| # | Institution | Articles | Citations | R ARank | R CRank |
|---|
| 1 | Kyoto University | 127 | 1,786 | 265 | 529 |
| 2 | Hokkaido University | 67 | 1,000 | 271 | 452 |
| 3 | Kyushu University | 31 | 475 | 775 | 1.0K |
| 4 | University of Tokyo | 30 | 458 | 1.6K | 2.0K |
| 5 | Nagoya University | 27 | 454 | 618 | 805 |
| 6 | Chukyo University | 13 | 376 | 19 | 4 |
| 7 | Osaka City University | 19 | 308 | 178 | 237 |
| 8 | Macquarie University | 11 | 253 | 693 | 921 |
| 9 | University of the Ryukyus | 15 | 244 | 170 | 237 |
| 10 | Tokyo Gakugei University | 13 | 241 | 38 | 37 |
| 11 | National Autonomous University of Mexico | 9 | 211 | 1.5K | 1.2K |
| 12 | University of Trnava | 8 | 177 | 11 | 17 |
| 13 | Okayama University | 17 | 175 | 401 | 741 |
| 14 | Rissho University | 9 | 173 | 13 | 4 |
| 15 | Polish Academy of Sciences | 9 | 170 | 1.4K | 1.6K |
| 16 | Universidade Estadual Paulista | 6 | 166 | 1.2K | 771 |
| 17 | Ibaraki University | 17 | 155 | 127 | 243 |
| 17 | Gifu University | 7 | 155 | 556 | 486 |
| 19 | Utah State University | 7 | 150 | 675 | 975 |
| 20 | Tokyo Metropolitan University | 11 | 147 | 403 | 486 |
| 21 | Adam Mickiewicz University in Poznań | 7 | 146 | 491 | 396 |
| 22 | University of California | 10 | 141 | 8.1K | 8.6K |
| 23 | Nagasaki University | 7 | 137 | 526 | 556 |
| 24 | Federal University of Pernambuco | 7 | 123 | 790 | 638 |
| 25 | University of South Bohemia in České Budějovice | 9 | 119 | 136 | 304 |
| 26 | Federal University of Rio Grande do Sul | 5 | 116 | 2.1K | 1.5K |
| 27 | National Institute of Polar Research | 3 | 113 | 114 | 98 |
| 28 | Ehime University | 9 | 112 | 379 | 550 |
| 29 | Michigan State University | 2 | 111 | 6.1K | 3.8K |
| 30 | National Zoological Park | 5 | 107 | 82 | 128 |
| 31 | University of Miyazaki | 10 | 104 | 281 | 547 |
| 32 | National University of Mar del Plata | 4 | 103 | 208 | 194 |
| 33 | Indian Institute of Science | 2 | 102 | 2.2K | 1.2K |
| 34 | Czech Academy of Sciences | 3 | 100 | 2.1K | 1.6K |
| 34 | Chinese Academy of Sciences | 11 | 100 | 3.5K | 4.3K |
| 34 | University of the Witwatersrand | 5 | 100 | 1.5K | 1.5K |
| 37 | University of Tsukuba | 10 | 99 | 1.3K | 2.0K |
| 38 | University of Southern Mississippi | 3 | 97 | 679 | 546 |
| 39 | Moscow State University | 6 | 96 | 1.6K | 1.4K |
| 40 | Australian National University | 4 | 95 | 2.7K | 2.8K |
| 40 | University of Melbourne | 6 | 95 | 4.1K | 4.7K |
| 42 | Flinders University | 5 | 94 | 1.1K | 1.5K |
| 42 | Miyagi University | 3 | 94 | 45 | 33 |
| 44 | Seoul National University | 11 | 93 | 1.7K | 3.0K |
| 45 | Karnatak University | 5 | 88 | 164 | 186 |
| 46 | National Center for Scientific Research | 8 | 87 | 4.1K | 4.9K |
| 46 | Slovak Academy of Sciences | 4 | 87 | 1.0K | 978 |
| 48 | Utrecht University | 3 | 85 | 3.8K | 3.6K |
| 48 | Keio University | 7 | 85 | 1.2K | 1.6K |
| 48 | University of Stirling | 1 | 85 | 2.0K | 1.1K |
| 51 | Mie University | 8 | 84 | 310 | 576 |
| 52 | Goethe University Frankfurt | 1 | 83 | 3.8K | 1.9K |
| 52 | Smithsonian Institution | 3 | 83 | 475 | 469 |
| 52 | University of Padua | 1 | 83 | 6.6K | 3.6K |
| 55 | University of Regensburg | 8 | 82 | 742 | 1.6K |
| 55 | University of São Paulo | 9 | 82 | 3.5K | 4.4K |
| 55 | Nara Women's University | 9 | 82 | 87 | 192 |
| 58 | North Carolina State University | 3 | 81 | 3.5K | 3.2K |
| 59 | University of Memphis | 2 | 80 | 1.6K | 1.2K |
| 60 | University of Hamburg | 5 | 79 | 1.6K | 2.2K |
| 60 | Tamagawa University | 5 | 79 | 61 | 65 |
| 62 | University of Salamanca | 6 | 78 | 677 | 1.0K |
| 62 | Kagoshima University | 10 | 78 | 438 | 907 |
| 62 | Saga University | 3 | 78 | 788 | 592 |
| 65 | Tokyo University of Agriculture and Technology | 9 | 77 | 469 | 798 |
| 66 | Ohio State University | 2 | 76 | 7.6K | 5.4K |
| 67 | Wuhan University | 4 | 74 | 2.2K | 2.1K |
| 67 | University of Victoria | 2 | 74 | 2.5K | 1.9K |
| 69 | University of Buenos Aires | 5 | 72 | 1.7K | 1.8K |
| 69 | Maria Curie-Skłodowska University | 3 | 72 | 525 | 360 |
