67 | Japan Women's University | 5 | 304 | 37 | 13 |
| 67 | Showa University | 2 | 304 | 698 | 156 |
| 70 | Eunice Kennedy Shriver National Institute of Child Health and Human Development | 6 | 295 | 451 | 463 |
| 71 | Kagoshima University | 5 | 289 | 858 | 292 |
| 72 | Kyoto Institute of Technology | 16 | 282 | 156 | 150 |
| 72 | Michigan Medicine | 8 | 282 | 1.0K | 959 |
| 74 | The Rockefeller University | 7 | 281 | 440 | 553 |
| 75 | Massachusetts General Hospital | 1 | 274 | 4.5K | 1.7K |
| 75 | Osaka Bioscience Institute | 7 | 274 | 25 | 49 |
| 75 | The Graduate University for Advanced Studies | 11 | 274 | 54 | 80 |
| 78 | Hungarian Academy of Sciences | 2 | 271 | 2.2K | 733 |
| 79 | University of Basel | 2 | 265 | 2.7K | 1.1K |
| 79 | National Institute of Advanced Industrial Science and Technology | 10 | 265 | 423 | 393 |
| 81 | Okinawa Institute of Science and Technology | 8 | 264 | 55 | 50 |
| 82 | University of Tokushima | 8 | 263 | 357 | 248 |
| 83 | Gunma University | 12 | 261 | 285 | 257 |
| 84 | Nara Medical University | 1 | 258 | 1.2K | 186 |
| 85 | Matsumoto Dental University | 2 | 253 | 85 | 22 |
| 86 | National Institute of Radiological Sciences | 2 | 252 | 462 | 141 |
| 87 | Biozentrum University of Basel | 1 | 250 | 107 | 52 |
| 87 | Setsunan University | 3 | 250 | 175 | 40 |
| 89 | Washington University School of Medicine | 3 | 247 | 2.4K | 1.4K |
| 90 | University of Miyazaki | 8 | 246 | 372 | 265 |
| 90 | Tottori University | 5 | 246 | 656 | 301 |
| 92 | Peking University | 2 | 245 | 4.3K | 1.7K |
| 92 | Graduate University for Advanced Studies | 4 | 245 | 43 | 31 |
| 94 | Kyoto Prefectural University of Medicine | 9 | 244 | 412 | 384 |
| 95 | Kitasato University | 7 | 243 | 417 | 259 |
| 96 | Mie University | 5 | 239 | 475 | 216 |
| 97 | National University of Singapore | 5 | 238 | 3.6K | 2.4K |
| 98 | Nagasaki University | 10 | 237 | 359 | 334 |
| 99 | Doshisha University | 8 | 235 | 196 | 97 |
| 99 | National Institute of Health Sciences | 10 | 235 | 107 | 130 |
| 101 | University of Würzburg | 4 | 230 | 1.7K | 1.2K |
| 102 | National Institutes of Natural Sciences, Japan | 11 | 226 | 53 | 93 |
| 103 | Fukushima Medical University | 9 | 225 | 178 | 150 |
| 103 | Indian Institute of Science | 6 | 225 | 1.1K | 739 |
| 105 | University of Utah | 2 | 223 | 5.1K | 2.2K |
| 106 | Jichi Medical University | 3 | 221 | 613 | 206 |
| 107 | Yamaguchi University | 11 | 210 | 299 | 292 |
| 108 | Ritsumeikan University | 10 | 209 | 255 | 193 |
| 109 | University of Patras | 1 | 208 | 3.0K | 629 |
| 109 | Albert Einstein College of Medicine | 4 | 208 | 1.5K | 1.1K |
| 111 | Memorial Sloan Kettering Cancer Center | 1 | 207 | 2.4K | 985 |
| 111 | Weill Cornell Medicine | 1 | 207 | 3.3K | 1.3K |
| 113 | Baylor College of Medicine | 3 | 200 | 2.8K | 1.7K |
| 113 | Research Organization of Information and Systems | 5 | 200 | 9 | 21 |
| 113 | Cancer Research Institute | 11 | 200 | 120 | 232 |
| 116 | Kihara Institute for Biological Research | 5 | 197 | 33 | 36 |
| 117 | French National Institute of Health and Medical Research | 3 | 195 | 2.0K | 1.2K |
| 118 | Umeå University | 2 | 190 | 2.9K | 1.3K |
| 119 | NYU Langone Medical Center | 1 | 189 | 1.7K | 619 |
| 120 | Gakushuin University | 8 | 185 | 77 | 71 |
| 121 | University of Edinburgh | 3 | 183 | 4.9K | 2.7K |
