| # | Institution | Articles | Citations | R ARank | R CRank |
|---|
| 1 | Swedish Museum of Natural History | 75 | 1,962 | 6 | 20 |
| 2 | University of Gothenburg | 64 | 1,335 | 152 | 448 |
| 3 | University of Sydney | 9 | 1,319 | 3.1K | 869 |
| 4 | University of Copenhagen | 46 | 1,115 | 506 | 875 |
| 5 | University of Oxford | 6 | 1,093 | 4.3K | 1.3K |
| 6 | Uppsala University | 23 | 1,010 | 812 | 739 |
| 7 | Natural History Museum | 41 | 769 | 91 | 155 |
| 8 | Muséum national d'histoire naturelle | 23 | 507 | 59 | 124 |
| 9 | University of Queensland | 21 | 470 | 1.4K | 1.9K |
| 10 | Smithsonian Institution | 13 | 432 | 155 | 157 |
| 11 | Chinese Academy of Sciences | 30 | 417 | 2.1K | 2.5K |
| 12 | Russian Academy of Sciences | 15 | 414 | 1.6K | 1.1K |
| 13 | University of Oslo | 24 | 338 | 624 | 1.5K |
| 14 | Harvard University | 11 | 326 | 1.8K | 2.3K |
| 15 | Stockholm University | 18 | 304 | 513 | 1.1K |
| 16 | University of California | 13 | 285 | 7.1K | 6.8K |
| 17 | University of Bergen | 21 | 282 | 409 | 1.1K |
| 18 | American Museum of Natural History | 11 | 275 | 102 | 144 |
| 19 | University of São Paulo | 17 | 265 | 2.2K | 2.4K |
| 20 | Lund University | 18 | 261 | 1.2K | 2.1K |
| 21 | University of Vienna | 12 | 258 | 1.2K | 1.5K |
| 22 | National Center for Scientific Research | 16 | 252 | 2.8K | 3.2K |
| 23 | Universidade Estadual Paulista | 10 | 242 | 803 | 559 |
| 24 | McGill University | 3 | 229 | 5.7K | 3.0K |
| 25 | Ocean University of China | 11 | 227 | 415 | 445 |
| 26 | Naturalis Biodiversity Center | 6 | 213 | 57 | 40 |
| 27 | University of Turku | 5 | 211 | 1.5K | 1.1K |
| 28 | Nanjing Agricultural University | 8 | 207 | 534 | 508 |
| 29 | California Academy of Sciences | 8 | 198 | 45 | 55 |
| 30 | Griffith University | 5 | 197 | 1.7K | 1.3K |
| 31 | Macdonald Campus | 1 | 182 | 137 | 31 |
| 32 | Hebrew University of Jerusalem | 10 | 181 | 1.1K | 1.7K |
| 33 | University of Barcelona | 10 | 179 | 1.6K | 2.0K |
| 34 | Australian Museum | 12 | 178 | 29 | 73 |
| 35 | Polish Academy of Sciences | 7 | 175 | 1.7K | 1.6K |
| 36 | Universidade Federal do Paraná | 8 | 173 | 790 | 531 |
| 37 | University of Georgia | 6 | 169 | 2.5K | 2.3K |
| 38 | University of North Carolina at Greensboro | 1 | 162 | 1.7K | 450 |
| 38 | Mongolian Academy of Sciences | 2 | 162 | 121 | 38 |
| 40 | University of Florida | 11 | 159 | 2.8K | 3.9K |
| 41 | University of Sheffield | 8 | 158 | 2.1K | 2.5K |
| 41 | University of North Carolina | 4 | 158 | 3.3K | 2.7K |
| 43 | Norwegian University of Science and Technology | 7 | 156 | 1.2K | 1.6K |
| 44 | Università degli Studi di Milano Bicocca | 5 | 154 | 502 | 513 |
| 45 | University of Michigan | 5 | 152 | 5.7K | 4.9K |
| 45 | University of Oulu | 3 | 152 | 1.9K | 1.2K |
| 47 | University of Salzburg | 6 | 149 | 248 | 395 |
| 48 | Swansea University | 5 | 147 | 1.0K | 982 |
| 49 | National Taiwan Ocean University | 8 | 143 | 223 | 332 |