| 69 | Indian Institutes of Science Education and Research | 2 | 72 | 294 | 193 |
| 72 | University of Adelaide | 3 | 69 | 3.4K | 3.4K |
| 72 | University of Auckland | 4 | 69 | 3.0K | 3.6K |
| 72 | Kanazawa University | 3 | 69 | 1.7K | 1.4K |
| 72 | Katholieke Universiteit Leuven | 2 | 69 | 5.4K | 4.3K |
| 72 | National University of Córdoba | 5 | 69 | 535 | 683 |
| 77 | University of Canterbury | 6 | 66 | 748 | 1.5K |
| 77 | Palacký University Olomouc | 5 | 66 | 434 | 714 |
| 79 | Ben-Gurion University of the Negev | 2 | 64 | 3.3K | 2.6K |
| 80 | Rikkyo University | 4 | 63 | 184 | 215 |
| 80 | Kobe University | 5 | 63 | 1.2K | 1.5K |
| 80 | Akvaplan-niva | 1 | 63 | 47 | 42 |
| 83 | Tel Aviv University | 4 | 61 | 3.8K | 4.2K |
| 83 | Smithsonian Tropical Research Institute | 5 | 61 | 147 | 315 |
| 83 | Mammal Research Institute of the Polish Academy of Sciences | 4 | 61 | 34 | 68 |
| 83 | National Scientific and Technical Research Council | 5 | 61 | 789 | 1.2K |
| 87 | Complutense University of Madrid | 4 | 59 | 2.8K | 2.9K |
| 87 | University of Pretoria | 6 | 59 | 1.1K | 1.8K |
| 87 | Kyoto Prefectural University | 3 | 59 | 211 | 233 |
| 87 | Tohoku University | 4 | 59 | 3.0K | 3.0K |
| 87 | University of Granada | 2 | 59 | 3.4K | 2.5K |
| 92 | University of Zurich | 3 | 58 | 3.5K | 3.7K |
| 93 | Hiroshima University | 8 | 57 | 1.3K | 2.2K |
| 94 | University of Texas at Austin | 2 | 56 | 7.9K | 6.3K |
| 94 | University of Glasgow | 1 | 56 | 6.4K | 3.9K |
| 94 | University of Lausanne | 1 | 56 | 3.3K | 2.2K |
| 94 | Jacob Blaustein Institutes for Desert Research | 1 | 56 | 256 | 207 |
| 98 | University of Turin | 3 | 54 | 3.3K | 3.4K |
| 98 | Toho University | 5 | 54 | 410 | 619 |
| 100 | Autonomous University of Madrid | 2 | 53 | 2.8K | 2.3K |
| 100 | Swedish University of Agricultural Sciences | 2 | 53 | 1.4K | 1.4K |
| 102 | Masaryk University | 3 | 52 | 1.2K | 1.3K |
| 102 | University of Windsor | 1 | 52 | 2.2K | 1.2K |
| 104 | University of Florence | 3 | 51 | 3.1K | 3.3K |
| 104 | Tokyo University of Marine Science and Technology | 5 | 51 | 156 | 271 |
| 104 | Nihon University | 5 | 51 | 1.2K | 1.5K |
| 104 | University of Campinas | 4 | 51 | 2.3K | 2.4K |
| 108 | University of Würzburg | 2 | 50 | 2.6K | 2.5K |
| 108 | University of Maryland | 3 | 50 | 5.7K | 5.8K |
| 108 | Kyushu International University | 2 | 50 | 6 | 3 |
| 108 | Iowa State University | 3 | 50 | 3.3K | 3.6K |
| 112 | University of Bristol | 2 | 49 | 4.8K | 4.3K |
| 112 | University of Lisbon | 4 | 49 | 1.9K | 2.6K |
| 112 | Federal University of Paraíba | 2 | 49 | 1.3K | 717 |
| 115 | University of Cincinnati | 1 | 48 | 5.3K | 3.5K |
| 115 | Tokyo Institute of Technology | 3 | 48 | 2.3K | 2.2K |
| 115 | National University of Singapore | 4 | 48 | 4.2K | 5.1K |
| 115 | University of Wisconsin–Madison | 3 | 48 | 6.7K | 6.8K |
| 115 | Shaanxi Normal University | 3 | 48 | 823 | 855 |
| 120 | University of Queensland | 2 | 47 | 6.8K | 6.0K |
| 120 | Aarhus University | 2 | 47 | 4.6K | 4.1K |
| 122 | Austrian Academy of Sciences | 2 | 45 | 699 | 855 |
| 122 | University of South Florida | 2 | 45 | 4.2K | 3.8K |
| 122 | University of Florida | 3 | 45 | 6.4K | 6.5K |
| 122 | University of Jyväskylä | 2 | 45 | 1.6K | 1.7K |
| 126 | The Graduate University for Advanced Studies | 2 | 44 | 277 | 304 |
| 127 | University of Potsdam | 4 | 43 | 803 | 1.5K |
| 127 | Uppsala University | 2 | 43 | 4.9K | 4.6K |
| 127 | University of Missouri | 2 | 43 | 5.0K | 4.5K |
| 127 | University of Roehampton | 1 | 43 | 699 | 427 |
| 127 | Universidad de la República | 4 | 43 | 526 | 902 |
| 132 | University of Milan | 3 | 42 | 3.8K | 4.2K |
| 132 | Yamagata University | 2 | 42 | 1.2K | 979 |
| 132 | Chiba University | 6 | 42 | 1.3K | 2.1K |
| 132 | Joetsu University of Education | 5 | 42 | 31 | 69 |
| 136 | University of Oslo | 4 | 41 | 3.2K | 4.5K |
| 136 | Rutgers University | 3 | 41 | 4.5K | 5.3K |
| 136 | University of Pisa | 4 | 41 | 2.7K | 3.7K |
| 136 | University of Lorraine | 3 | 41 | 1.0K | 1.3K |