| 122 | London Research Institute | 4 | 179 | 68 | 108 |
| 123 | National Institute of Mental Health | 2 | 177 | 991 | 640 |
| 124 | National University Corporation Tsukuba University of Technology | 1 | 175 | 58 | 2 |
| 124 | Hokuriku University | 1 | 175 | 211 | 65 |
| 126 | Meiji University | 3 | 172 | 495 | 109 |
| 127 | Yokohama National University | 4 | 170 | 660 | 312 |
| 128 | Nihon University | 12 | 166 | 537 | 625 |
| 129 | Shiga University of Medical Science | 4 | 163 | 514 | 330 |
| 130 | University of Adelaide | 2 | 160 | 4.2K | 2.2K |
| 130 | University of Washington | 1 | 160 | 10.2K | 4.6K |
| 132 | University of Oxford | 2 | 157 | 7.6K | 4.1K |
| 132 | Hirosaki University | 4 | 157 | 408 | 201 |
| 132 | Oita University | 13 | 157 | 163 | 289 |
| 132 | Salk Institute for Biological Studies | 2 | 157 | 267 | 245 |
| 136 | National Institute of Technology and Evaluation | 1 | 153 | 60 | 25 |
| 137 | University of Queensland | 2 | 147 | 6.8K | 3.8K |
| 138 | Wakayama Medical University | 5 | 144 | 360 | 279 |
| 139 | Central Institute for Experimental Animals | 2 | 139 | 71 | 42 |
| 140 | University of East Asia | 1 | 137 | 69 | 9 |
| 141 | University of Missouri | 2 | 136 | 5.0K | 2.6K |
| 142 | Seoul National University | 2 | 135 | 4.6K | 2.5K |
| 143 | Autonomous University of Barcelona | 4 | 134 | 2.4K | 1.9K |
| 144 | Fudan University | 4 | 131 | 3.1K | 2.2K |
| 145 | Kagawa University | 4 | 129 | 506 | 349 |
| 146 | Chubu University | 6 | 128 | 152 | 150 |
| 147 | University of Texas Southwestern Medical Center | 1 | 126 | 3.4K | 1.7K |
| 148 | University of Wisconsin–Madison | 1 | 125 | 10.0K | 4.8K |
| 149 | Massachusetts Institute of Technology | 4 | 122 | 3.0K | 3.1K |
| 149 | Brown University | 2 | 122 | 3.6K | 2.2K |
| 149 | Ludwig Cancer Research | 2 | 122 | 450 | 406 |
| 152 | Tohoku Medical Megabank Organization | 3 | 121 | 56 | 42 |
| 153 | University of Zurich | 2 | 120 | 4.3K | 2.7K |
| 153 | University of Yamanashi | 7 | 120 | 421 | 496 |
| 155 | University of Fukui | 6 | 119 | 369 | 322 |
| 156 | Tata Institute of Fundamental Research | 1 | 115 | 1.0K | 357 |
| 157 | Yamagata University | 5 | 112 | 618 | 534 |
| 158 | National Defense Medical College | 2 | 111 | 703 | 338 |
| 159 | Istanbul University | 1 | 108 | 3.5K | 1.3K |
| 159 | Toyo University | 1 | 108 | 648 | 140 |
| 159 | National Institute of Technology | 1 | 108 | 3.2K | 1.0K |
| 159 | The Nippon Dental University | 1 | 108 | 279 | 85 |
| 159 | Tokyo Women's Medical University | 6 | 108 | 370 | 492 |
| 159 | National Institute of Agrobiological Sciences | 4 | 108 | 202 | 238 |
| 165 | Chinese Academy of Sciences | 3 | 107 | 5.6K | 4.2K |
| 166 | Rutgers University | 3 | 106 | 4.5K | 3.5K |
| 167 | University of Freiburg | 2 | 104 | 2.8K | 1.9K |
| 168 | University of Barcelona | 1 | 103 | 6.1K | 2.8K |
| 169 | Erasmus University Rotterdam | 2 | 102 | 2.2K | 1.5K |
| 170 | Oregon Health & Science University | 3 | 101 | 1.7K | 1.6K |
| 171 | Aichi Gakuin University | 2 | 100 | 169 | 74 |
| 171 | Leonard M. Miller School of Medicine | 2 | 100 | 2.1K | 1.4K |
| 171 | University of Washington Medical Center | 1 | 100 | 660 | 349 |
| 171 | Tampere University Hospital | 2 | 100 | 746 | 597 |
| 175 | Pasteur Institute | 3 | 98 | 983 | 945 |