| 50 | University of Crete | 8 | 141 | 401 | 730 |
| 51 | University of Valencia | 7 | 139 | 1.4K | 1.6K |
| 52 | Moscow State University | 8 | 137 | 1.3K | 1.2K |
| 53 | Natural History Museum of Los Angeles County | 5 | 133 | 46 | 46 |
| 54 | Austral University of Chile | 6 | 130 | 242 | 262 |
| 54 | Natural History Museum of Crete | 6 | 130 | 6 | 8 |
| 56 | Nankai University | 6 | 128 | 1.0K | 958 |
| 57 | University of Miyazaki | 2 | 127 | 1.2K | 474 |
| 58 | University of Louisiana at Lafayette | 3 | 125 | 427 | 271 |
| 58 | La Trobe University | 2 | 125 | 2.8K | 1.5K |
| 58 | (former) Institut National de Recherche pour l'Agriculture, l'Alimentation et l'Environnement | 5 | 125 | 1.1K | 1.3K |
| 61 | University of Brasília | 5 | 124 | 1.2K | 716 |
| 62 | National University of Singapore | 8 | 122 | 2.7K | 3.5K |
| 62 | University of Innsbruck | 8 | 122 | 686 | 1.2K |
| 62 | Commonwealth Scientific and Industrial Research Organisation | 1 | 122 | 2.0K | 975 |
| 65 | University of Helsinki | 6 | 119 | 2.6K | 3.0K |
| 65 | Comenius University in Bratislava | 4 | 119 | 154 | 89 |
| 65 | Liverpool John Moores University | 9 | 119 | 277 | 675 |
| 68 | Czech Academy of Sciences | 5 | 115 | 1.6K | 1.5K |
| 69 | Stellenbosch University | 9 | 108 | 775 | 1.4K |
| 70 | Australian National University | 2 | 107 | 4.1K | 2.6K |
| 71 | Ruhr University Bochum | 1 | 106 | 4.1K | 1.8K |
| 71 | Auburn University | 6 | 106 | 1.3K | 1.6K |
| 73 | Paris Diderot University | 6 | 104 | 644 | 1.1K |
| 74 | Osnabrück University | 6 | 102 | 326 | 536 |
| 75 | University of Adelaide | 5 | 99 | 2.4K | 2.9K |
| 75 | Tohoku University | 2 | 99 | 4.0K | 2.4K |
| 77 | Villanova University | 4 | 97 | 425 | 438 |
| 78 | University of Hamburg | 8 | 96 | 1.0K | 1.9K |
| 79 | Ghent University | 7 | 95 | 2.4K | 3.4K |
| 80 | University of Kansas | 5 | 94 | 2.3K | 2.7K |
| 81 | University of South Bohemia in České Budějovice | 3 | 93 | 398 | 361 |
| 82 | University of Alcalá | 4 | 92 | 856 | 865 |
| 83 | Carnegie Museum of Natural History | 2 | 91 | 62 | 50 |
| 84 | National Institute of Amazonian Research | 2 | 90 | 282 | 169 |
| 84 | Senckenberg Natural History Collections Dresden | 2 | 90 | 27 | 17 |
| 86 | National Scientific and Technical Research Council | 7 | 88 | 539 | 906 |
| 87 | University of Amsterdam | 5 | 86 | 3.3K | 3.8K |
| 88 | University of Siena | 4 | 85 | 1.3K | 1.5K |
| 89 | Charles University | 5 | 83 | 1.9K | 2.1K |
| 90 | National Chung Hsing University | 1 | 81 | 2.2K | 1.0K |
| 90 | Santa Barbara Museum of Natural History | 1 | 81 | 37 | 17 |
| 92 | Federal University of Rio de Janeiro | 4 | 80 | 2.4K | 2.0K |
| 92 | Macquarie University | 3 | 80 | 2.3K | 2.1K |
| 92 | University of Guelph | 4 | 80 | 1.8K | 2.1K |
| 92 | Federal University of ABC | 3 | 80 | 374 | 243 |
| 92 | Royal Ontario Museum | 2 | 80 | 123 | 100 |
| 92 | Queensland Museum | 5 | 80 | 28 | 55 |
| 98 | University of Ljubljana | 3 | 79 | 2.2K | 1.9K |