| 136 | Japan Weather Association | 1 | 41 | 18 | 11 |
| 141 | University of Hong Kong | 2 | 40 | 5.1K | 4.8K |
| 141 | Wakayama University | 4 | 40 | 115 | 165 |
| 143 | University of Chile | 3 | 39 | 2.4K | 2.8K |
| 144 | University of Sydney | 1 | 38 | 9.3K | 6.8K |
| 144 | National Institute of Agrobiological Sciences | 1 | 38 | 420 | 411 |
| 146 | University of Vigo | 2 | 37 | 1.1K | 1.2K |
| 147 | University of Giessen | 1 | 36 | 3.2K | 2.5K |
| 147 | Aristotle University of Thessaloniki | 2 | 36 | 4.0K | 3.7K |
| 147 | University of Georgia | 3 | 36 | 3.8K | 4.5K |
| 147 | Muséum national d'histoire naturelle | 1 | 36 | 934 | 829 |
| 147 | Czech University of Life Sciences Prague | 3 | 36 | 267 | 391 |
| 147 | Konrad Lorenz Institute of Ethology | 2 | 36 | 20 | 28 |
| 147 | Federal University of Uberlândia | 5 | 36 | 509 | 833 |
| 147 | Hainan Normal University | 5 | 36 | 89 | 206 |
| 147 | Tokiwa University | 1 | 36 | 24 | 14 |
| 156 | Tottori University | 2 | 35 | 1.3K | 1.2K |
| 157 | Freie Universität Berlin | 2 | 34 | 3.1K | 3.4K |
| 157 | Lund University | 2 | 34 | 5.3K | 5.4K |
| 157 | Sophia University | 3 | 34 | 387 | 484 |
| 157 | Universidade Federal do Paraná | 3 | 34 | 1.8K | 1.7K |
| 157 | Tethys Research Institute | 1 | 34 | 24 | 28 |
| 157 | California State University, Long Beach | 1 | 34 | 673 | 515 |
| 163 | Virginia Tech | 1 | 33 | 5.9K | 4.5K |
| 163 | Rhodes University | 2 | 33 | 786 | 856 |
| 163 | Shizuoka University | 6 | 33 | 561 | 1.1K |
| 166 | University of Cambridge | 2 | 32 | 7.1K | 7.0K |
| 166 | McMaster University | 2 | 32 | 5.1K | 5.2K |
| 166 | Kindai University | 3 | 32 | 1.1K | 1.4K |
| 166 | Carleton University | 2 | 32 | 2.4K | 2.6K |
| 166 | Memorial University of Newfoundland | 2 | 32 | 2.2K | 2.5K |
| 166 | Simón Bolívar University | 2 | 32 | 427 | 463 |
| 172 | University of Texas–Pan American | 1 | 31 | 453 | 452 |
| 172 | National University of Río Cuarto | 2 | 31 | 277 | 353 |
| 172 | Griffith University | 2 | 31 | 3.3K | 3.7K |
| 172 | University of Montana | 1 | 31 | 1.5K | 1.2K |
| 172 | Azabu University | 3 | 31 | 191 | 295 |
| 177 | Duke University | 1 | 30 | 6.1K | 5.0K |
| 177 | Mbarara University of Science and Technology | 2 | 30 | 143 | 182 |
| 177 | Pontifical Catholic University of Chile | 3 | 30 | 1.5K | 2.3K |
| 177 | University of Lucknow | 3 | 30 | 324 | 507 |
| 177 | Duke University Hospital | 1 | 30 | 3.7K | 3.3K |
| 182 | Cornell University | 5 | 29 | 3.6K | 6.3K |
| 182 | Volcani Institute of Agricultural Research | 1 | 29 | 229 | 278 |
| 182 | Federal University of Minas Gerais | 3 | 29 | 2.4K | 3.0K |
| 182 | Humboldt State University | 1 | 29 | 416 | 381 |
| 182 | University of the Philippines | 1 | 29 | 850 | 691 |
| 182 | Nara Institute of Science and Technology | 2 | 29 | 600 | 712 |
| 182 | Natural History Museum | 2 | 29 | 838 | 968 |
| 182 | Museo Nacional de Ciencias Naturales | 3 | 29 | 140 | 292 |
| 190 | Charles University | 4 | 28 | 2.3K | 3.4K |
| 190 | Eötvös Loránd University | 1 | 28 | 2.1K | 1.6K |
| 190 | University of Oklahoma | 2 | 28 | 3.1K | 3.5K |
| 190 | Florida International University | 2 | 28 | 2.4K | 2.9K |
| 190 | RIKEN | 2 | 28 | 1.0K | 1.3K |
| 190 | Iwate University | 1 | 28 | 1.0K | 742 |
| 190 | Hungarian University of Agriculture and Life Sciences | 1 | 28 | 695 | 529 |
| 190 | University of Wyoming | 3 | 28 | 1.3K | 2.2K |
| 190 | Primate Research Institute | 2 | 28 | 40 | 68 |
| 190 | Instituto Biológico | 2 | 28 | 218 | 292 |
| 190 | Indiana University | 3 | 28 | 3.9K | 5.1K |
| 201 | University of Sassari | 1 | 27 | 1.6K | 1.5K |
| 201 | University of Santiago de Compostela | 1 | 27 | 2.4K | 2.0K |
| 201 | Universidade de Santiago de Compostela | 1 | 27 | 1.4K | 1.1K |
| 204 | Ludwig Maximilian University of Munich | 2 | 26 | 4.2K | 4.8K |
| 204 | University of Bonn | 1 | 26 | 4.5K | 3.9K |
| 204 | University of Derby | 1 | 26 | 707 | 601 |
| 207 | Harvard University | 2 | 25 | 5.5K | 6.4K |
| 207 | University of Washington | 2 | 25 | 8.1K | 8.5K |