| 175 | Mukogawa Women's University | 2 | 98 | 132 | 72 |
| 177 | Sojo University | 4 | 95 | 86 | 78 |
| 178 | Czech Academy of Sciences | 1 | 92 | 3.4K | 1.7K |
| 179 | University of Cologne | 6 | 90 | 1.7K | 2.3K |
| 180 | Ibaraki University | 3 | 89 | 652 | 405 |
| 181 | Washington State University | 2 | 87 | 3.5K | 2.5K |
| 181 | Kwansei Gakuin University | 5 | 87 | 192 | 211 |
| 183 | Takasaki University of Health and Welfare | 3 | 85 | 31 | 25 |
| 184 | Tel Aviv University | 3 | 84 | 4.4K | 3.7K |
| 185 | University of California, Berkeley | 1 | 83 | 4.9K | 3.0K |
| 185 | University of California, San Diego | 2 | 83 | 4.2K | 3.3K |
| 187 | Kyoto Sangyo University | 4 | 82 | 104 | 136 |
| 187 | Asahikawa Medical College | 2 | 82 | 395 | 290 |
| 189 | Gwangju Institute of Science and Technology | 3 | 81 | 473 | 460 |
| 189 | Institut Jacques Monod | 3 | 81 | 72 | 130 |
| 189 | Toho University | 3 | 81 | 624 | 464 |
| 189 | Academia Sinica | 5 | 81 | 1.6K | 1.8K |
| 193 | University of Hawaii | 1 | 80 | 3.0K | 1.6K |
| 193 | University of Nottingham | 1 | 80 | 6.9K | 3.7K |
| 193 | Ochanomizu University | 2 | 80 | 426 | 271 |
| 193 | Robert Jones and Agnes Hunt Orthopaedic Hospital | 1 | 80 | 79 | 45 |
| 193 | Hoshi University | 3 | 80 | 166 | 176 |
| 198 | University at Buffalo | 3 | 79 | 3.4K | 3.0K |
| 198 | Kyorin University | 3 | 79 | 236 | 225 |
| 200 | North Carolina State University | 1 | 78 | 5.8K | 3.2K |
| 200 | Shinshu University | 2 | 78 | 1.3K | 818 |
| 200 | University of Western Australia | 2 | 78 | 4.0K | 3.2K |
| 200 | Chung Shan Medical University | 4 | 78 | 310 | 397 |
| 200 | Nagahama Institute of Bio-Science and Technology | 6 | 78 | 13 | 61 |
| 205 | Cornell University | 4 | 76 | 4.1K | 4.6K |
| 205 | Rikkyo University | 5 | 76 | 150 | 189 |
| 207 | Fukuoka Dental College | 4 | 74 | 97 | 95 |
| 208 | Niigata University of Pharmacy and Applied Life Sciences | 2 | 73 | 129 | 100 |
| 209 | Saitama University | 6 | 72 | 391 | 485 |
| 210 | Thomas Jefferson University | 2 | 71 | 2.3K | 1.8K |
| 211 | University of Pittsburgh | 1 | 70 | 8.0K | 4.7K |
| 212 | Carnegie Institution for Science | 2 | 69 | 433 | 472 |
| 213 | University of Texas at Austin | 1 | 68 | 10.0K | 5.9K |
| 214 | National Institute of Neurological Disorders and Stroke | 1 | 66 | 430 | 368 |
| 214 | National Institute of Allergy and Infectious Diseases | 1 | 66 | 1.1K | 853 |
| 216 | University of Vienna | 1 | 65 | 5.6K | 3.2K |
| 216 | Hebrew University of Jerusalem | 1 | 65 | 5.1K | 2.9K |
| 216 | Jikei University School of Medicine | 2 | 65 | 623 | 487 |
| 219 | Nara Women's University | 2 | 64 | 340 | 233 |
| 220 | University of Amsterdam | 2 | 62 | 5.3K | 4.3K |
| 220 | Harbin Institute of Technology | 5 | 62 | 1.6K | 1.7K |
| 220 | Japan Advanced Institute of Science and Technology | 2 | 62 | 363 | 278 |
| 223 | University of Delaware | 1 | 61 | 4.2K | 2.6K |
| 223 | Saitama Medical University | 4 | 61 | 316 | 398 |
| 225 | Azabu University | 3 | 60 | 191 | 177 |
| 225 | Korea Research Institute of Bioscience and Biotechnology | 2 | 60 | 428 | 468 |
| 225 | Korea University | 3 | 60 | 2.5K | 2.5K |
| 228 | Brandeis University | 2 | 59 | 1.1K | 1.1K |
| 228 | Johns Hopkins University | 2 | 59 | 5.7K | 4.8K |
| 228 | University of Hawaii at Manoa | 2 | 59 | 1.5K | 1.4K |
| 228 | Beth Israel Deaconess Medical Center | 2 | 59 | 2.2K | 1.9K |
| 232 | Yonsei University | 6 | 58 | 2.5K | 3.2K |
| 232 | NUI Galway | 1 | 58 | 1.4K | 811 |
| 232 | Shimane University | 2 | 58 | 977 | 650 |
| 232 | Peking Union Medical College | 5 | 58 | 1.4K | 1.7K |
| 236 | Tokyo Metropolitan Matsuzawa Hospital | 1 | 57 | 55 | 37 |
| 237 | Fukuoka University | 2 | 56 | 1.1K | 749 |
| 237 | Royal Perth Hospital | 1 | 56 | 783 | 528 |
| 239 | University of Marburg | 1 | 54 | 3.1K | 2.1K |
| 239 | University of Tartu | 1 | 54 | 2.5K | 1.5K |
| 239 | Leiden University Medical Center | 1 | 54 | 2.7K | 2.0K |
| 242 | University of New South Wales | 1 | 53 | 8.2K | 5.3K |
| 242 | University of Louisville | 1 | 53 | 4.1K | 2.4K |
| 242 | Chongqing Medical University | 2 | 53 | 876 | 771 |
| 242 | Kanazawa Medical University | 3 | 53 | 431 | 491 |
| 246 | Moscow State University | 1 | 52 | 3.8K | 1.9K |
| 246 | Konan University | 4 | 52 | 153 | 184 |
| 246 | Kobe Pharmaceutical University | 1 | 52 | 256 | 186 |
| 246 | Tohoku Medical and Pharmaceutical University | 5 | 52 | 106 | 174 |
| 246 | Josai University | 1 | 52 | 284 | 144 |
| 251 | Stockholm University | 1 | 51 | 4.4K | 2.8K |
| 251 | University of Colorado | 1 | 51 | 6.2K | 4.2K |
| 251 | University of Illinois College of Medicine | 2 | 51 | 650 | 693 |
| 251 | University of Cincinnati Academic Health Center | 1 | 51 | 1.9K | 1.4K |
| 255 | University College London | 1 | 50 | 8.4K | 5.7K |
| 255 | University of Manchester | 1 | 50 | 8.5K | 5.6K |
| 255 | Chungbuk National University | 1 | 50 | 1.8K | 1.1K |
| 255 | Radboud University Nijmegen | 2 | 50 | 3.1K | 3.1K |
| 255 | Bulgarian Academy of Sciences | 1 | 50 | 2.3K | 1.3K |
| 260 | University of Maryland | 1 | 49 | 8.8K | 5.9K |
| 261 | University of Bayreuth | 1 | 48 | 1.9K | 1.3K |
| 261 | Boston Medical Center | 1 | 48 | 1.2K | 858 |
| 263 | University of Bologna | 1 | 47 | 6.9K | 4.5K |
| 263 | University of Calabria | 1 | 47 | 2.0K | 1.4K |
| 263 | Agricultural University of Athens | 3 | 47 | 392 | 568 |
| 266 | Karolinska Institute | 2 | 46 | 3.6K | 3.5K |
| 266 | Sōka University | 2 | 46 | 181 | 139 |
| 266 | Akita University | 1 | 46 | 845 | 417 |
| 269 | St. Marianna University School of Medicine | 2 | 44 | 640 | 593 |
| 270 | University of Southern California | 1 | 43 | 7.9K | 5.6K |
| 270 | Wageningen University & Research | 1 | 43 | 3.1K | 2.5K |
| 270 | Fukuyama University | 1 | 43 | 272 | 179 |
| 270 | Claude Bernard University Lyon 1 | 1 | 43 | 2.6K | 1.9K |
| 274 | Keele University | 1 | 42 | 1.9K | 1.3K |
| 275 | University of Missouri–Kansas City | 1 | 41 | 1.8K | 1.3K |
| 275 | University of Shizuoka | 1 | 41 | 759 | 518 |
| 275 | University of Medicine and Dentistry of New Jersey | 2 | 41 | 520 | 822 |
| 275 | Texas Tech University Health Sciences Center | 1 | 41 | 1.5K | 1.1K |
| 275 | Université de Nantes | 1 | 41 | 1.5K | 1.1K |
| 280 | University of Illinois at Urbana–Champaign | 1 | 39 | 6.2K | 4.7K |
| 280 | Aarhus University | 2 | 39 | 4.6K | 4.5K |
| 280 | University of Arkansas for Medical Sciences | 1 | 39 | 2.5K | 1.9K |
| 280 | Kyushu Institute of Technology | 2 | 39 | 860 | 719 |
| 284 | Aichi Institute of Technology | 1 | 38 | 363 | 183 |
| 284 | Tokyo Dental College | 2 | 38 | 276 | 305 |