| 207 | California State University | 3 | 25 | 3.0K | 4.1K |
| 207 | Western Carolina University | 1 | 25 | 541 | 419 |
| 207 | University of San Francisco | 1 | 25 | 726 | 697 |
| 207 | Halmstad University | 1 | 25 | 315 | 338 |
| 213 | University of Chicago | 1 | 24 | 6.7K | 5.7K |
| 213 | University of Alberta | 2 | 24 | 7.2K | 7.5K |
| 213 | University of Sheffield | 1 | 24 | 6.6K | 5.7K |
| 213 | International Christian University | 3 | 24 | 68 | 166 |
| 213 | Yamashina Institute for Ornithology | 1 | 24 | 26 | 28 |
| 218 | Princeton University | 3 | 23 | 2.7K | 4.0K |
| 218 | James Cook University | 1 | 23 | 2.3K | 2.3K |
| 218 | University of Ottawa | 1 | 23 | 6.6K | 5.5K |
| 218 | University of Miami | 1 | 23 | 5.4K | 4.7K |
| 218 | Newcastle University | 2 | 23 | 4.0K | 4.8K |
| 223 | University of Kansas | 2 | 22 | 4.2K | 5.0K |
| 223 | Jagiellonian University | 2 | 22 | 2.5K | 2.6K |
| 223 | Osaka Prefecture University | 2 | 22 | 1.4K | 1.5K |
| 223 | University of Illinois at Urbana–Champaign | 1 | 22 | 6.2K | 5.6K |
| 223 | University of La Laguna | 1 | 22 | 1.8K | 1.6K |
| 223 | Université libre de Bruxelles | 3 | 22 | 2.2K | 3.7K |
| 223 | University of South Africa | 1 | 22 | 1.5K | 1.1K |
| 223 | Umutara Polytechnic | 1 | 22 | 1 | 1 |
| 223 | Tokyo University of Agriculture | 2 | 22 | 530 | 682 |
| 223 | Ghent University | 1 | 22 | 6.5K | 5.9K |
| 223 | Beijing Normal University | 3 | 22 | 1.7K | 2.5K |
| 223 | University of Lethbridge | 1 | 22 | 1.1K | 1.1K |
| 223 | University of Crete | 2 | 22 | 1.5K | 2.3K |
| 223 | Bucknell University | 1 | 22 | 805 | 725 |
| 223 | Okinawa Institute of Science and Technology | 1 | 22 | 367 | 376 |
| 223 | Nara University | 1 | 22 | 33 | 31 |
| 223 | Zoological Society of London | 1 | 22 | 253 | 325 |
| 223 | Poznań University of Life Sciences | 2 | 22 | 429 | 585 |
| 241 | University of Liverpool | 3 | 21 | 3.8K | 5.5K |
| 241 | University of Guelph | 1 | 21 | 3.9K | 3.6K |
| 241 | Hunter College | 3 | 21 | 544 | 1.2K |
| 244 | University of Antwerp | 4 | 20 | 1.6K | 3.7K |
| 244 | Sun Yat-sen University | 1 | 20 | 5.3K | 4.6K |
| 244 | Niigata University | 4 | 20 | 1.1K | 2.0K |
| 244 | Drexel University | 1 | 20 | 3.1K | 3.1K |
| 244 | Emory University | 1 | 20 | 5.2K | 4.8K |
| 244 | Central Washington University | 1 | 20 | 355 | 388 |
| 244 | University of Calcutta | 2 | 20 | 1.3K | 1.5K |
| 244 | Nagoya Institute of Technology | 2 | 20 | 905 | 955 |
| 244 | University of Dundee | 1 | 20 | 3.0K | 3.0K |
| 244 | University of Gdańsk | 2 | 20 | 1.1K | 1.1K |
| 244 | Catholic University of Colombia | 1 | 20 | 71 | 26 |
| 244 | Chongqing Normal University | 3 | 20 | 253 | 471 |
| 244 | Universidad Externado de Colombia | 1 | 20 | 148 | 26 |
| 244 | Kinjo Gakuin University | 3 | 20 | 33 | 91 |
| 258 | University of Vienna | 2 | 19 | 4.1K | 5.0K |
| 258 | NOAA | 1 | 19 | 809 | 862 |
| 258 | University of Leeds | 1 | 19 | 6.9K | 6.2K |
| 258 | Babeș-Bolyai University | 1 | 19 | 1.3K | 1.2K |
| 258 | Lehigh University | 2 | 19 | 1.4K | 2.0K |
| 258 | Meijo University | 2 | 19 | 542 | 699 |
| 258 | Doshisha University | 4 | 19 | 367 | 786 |
| 258 | Austral University of Chile | 1 | 19 | 1.0K | 975 |
| 258 | Kyoto Institute of Technology | 4 | 19 | 502 | 861 |
| 258 | University of Paris-Sud | 1 | 19 | 2.1K | 2.3K |
| 258 | (former) Institut National de Recherche pour l'Agriculture, l'Alimentation et l'Environnement | 2 | 19 | 1.8K | 2.6K |
| 258 | ComUE Paris-Saclay University | 1 | 19 | 2.8K | 2.4K |
| 270 | University of Central Florida | 1 | 18 | 3.8K | 3.6K |
| 270 | Purdue University | 1 | 18 | 7.4K | 6.7K |
| 270 | Durham University | 1 | 18 | 4.0K | 3.7K |
| 270 | Metropolitan State University of Denver | 2 | 18 | 91 | 238 |
| 270 | University of Manitoba | 1 | 18 | 5.6K | 5.1K |
| 270 | Kyoto Sangyo University | 2 | 18 | 214 | 346 |
| 270 | Savitribai Phule Pune University | 2 | 18 | 700 | 1.1K |
| 270 | Kuwait University | 1 | 18 | 2.2K | 2.1K |