| 284 | Shujitsu University | 1 | 38 | 73 | 53 |
| 287 | Nagasaki International University | 2 | 37 | 55 | 82 |
| 287 | Princess Alexandra Hospital | 1 | 37 | 1.0K | 855 |
| 289 | Tokyo University of Agriculture and Technology | 2 | 35 | 1.3K | 1.2K |
| 290 | China Medical University | 2 | 34 | 1.7K | 1.7K |
| 290 | Alternative Energies and Atomic Energy Commission | 1 | 34 | 307 | 336 |
| 290 | Saga University | 2 | 34 | 1.1K | 1.0K |
| 290 | Duke University Hospital | 1 | 34 | 3.7K | 3.2K |
| 290 | Rakuno Gakuen University | 1 | 34 | 311 | 219 |
| 295 | Pennsylvania State University | 1 | 33 | 9.3K | 7.0K |
| 295 | Tezpur University | 2 | 33 | 398 | 460 |
| 297 | National Taiwan University | 1 | 32 | 5.4K | 4.2K |
| 297 | Dokkyo Medical University | 2 | 32 | 336 | 354 |
| 297 | University of Duisburg-Essen | 1 | 32 | 3.4K | 2.8K |
| 297 | Fred Hutchinson Cancer Research Center | 1 | 32 | 1.3K | 1.3K |
| 297 | Hyogo University | 1 | 32 | 25 | 29 |
| 297 | Toyohashi University of Technology | 1 | 32 | 969 | 687 |
| 303 | Yale University | 2 | 31 | 5.0K | 5.3K |
| 303 | Polish Academy of Sciences | 1 | 31 | 4.5K | 3.4K |
| 303 | University of Gdańsk | 1 | 31 | 1.6K | 899 |
| 303 | Douglas Mental Health University Institute | 1 | 31 | 307 | 350 |
| 303 | Kobe Gakuin University | 2 | 31 | 159 | 179 |
| 303 | Musashino University | 1 | 31 | 169 | 113 |
| 309 | University of Naples Federico II | 1 | 30 | 4.9K | 4.0K |
| 309 | University of Oregon | 1 | 30 | 3.2K | 2.7K |
| 309 | University of Nice Sophia Antipolis | 1 | 30 | 1.2K | 1.1K |
| 309 | Wright State University | 1 | 30 | 1.9K | 1.5K |
| 309 | Himeji Dokkyo University | 1 | 30 | 91 | 70 |
| 314 | University of Michigan | 4 | 29 | 6.4K | 8.5K |
| 314 | Osaka Prefecture University | 5 | 29 | 806 | 1.4K |
| 314 | Ministry of Health | 2 | 29 | 2.4K | 2.2K |
| 317 | University of Melbourne | 1 | 28 | 9.1K | 7.3K |
| 317 | University of Rennes 1 | 1 | 28 | 1.8K | 1.7K |
| 317 | Lawrence Berkeley National Laboratory | 1 | 28 | 1.6K | 1.7K |
| 317 | Iwate Medical University | 1 | 28 | 864 | 689 |
| 321 | Xi'an Jiaotong University | 1 | 27 | 3.6K | 2.9K |
| 321 | Iwate University | 4 | 27 | 430 | 753 |
| 321 | Army Medical University | 1 | 27 | 1.6K | 1.4K |
| 321 | Kyoto University Hospital | 1 | 27 | 663 | 571 |
| 325 | University of Ljubljana | 1 | 26 | 4.1K | 3.2K |
| 325 | Royal Children's Hospital | 1 | 26 | 1.2K | 1.2K |
| 325 | Murdoch Children's Research Institute | 1 | 26 | 894 | 966 |
| 328 | Syracuse University | 1 | 25 | 3.4K | 2.9K |
| 328 | Kyoto Pharmaceutical University | 1 | 25 | 417 | 410 |
| 328 | National Cancer Center | 2 | 25 | 847 | 1.1K |
| 331 | Technion – Israel Institute of Technology | 2 | 24 | 2.8K | 3.4K |
| 331 | AstraZeneca | 1 | 24 | 922 | 1.0K |
| 331 | Case Western Reserve University | 2 | 24 | 4.1K | 4.8K |
| 331 | University of Paris-Sud | 1 | 24 | 2.1K | 2.1K |
| 335 | Mazda | 1 | 23 | 85 | 35 |
| 335 | Chalmers University of Technology | 1 | 23 | 2.8K | 2.6K |
| 335 | University of Texas Medical Branch | 2 | 23 | 2.2K | 2.7K |
| 335 | University of Tampere | 1 | 23 | 2.9K | 2.6K |
| 335 | Netherlands Institute for Neuroscience | 1 | 23 | 268 | 364 |
| 335 | Aichi Shukutoku University | 1 | 23 | 86 | 77 |
| 335 | University Health Network | 3 | 23 | 1.3K | 2.1K |
| 335 | Yong Loo Lin School of Medicine | 2 | 23 | 1.1K | 1.6K |
| 335 | KEK | 1 | 23 | 356 | 282 |
| 344 | Asahi Kasei | 1 | 22 | 55 | 64 |
| 344 | Kyowa Hakko Kirin | 1 | 22 | 21 | 47 |
| 344 | University of the Philippines Manila | 1 | 22 | 223 | 201 |
| 344 | University of Virginia | 1 | 22 | 6.4K | 5.5K |
| 348 | Pusan National University | 2 | 21 | 2.1K | 2.5K |
| 348 | University of Delhi | 1 | 21 | 3.1K | 2.4K |
| 348 | Asahikawa Medical University | 1 | 21 | 505 | 445 |
| 348 | Rice University | 1 | 21 | 3.1K | 3.1K |
| 348 | Indian Agricultural Research Institute | 1 | 21 | 511 | 557 |
| 348 | Institute for Environmental Sciences | 1 | 21 | 42 | 61 |
| 354 | KAIST | 1 | 20 | 2.5K | 2.4K |
| 354 | Aoyama Gakuin University | 1 | 20 | 611 | 457 |
| 354 | Sichuan University | 1 | 20 | 4.9K | 4.2K |
| 354 | State University of New York Upstate Medical University | 1 | 20 | 1.6K | 1.6K |
| 354 | Garvan Institute of Medical Research | 1 | 20 | 480 | 637 |
| 354 | Meiji Pharmaceutical University | 1 | 20 | 281 | 319 |
| 354 | St. Joseph's Hospital and Medical Center | 1 | 20 | 236 | 299 |
| 354 | National Institute on Alcohol Abuse and Alcoholism | 1 | 20 | 439 | 642 |
| 354 | National Institute of Diabetes and Digestive and Kidney Diseases | 1 | 20 | 501 | 674 |
| 363 | Pompeu Fabra University | 1 | 19 | 1.6K | 1.7K |
| 363 | Chungnam National University | 1 | 19 | 2.4K | 2.2K |
| 363 | University of Arizona | 1 | 19 | 7.7K | 6.9K |
| 363 | Oregon State University | 1 | 19 | 4.4K | 4.3K |
| 363 | University of Exeter | 1 | 19 | 4.4K | 4.1K |
| 363 | National Human Genome Research Institute | 1 | 19 | 603 | 812 |
| 363 | Aomori University | 1 | 19 | 60 | 66 |
| 363 | WEHI | 1 | 19 | 450 | 566 |
| 371 | University of Geneva | 1 | 18 | 4.1K | 4.1K |
| 371 | Helmholtz Zentrum München | 1 | 18 | 973 | 1.2K |
| 371 | Utsunomiya University | 1 | 18 | 808 | 651 |
| 371 | Ninewells Hospital | 1 | 18 | 813 | 962 |
| 375 | ETH Zürich | 1 | 17 | 4.8K | 4.8K |
| 375 | Saint Petersburg State University | 1 | 17 | 1.5K | 1.1K |
| 375 | University Hospital Zurich | 1 | 17 | 1.9K | 2.0K |
| 375 | National Institute of Environmental Health Sciences | 1 | 17 | 1.1K | 1.3K |
| 375 | Kagawa Prefectural College of Health Sciences | 1 | 17 | 19 | 24 |
| 375 | University of California, Irvine | 1 | 17 | 3.0K | 3.4K |
| 375 | Daiichi University of Pharmacy | 2 | 17 | 21 | 45 |
| 375 | Gunma Prefectural College of Health Sciences | 1 | 17 | 30 | 30 |
| 383 | University of Perugia | 1 | 16 | 3.4K | 3.5K |
| 383 | University of Illinois at Chicago | 1 | 16 | 5.8K | 5.8K |
| 383 | Ontario Cancer Institute | 1 | 16 | 386 | 575 |
| 386 | Katholieke Universiteit Leuven | 1 | 15 | 7.2K | 7.0K |
| 386 | Colorado State University | 1 | 15 | 4.7K | 4.9K |
| 386 | Vertex Pharmaceuticals | 1 | 15 | 146 | 247 |
| 386 | Aichi Medical University | 1 | 15 | 832 | 826 |
| 386 | University of Rochester Medical Center | 1 | 15 | 2.1K | 2.3K |
| 391 | University of Bonn | 1 | 14 | 4.5K | 4.6K |
| 391 | Emory University | 2 | 14 | 3.9K | 5.3K |
| 391 | Tokyo Metropolitan Police Hospital | 1 | 14 | 103 | 126 |
| 394 | Lehigh University | 1 | 13 | 2.1K | 2.3K |
| 394 | Kōchi University | 1 | 13 | 1.1K | 1.2K |
| 394 | Nanjing Agricultural University | 2 | 13 | 1.2K | 1.7K |