| 270 | Jawaharlal Nehru Centre for Advanced Scientific Research | 1 | 18 | 392 | 460 |
| 270 | Universidade Federal do Pampa | 1 | 18 | 688 | 405 |
| 270 | Kurume University | 2 | 18 | 408 | 661 |
| 270 | State University of Santa Cruz | 1 | 18 | 630 | 408 |
| 282 | Kansas State University | 2 | 17 | 2.5K | 3.5K |
| 282 | Ewha Womans University | 3 | 17 | 1.2K | 2.3K |
| 282 | University of Southampton | 1 | 17 | 6.1K | 5.9K |
| 282 | University of Strasbourg | 1 | 17 | 2.3K | 2.4K |
| 282 | University of Birmingham | 1 | 17 | 7.2K | 6.7K |
| 282 | Charles Sturt University | 3 | 17 | 667 | 1.7K |
| 282 | Eastern Kentucky University | 1 | 17 | 418 | 438 |
| 282 | Hubert Curien Multi-disciplinary Institute | 1 | 17 | 164 | 213 |
| 282 | Okayama University of Science | 1 | 17 | 747 | 601 |
| 282 | National University of Jujuy | 2 | 17 | 41 | 76 |
| 282 | National Institute for Agro-Environmental Sciences | 1 | 17 | 227 | 295 |
| 282 | University of Tennessee | 1 | 17 | 6.2K | 5.8K |
| 282 | Universidade Federal da Bahia | 4 | 17 | 980 | 1.8K |
| 282 | Università degli Studi di Milano Bicocca | 1 | 17 | 2.0K | 2.2K |
| 296 | University of London | 1 | 16 | 7.7K | 7.0K |
| 296 | Hebrew University of Jerusalem | 2 | 16 | 3.7K | 4.9K |
| 296 | Simon Fraser University | 2 | 16 | 3.1K | 4.4K |
| 296 | Université catholique de Louvain | 1 | 16 | 4.5K | 4.7K |
| 296 | University of Tehran | 1 | 16 | 4.5K | 4.1K |
| 296 | Rice University | 1 | 16 | 3.1K | 3.3K |
| 296 | University of South Carolina | 2 | 16 | 4.0K | 5.3K |
| 296 | Shiga University | 1 | 16 | 183 | 165 |
| 296 | University of Parma | 1 | 16 | 2.9K | 3.1K |
| 296 | Okinawa University | 2 | 16 | 2 | 14 |
| 296 | University of Michigan–Dearborn | 1 | 16 | 596 | 682 |
| 296 | University of Veterinary Medicine Hanover, Foundation | 1 | 16 | 490 | 631 |
| 296 | Universidad Autónoma del Estado de Morelos | 2 | 16 | 414 | 621 |
| 296 | Royal Veterinary College | 1 | 16 | 527 | 643 |
| 310 | University of Lyon | 2 | 15 | 470 | 1.2K |
| 310 | Leiden University | 1 | 15 | 4.3K | 4.3K |
| 310 | Université du Québec à Rimouski | 1 | 15 | 392 | 468 |
| 310 | United States Geological Survey | 1 | 15 | 792 | 940 |
| 310 | University of Valencia | 1 | 15 | 4.9K | 4.4K |
| 310 | City University of New York | 2 | 15 | 3.5K | 4.7K |
| 310 | Pontifícia Universidade Católica de Minas Gerais | 2 | 15 | 258 | 286 |
| 310 | Laval University | 1 | 15 | 5.2K | 5.2K |
| 310 | University of New Orleans | 1 | 15 | 1.2K | 1.4K |
| 310 | Université du Québec à Montréal | 1 | 15 | 2.3K | 2.5K |
| 310 | University of Idaho | 2 | 15 | 1.4K | 2.2K |
| 310 | Canterbury Museum | 1 | 15 | 36 | 50 |
| 310 | Instituto Superior de Psicologia Aplicada | 1 | 15 | 46 | 96 |
| 323 | Stockholm University | 1 | 14 | 4.4K | 4.5K |
| 323 | Haskell Indian Nations University | 1 | 14 | 4 | 12 |
| 323 | Kagawa University | 6 | 14 | 339 | 1.3K |
| 323 | Coventry University | 2 | 14 | 1.0K | 1.9K |
| 323 | Japan Women's University | 1 | 14 | 188 | 241 |
| 323 | Kumamoto University | 5 | 14 | 1.0K | 2.4K |
| 323 | University of Hawaii at Manoa | 1 | 14 | 2.3K | 2.8K |
| 323 | Louis Pasteur University | 1 | 14 | 1.2K | 1.6K |
| 323 | Kobe College | 1 | 14 | 45 | 48 |
| 323 | Universidade Federal de Sergipe | 1 | 14 | 1.4K | 976 |
| 323 | Fundación Universitaria Konrad Lorenz | 1 | 14 | 47 | 45 |
| 323 | National Geographic Society | 1 | 14 | 43 | 82 |
| 335 | University of Arizona | 1 | 13 | 7.7K | 7.6K |
| 335 | University of Oldenburg | 1 | 13 | 1.1K | 1.3K |
| 335 | The University of Tampa | 1 | 13 | 293 | 430 |
| 335 | University of Texas at Arlington | 1 | 13 | 2.4K | 2.6K |
| 335 | Sichuan University | 1 | 13 | 4.9K | 4.7K |
| 335 | Smithsonian Environmental Research Center | 1 | 13 | 201 | 334 |
| 335 | Federal University of Rio Grande do Norte | 6 | 13 | 636 | 1.9K |
| 335 | Natural History Museum of Crete | 1 | 13 | 36 | 62 |
| 335 | Shobi University | 1 | 13 | 4 | 4 |
| 344 | Leipzig University | 1 | 12 | 3.9K | 4.2K |
| 344 | Georgetown University | 1 | 12 | 3.5K | 3.7K |