| 394 | Peking University Third Hospital | 1 | 13 | 974 | 1.1K |
| 394 | Peking University First Hospital | 1 | 13 | 960 | 1.1K |
| 394 | ComUE Paris-Saclay University | 1 | 13 | 2.8K | 2.8K |
| 400 | University of Tübingen | 1 | 12 | 4.6K | 5.0K |
| 400 | University of Münster | 1 | 12 | 4.0K | 4.4K |
| 400 | Research Institute of Molecular Pathology | 1 | 12 | 114 | 201 |
| 400 | Silesian University of Technology | 1 | 12 | 1.2K | 1.3K |
| 400 | VU University Medical Center | 1 | 12 | 1.8K | 2.3K |
| 400 | Shenzhen University | 1 | 12 | 2.2K | 2.4K |
| 400 | Madurai Kamaraj University | 2 | 12 | 409 | 744 |
| 400 | The Christie NHS Foundation Trust | 1 | 12 | 455 | 620 |
| 400 | Mitsubishi Tanabe Pharma | 2 | 12 | 77 | 238 |
| 400 | Osaka Dental University | 1 | 12 | 216 | 265 |
| 400 | Hyogo Cancer Center | 1 | 12 | 150 | 154 |
| 400 | Hebei General Hospital | 1 | 12 | 185 | 238 |
| 412 | New York University | 1 | 11 | 6.1K | 6.2K |
| 412 | Sapporo Medical University | 2 | 11 | 577 | 1.0K |
| 412 | Beppu University | 1 | 11 | 27 | 34 |
| 412 | University of Sussex | 1 | 11 | 3.6K | 4.0K |
| 412 | Hokkaido University of Education | 1 | 11 | 213 | 232 |
| 412 | Texas Christian University | 1 | 11 | 1.2K | 1.4K |
| 412 | Meisei University | 1 | 11 | 235 | 238 |
| 412 | Anschutz Medical Campus | 1 | 11 | 2.0K | 2.3K |
| 412 | Ishikawa Prefectural Nursing University | 1 | 11 | 30 | 46 |
| 421 | Ajou University | 1 | 10 | 1.6K | 2.0K |
| 421 | Tianjin Medical University | 1 | 10 | 1.2K | 1.4K |
| 421 | Rajshahi University | 1 | 10 | 645 | 851 |
| 421 | Maebashi Institute of Technology | 1 | 10 | 103 | 128 |
| 421 | Suzuka University of Medical Science | 1 | 10 | 177 | 250 |
| 421 | Symbiosis International University | 1 | 10 | 266 | 308 |
| 421 | National Agricultural Research Center for Hokkaido Region | 1 | 10 | 112 | 193 |
| 421 | Tianjin Medical University Cancer Institute and Hospital | 1 | 10 | 422 | 615 |
| 429 | University of Alberta | 1 | 9 | 9.2K | 9.4K |
| 429 | University of Málaga | 1 | 9 | 2.7K | 3.1K |
| 429 | Obihiro University of Agriculture and Veterinary Medicine | 1 | 9 | 470 | 630 |
| 432 | University of Birmingham | 1 | 8 | 7.2K | 7.8K |
| 432 | Mayo Clinic | 1 | 8 | 4.5K | 4.7K |
| 432 | Guangxi University | 1 | 8 | 1.6K | 1.8K |
| 432 | Huazhong Agricultural University | 1 | 8 | 1.5K | 1.9K |
| 432 | Kanagawa University of Human Services | 1 | 8 | 74 | 120 |
| 432 | Agricultural University of Hebei | 1 | 8 | 568 | 769 |
| 432 | California Pacific Medical Center | 1 | 8 | 427 | 709 |
| 432 | Meikai University | 1 | 8 | 81 | 124 |
| 432 | National Institute of Public Health | 1 | 8 | 1.2K | 1.6K |
| 441 | University of Florida | 1 | 7 | 9.6K | 10.2K |
| 441 | Airlangga University | 1 | 7 | 1.5K | 996 |
| 441 | Tokyo Medical University | 1 | 7 | 1.2K | 1.5K |
| 441 | International University of Health and Welfare | 4 | 7 | 182 | 814 |
| 445 | Canadian Pacific Railway | 1 | 6 | 10 | 22 |
| 445 | Great Ormond Street Hospital | 1 | 6 | 1.4K | 1.9K |
| 445 | Portland State University | 1 | 6 | 2.0K | 2.9K |
| 445 | Alfaisal University | 1 | 6 | 392 | 693 |
| 445 | Johns Hopkins All Children's Hospital | 1 | 6 | 191 | 297 |
| 445 | Jawaharlal Nehru Centre for Advanced Scientific Research | 1 | 6 | 392 | 612 |
| 445 | Tata Memorial Centre | 1 | 6 | 678 | 869 |
| 445 | Ludong University | 1 | 6 | 657 | 879 |
| 445 | National Institute of Animal Health | 1 | 6 | 256 | 381 |
| 445 | National Institute for Physiological Sciences | 2 | 6 | 223 | 515 |
| 455 | University of Stuttgart | 1 | 5 | 2.7K | 3.4K |
| 455 | University of Strasbourg | 1 | 5 | 2.3K | 3.3K |
| 455 | Kochi University of Technology | 1 | 5 | 409 | 527 |
| 455 | University of Indonesia | 1 | 5 | 1.9K | 2.0K |
| 455 | Tokyo Gakugei University | 1 | 5 | 434 | 608 |
| 455 | Tenshi College | 1 | 5 | 18 | 53 |
| 455 | National Health Research Institutes | 2 | 5 | 538 | 1.3K |
| 455 | Tokoha Gakuen University | 1 | 5 | 52 | 87 |
| 455 | Veer Bahadur Singh Purvanchal University | 1 | 5 | 41 | 92 |
| 455 | Biotechnology Research Center | 1 | 5 | 37 | 119 |
| 455 | Amity University, Kolkata | 1 | 5 | 19 | 44 |
| 455 | Indian Institute of Technology Guwahati | 1 | 5 | 1.4K | 1.8K |
| 467 | University of London | 1 | 4 | 7.7K | 9.1K |
| 467 | Kyung Hee University | 1 | 4 | 3.1K | 4.1K |
| 467 | Dong-A University | 1 | 4 | 937 | 1.5K |
| 467 | Banaras Hindu University | 1 | 4 | 2.6K | 3.5K |
| 467 | Southwest University | 1 | 4 | 2.2K | 2.9K |
| 467 | University of Science and Technology of China | 1 | 4 | 3.2K | 4.0K |
| 467 | Kibi International University | 1 | 4 | 67 | 139 |
| 467 | Sanford Burnham Prebys Medical Discovery Institute | 1 | 4 | 420 | 766 |
| 467 | Tamagawa University | 1 | 4 | 235 | 352 |
| 467 | St George's, University of London | 1 | 4 | 916 | 1.5K |
| 477 | Akita Prefectural University | 1 | 3 | 469 | 733 |
| 477 | Austrian Academy of Sciences | 1 | 3 | 1.1K | 1.8K |
| 477 | Meharry Medical College | 1 | 3 | 474 | 957 |
| 477 | Dayeh University | 1 | 3 | 458 | 849 |
| 477 | Temasek Life Sciences Laboratory | 1 | 3 | 7 | 41 |
| 477 | Eastern Hepatobiliary Surgery Hospital | 1 | 3 | 265 | 484 |
| 477 | University of California, Santa Barbara | 1 | 3 | 2.3K | 3.5K |
| 484 | Sun Yat-sen University | 1 | 2 | 5.3K | 6.7K |
| 484 | Soochow University | 1 | 2 | 2.6K | 3.8K |
| 484 | Mahidol University | 1 | 2 | 3.5K | 5.0K |
| 484 | University of Florence | 1 | 2 | 5.1K | 6.9K |
| 484 | Consiglio Nazionale delle Ricerche | 1 | 2 | 2.0K | 3.2K |
| 484 | University of the Ryukyus | 2 | 2 | 1.2K | 2.5K |
| 484 | Siriraj Hospital | 1 | 2 | 693 | 1.3K |
| 484 | National Heart, Lung, and Blood Institute | 1 | 2 | 1.2K | 1.8K |
| 492 | University of British Columbia | 1 | 1 | 10.0K | 12.8K |
| 492 | Tokyo University of Technology | 1 | 1 | 325 | 634 |
| 492 | University of Dundee | 1 | 1 | 3.0K | 4.8K |
| 492 | King Abdullah University of Science and Technology | 1 | 1 | 1.4K | 2.1K |
| 492 | Hyogo University of Health Sciences | 1 | 1 | 112 | 299 |
| 492 | National Hospital Organization Kyushu Cancer Center | 1 | 1 | 100 | 164 |
| 492 | Sanyo-onoda City University | 1 | 1 | 89 | 199 |
| 499 | Monash University | 1 | 0 | 8.0K | 11.1K |
| 499 | Occidental College | 1 | 0 | 344 | 848 |
| 499 | La Trobe University | 1 | 0 | 3.9K | 6.3K |
| 499 | University of the Punjab | 1 | 0 | 1.4K | 2.5K |
| 499 | Kagawa Nutrition University | 1 | 0 | 110 | 232 |
| 499 | Kawasaki Medical School | 1 | 0 | 64 | 146 |
| 499 | Japan Biological Informatics Consortium | 1 | 0 | 16 | 59 |
| 499 | Osaka Metropolitan University | 1 | 0 | 110 | 316 |