| 344 | University of Botswana | 2 | 12 | 491 | 920 |
| 344 | University of Waikato | 2 | 12 | 1.2K | 2.2K |
| 344 | University of New South Wales | 1 | 12 | 8.2K | 8.3K |
| 344 | College of Wooster | 1 | 12 | 185 | 268 |
| 344 | Wildlife Conservation Society | 1 | 12 | 180 | 297 |
| 344 | University of Southern Denmark | 1 | 12 | 3.2K | 3.7K |
| 344 | University of Toronto Mississauga | 1 | 12 | 675 | 968 |
| 344 | Ohio State University, Newark Campus | 1 | 12 | 43 | 54 |
| 344 | National Museum of Nature and Science | 2 | 12 | 148 | 240 |
| 344 | Botswana Predator Conservation Trust | 1 | 12 | 16 | 29 |
| 356 | University of Texas at Tyler | 1 | 11 | 345 | 500 |
| 356 | Ruhr University Bochum | 1 | 11 | 4.1K | 4.4K |
| 356 | University of Bayreuth | 3 | 11 | 1.0K | 2.2K |
| 356 | Chulalongkorn University | 2 | 11 | 2.2K | 3.4K |
| 356 | Charles Darwin University | 1 | 11 | 919 | 1.3K |
| 356 | University of Toyama | 1 | 11 | 2.2K | 2.3K |
| 356 | Shinshu University | 1 | 11 | 2.0K | 2.1K |
| 356 | Stetson University | 1 | 11 | 183 | 250 |
| 356 | National Institute for Environmental Studies | 2 | 11 | 641 | 1.1K |
| 356 | Rottenburg University of Applied Forest Sciences | 1 | 11 | 9 | 15 |
| 356 | University of Hyogo | 1 | 11 | 1.2K | 1.2K |
| 367 | University of New Hampshire | 1 | 10 | 2.4K | 2.9K |
| 367 | University of Łódź | 1 | 10 | 1.8K | 1.7K |
| 367 | Sapienza University of Rome | 1 | 10 | 6.2K | 6.5K |
| 367 | United States Fish and Wildlife Service | 1 | 10 | 136 | 212 |
| 367 | Bedford Institute of Oceanography | 1 | 10 | 299 | 407 |
| 367 | University of Massachusetts | 1 | 10 | 5.6K | 6.2K |
| 367 | Senshu University | 1 | 10 | 161 | 169 |
| 367 | University of Delaware | 1 | 10 | 4.2K | 4.8K |
| 367 | University of Richmond | 1 | 10 | 694 | 881 |
| 367 | University of Costa Rica | 2 | 10 | 607 | 1.3K |
| 367 | University of Nebraska–Lincoln | 1 | 10 | 3.3K | 4.0K |
| 367 | Fukuoka Institute of Technology | 1 | 10 | 302 | 293 |
| 367 | Mississippi State University | 1 | 10 | 2.8K | 3.2K |
| 367 | St. Francis Xavier University | 1 | 10 | 660 | 895 |
| 367 | Venezuelan Institute for Scientific Research | 1 | 10 | 544 | 699 |
| 367 | Madurai Kamaraj University | 2 | 10 | 409 | 791 |
| 367 | Bialystok University of Technology | 1 | 10 | 450 | 483 |
| 367 | Hyogo College of Medicine | 2 | 10 | 848 | 1.3K |
| 367 | Patuxent Wildlife Research Center | 1 | 10 | 149 | 212 |
| 367 | Siedlce University of Natural Sciences and Humanities | 1 | 10 | 228 | 156 |
| 387 | University of St Andrews | 1 | 9 | 2.2K | 2.7K |
| 387 | University of Malawi | 1 | 9 | 458 | 686 |
| 387 | Kwansei Gakuin University | 2 | 9 | 426 | 794 |
| 387 | Edinburgh Napier University | 1 | 9 | 991 | 1.4K |
| 387 | King Saud University | 1 | 9 | 4.5K | 5.0K |
| 387 | Royal Zoological Society of Scotland | 1 | 9 | 26 | 59 |
| 387 | Fuji Tokoha University | 1 | 9 | 6 | 12 |
| 387 | National University of General San Martín | 1 | 9 | 343 | 398 |
| 387 | National University of Luján | 1 | 9 | 127 | 183 |
| 387 | Claude Bernard University Lyon 1 | 1 | 9 | 2.6K | 3.3K |
| 397 | National Taiwan University | 2 | 8 | 4.2K | 5.9K |
| 397 | University of Connecticut | 1 | 8 | 6.0K | 6.8K |
| 397 | University of Göttingen | 1 | 8 | 4.3K | 5.1K |
| 397 | University of Toronto | 2 | 8 | 9.5K | 11.8K |
| 397 | École nationale vétérinaire d'Alfort | 1 | 8 | 175 | 312 |
| 397 | Franklin & Marshall College | 1 | 8 | 317 | 511 |
| 397 | University of Extremadura | 1 | 8 | 2.1K | 2.5K |
| 397 | Ryukoku University | 1 | 8 | 488 | 539 |
| 397 | University of Colorado | 1 | 8 | 6.2K | 7.1K |
| 397 | University of Fort Hare | 1 | 8 | 567 | 704 |
| 397 | Catholic University of the North | 1 | 8 | 594 | 808 |
| 397 | Huazhong Agricultural University | 1 | 8 | 1.5K | 1.9K |
| 397 | Arturo Prat University | 1 | 8 | 152 | 215 |
| 397 | University of Paris-Est | 1 | 8 | 58 | 181 |
| 397 | Bharathidasan University | 1 | 8 | 758 | 953 |
| 397 | Chiba Institute of Science | 1 | 8 | 100 | 190 |
| 397 | Kyoto University of Advanced Science | 1 | 8 | 93 | 112 |
| 397 | list of women's colleges | 1 | 8 | 62 | 140 |
| 397 | Spanish National Research Council | 1 | 8 | 1.6K | 2.3K |
| 416 | Russian Academy of Sciences | 1 | 7 | 5.8K | 5.7K |
| 416 | University of Tartu | 1 | 7 | 2.5K | 3.3K |
| 416 | Wageningen University & Research | 1 | 7 | 3.1K | 4.1K |
| 416 | University of Turku | 1 | 7 | 4.1K | 5.0K |
| 416 | Bangor University | 1 | 7 | 1.7K | 2.4K |
| 416 | Naruto University of Education | 1 | 7 | 146 | 195 |
| 416 | Colorado State University | 1 | 7 | 4.7K | 5.8K |
| 416 | University of Exeter | 1 | 7 | 4.4K | 5.3K |
| 416 | Bangalore University | 1 | 7 | 593 | 760 |
| 416 | University of Shiga Prefecture | 1 | 7 | 394 | 518 |
| 416 | Beijing Forestry University | 1 | 7 | 1.0K | 1.3K |
| 416 | Universidad Autónoma del Estado de Hidalgo | 1 | 7 | 534 | 667 |
| 416 | Universidade Federal de Goiás | 3 | 7 | 1.1K | 2.1K |
| 416 | North Minzu University | 1 | 7 | 224 | 294 |
| 430 | Yonsei University | 1 | 6 | 5.3K | 6.0K |
| 430 | Max Planck Society | 1 | 6 | 451 | 696 |
| 430 | Central University of Venezuela | 1 | 6 | 1.1K | 1.5K |
| 430 | Instituto Butantan | 1 | 6 | 473 | 767 |
| 430 | Umeå University | 1 | 6 | 3.9K | 5.2K |
| 430 | Xiamen University | 1 | 6 | 3.2K | 3.9K |
| 430 | Davidson College | 1 | 6 | 286 | 491 |
| 430 | Graduate University for Advanced Studies | 1 | 6 | 139 | 252 |
| 430 | Ishikawa Prefectural University | 2 | 6 | 99 | 260 |
| 430 | Hanoi University of Science | 1 | 6 | 71 | 171 |
| 430 | RIKEN Brain Science Institute | 2 | 6 | 201 | 545 |
| 430 | Federal Rural University of Pernambuco | 2 | 6 | 652 | 1.1K |
| 442 | University of Aveiro | 1 | 5 | 2.7K | 3.6K |
| 442 | McGill University | 1 | 5 | 8.9K | 10.2K |
| 442 | University of Minnesota | 1 | 5 | 10.0K | 11.2K |
| 442 | University of Haifa | 1 | 5 | 3.0K | 4.0K |
| 442 | Kyushu Institute of Technology | 1 | 5 | 1.2K | 1.5K |
| 442 | University of North Carolina | 2 | 5 | 5.0K | 7.9K |
| 442 | Missouri State University | 1 | 5 | 947 | 1.5K |
| 442 | University of Kent | 1 | 5 | 2.9K | 3.9K |
| 442 | Miyagi University of Education | 1 | 5 | 133 | 217 |
| 442 | Université Sainte-Anne | 1 | 5 | 16 | 31 |
| 442 | Capital Normal University | 1 | 5 | 1.1K | 1.5K |
| 442 | Southwest Minnesota State University | 1 | 5 | 137 | 154 |
| 442 | Centro Universitário de Brasília | 1 | 5 | 65 | 69 |
| 442 | Mato Grosso State University | 1 | 5 | 176 | 235 |
| 442 | Ohio State University, Lima Campus | 1 | 5 | 16 | 52 |
| 442 | Universidade Estadual de Ponta Grossa | 1 | 5 | 530 | 542 |
| 458 | Autonomous University of Barcelona | 1 | 4 | 5.0K | 6.4K |
| 458 | University of Warsaw | 1 | 4 | 3.3K | 3.9K |
| 458 | University of Pavia | 2 | 4 | 3.0K | 5.3K |
| 458 | University of Michigan | 2 | 4 | 8.6K | 12.0K |
| 458 | China Agricultural University | 1 | 4 | 2.1K | 2.7K |
| 458 | Pennsylvania State University | 1 | 4 | 9.3K | 10.7K |
| 458 | Shimane University | 1 | 4 | 1.5K | 2.0K |
| 458 | Indiana University Bloomington | 1 | 4 | 1.7K | 2.8K |
| 458 | Maharshi Dayanand University | 1 | 4 | 464 | 774 |
| 458 | University of Liège | 1 | 4 | 3.9K | 5.2K |
| 458 | University of Western Australia | 2 | 4 | 4.0K | 7.1K |
| 458 | Nara University of Education | 2 | 4 | 87 | 248 |
| 458 | Nicolaus Copernicus University in Toruń | 1 | 4 | 1.8K | 2.3K |
| 458 | Universidad Veracruzana | 1 | 4 | 965 | 1.3K |
| 458 | Virginia Commonwealth University | 1 | 4 | 4.7K | 6.0K |
| 458 | Wenzhou University | 1 | 4 | 866 | 1.3K |
| 458 | Universidad de Córdoba | 1 | 4 | 1.4K | 1.9K |
| 475 | Stanford University | 2 | 3 | 6.7K | 10.7K |
| 475 | University of Tübingen | 2 | 3 | 3.4K | 6.2K |
| 475 | University of Bergen | 1 | 3 | 4.1K | 5.8K |
| 475 | University of Basel | 1 | 3 | 3.7K | 5.2K |
| 475 | University of British Columbia | 1 | 3 | 10.0K | 11.9K |
| 475 | Inha University | 1 | 3 | 2.0K | 3.0K |
| 475 | Kyung Hee University | 1 | 3 | 3.1K | 4.3K |
| 475 | Incheon National University | 1 | 3 | 736 | 1.2K |
| 475 | National Institute of Genetics | 1 | 3 | 425 | 709 |
| 475 | Åbo Akademi University | 1 | 3 | 1.3K | 2.3K |
| 475 | Hokkaido University of Education | 1 | 3 | 213 | 367 |
| 475 | Clermont Auvergne University | 1 | 3 | 929 | 1.7K |
| 475 | University of Texas at El Paso | 1 | 3 | 1.8K | 2.9K |
| 475 | Northeastern State University | 1 | 3 | 91 | 226 |
| 475 | Pacific Union College | 1 | 3 | 17 | 35 |
| 475 | Universidad Nacional del Comahue | 1 | 3 | 380 | 645 |
| 475 | Adelphi University | 1 | 3 | 602 | 1.1K |
| 475 | Universidade Federal de Alagoas | 2 | 3 | 722 | 1.5K |
| 475 | Shahrekord University | 1 | 3 | 620 | 1.1K |
| 475 | Korea Polar Research Institute | 1 | 3 | 269 | 460 |
| 475 | Western Australian Museum | 2 | 3 | 81 | 218 |
| 496 | Sungkyunkwan University | 1 | 2 | 4.3K | 5.5K |
| 496 | Makerere University | 1 | 2 | 1.3K | 2.3K |
| 496 | Waseda University | 2 | 2 | 2.1K | 4.3K |
| 496 | Brigham Young University | 1 | 2 | 3.4K | 5.0K |
| 496 | University of Gothenburg | 1 | 2 | 5.6K | 7.7K |
| 496 | Clemson University | 1 | 2 | 3.7K | 5.5K |
| 496 | Université Cheikh Anta Diop | 1 | 2 | 447 | 911 |
| 496 | Dokkyo Medical University | 1 | 2 | 522 | 873 |
| 496 | University of Rennes 1 | 2 | 2 | 1.3K | 3.1K |
| 496 | University of Oregon | 1 | 2 | 3.2K | 4.7K |
| 496 | University of Tasmania | 2 | 2 | 2.4K | 5.2K |
| 496 | University of Maine | 1 | 2 | 2.0K | 3.3K |
| 496 | University of Louisville | 1 | 2 | 4.1K | 6.1K |
| 496 | University College Cork | 1 | 2 | 2.7K | 4.5K |
| 496 | Hampshire College | 1 | 2 | 127 | 360 |
| 496 | University of Northern Colorado | 1 | 2 | 706 | 1.3K |
| 496 | United States Forest Service | 1 | 2 | 838 | 1.3K |
| 496 | Aichi Gakuin University | 1 | 2 | 270 | 529 |
| 496 | Azim Premji University | 1 | 2 | 64 | 153 |
| 496 | DeSales University | 1 | 2 | 33 | 90 |
| 496 | Periyar University | 1 | 2 | 425 | 723 |
| 496 | Hopkins Marine Station | 1 | 2 | 152 | 291 |
| 496 | Taiwan Forestry Research Institute | 1 | 2 | 98 | 194 |
| 496 | Hospital General Universitario Gregorio Marañón | 1 | 2 | 822 | 1.3K |
| 496 | University of Nevada, Reno | 1 | 2 | 4.3K | 6.2K |
| 496 | Universidade Estadual de Londrina | 1 | 2 | 1.2K | 1.5K |
| 496 | University of Rzeszow | 1 | 2 | 797 | 1.2K |
| 523 | University of Coimbra | 1 | 1 | 3.6K | 5.5K |
| 523 | Tours University | 1 | 1 | 826 | 1.7K |
| 523 | National Taiwan Normal University | 1 | 1 | 1.4K | 2.5K |
| 523 | Yamaguchi University | 1 | 1 | 1.8K | 2.7K |
| 523 | Western Sydney University | 1 | 1 | 2.8K | 4.9K |
| 523 | Hofstra University | 1 | 1 | 864 | 1.8K |
| 523 | Okinawa International University | 1 | 1 | 18 | 55 |
| 523 | Kyoto Women's University | 4 | 1 | 40 | 297 |
| 523 | Archbold Biological Station | 1 | 1 | 63 | 124 |
| 523 | Chinese Academy of Fishery Sciences | 1 | 1 | 475 | 813 |
| 523 | Kashi University | 1 | 1 | 35 | 134 |
| 523 | Nakhon Si Thammarat Rajabhat University | 1 | 1 | 15 | 52 |
| 523 | PanThera Biopharma | 1 | 1 | 6 | 25 |
| 536 | University of Aberdeen | 1 | 0 | 4.4K | 6.9K |
| 536 | University of A Coruña | 1 | 0 | 1.4K | 2.7K |
| 536 | Tribhuvan University | 1 | 0 | 950 | 1.9K |
| 536 | Osaka University | 1 | 0 | 5.3K | 7.3K |
| 536 | Osaka Gakuin University | 1 | 0 | 40 | 105 |
| 536 | University of Veterinary Medicine Vienna | 1 | 0 | 469 | 920 |
| 536 | University of Kentucky | 1 | 0 | 6.3K | 9.2K |
| 536 | Universidad de las Américas Puebla | 1 | 0 | 186 | 530 |
| 536 | National Institute of Amazonian Research | 1 | 0 | 402 | 708 |
| 536 | Autonomous University of Tlaxcala | 1 | 0 | 166 | 405 |
| 536 | Universidade Federal de Mato Grosso do Sul | 1 | 0 | 1.1K | 1.7K |
| 536 | Autonomous University of Guerrero | 1 | 0 | 243 | 610 |
| 536 | Montes Claros State University | 1 | 0 | 309 | 506 |
| 536 | Escola Superior de Educação de Paula Frassinetti | 1 | 0 | 3 | 7 |
| 536 | University of California, Los Angeles | 1 | 0 | 5.5K | 8.3K |
| 536 | University of California, Riverside | 1 | 0 | 1.7K | 3.3K |
| 536 | Universidade do Porto | 1 | 0 | 2.4K | 4.2K |