| 98 | Natural History Museum of Geneva | 5 | 79 | 9 | 19 |
| 100 | University of New England | 6 | 78 | 441 | 971 |
| 100 | University of Porto | 3 | 78 | 1.8K | 1.6K |
| 102 | Chinese University of Hong Kong | 3 | 77 | 3.7K | 3.3K |
| 103 | University of Arizona | 2 | 76 | 5.8K | 4.3K |
| 103 | Binghamton University | 3 | 76 | 761 | 796 |
| 105 | Complutense University of Madrid | 2 | 75 | 4.3K | 2.6K |
| 105 | Venezuelan Institute for Scientific Research | 1 | 75 | 544 | 239 |
| 105 | Universidad Nacional de San Antonio Abad del Cusco | 1 | 75 | 71 | 17 |
| 108 | James Cook University | 2 | 74 | 1.5K | 1.2K |
| 109 | Chungbuk National University | 3 | 72 | 937 | 858 |
| 109 | Chonnam National University | 3 | 72 | 1.3K | 1.1K |
| 111 | University of Minnesota | 2 | 71 | 8.0K | 6.1K |
| 111 | Canadian Food Inspection Agency | 1 | 71 | 179 | 118 |
| 113 | University of Bonn | 3 | 70 | 2.7K | 2.7K |
| 113 | Hokkaido University | 5 | 70 | 2.5K | 2.7K |
| 115 | University of Toronto | 4 | 68 | 7.1K | 7.0K |
| 115 | National Autonomous University of Mexico | 5 | 68 | 2.3K | 2.4K |
| 115 | King Abdullah University of Science and Technology | 2 | 68 | 967 | 828 |
| 115 | University of Parma | 1 | 68 | 2.9K | 1.6K |
| 115 | University of Iowa | 2 | 68 | 4.9K | 3.8K |
| 120 | University of Naples Federico II | 3 | 67 | 2.9K | 2.9K |
| 120 | University of Mons | 3 | 67 | 186 | 261 |
| 122 | University of Oldenburg | 7 | 66 | 193 | 603 |
| 123 | Heidelberg University | 2 | 65 | 4.1K | 3.4K |
| 124 | University of Glasgow | 3 | 64 | 3.9K | 3.7K |
| 124 | Texas Tech University | 1 | 64 | 3.8K | 1.9K |
| 124 | Oklahoma State University | 2 | 64 | 2.5K | 2.0K |
| 127 | National Taiwan Normal University | 3 | 62 | 650 | 672 |
| 127 | University of the South Pacific | 1 | 62 | 436 | 192 |
| 127 | De La Salle University | 1 | 62 | 401 | 136 |
| 127 | Universidade Federal do Pará | 5 | 62 | 493 | 569 |
| 127 | Hungarian Natural History Museum | 3 | 62 | 67 | 78 |
| 132 | University of Tokyo | 3 | 60 | 5.9K | 5.1K |
| 133 | Iowa State University | 2 | 58 | 4.0K | 3.3K |
| 133 | University of Chinese Academy of Sciences | 4 | 58 | 1.8K | 1.8K |
| 135 | National Science Museum | 1 | 57 | 149 | 93 |
| 136 | Brigham Young University | 2 | 56 | 2.3K | 2.0K |
| 136 | Randolph–Macon College | 2 | 56 | 36 | 44 |
| 138 | University of Texas at Austin | 2 | 55 | 7.9K | 6.4K |
| 138 | Palacký University Olomouc | 2 | 55 | 939 | 806 |
| 138 | University of California, San Diego | 3 | 55 | 3.4K | 3.9K |
| 138 | Universidad de la República | 4 | 55 | 526 | 757 |
| 142 | University of Perpignan | 1 | 54 | 253 | 162 |
| 142 | California State Polytechnic University, Pomona | 3 | 54 | 213 | 344 |
| 142 | Natural History Museum of Bern | 3 | 54 | 13 | 28 |
| 145 | Hungarian Academy of Sciences | 2 | 53 | 2.2K | 2.0K |
| 145 | Inha University | 2 | 53 | 1.4K | 1.2K |
| 145 | National University of Kharkiv | 2 | 53 | 330 | 162 |
| 145 | Staatliches Museum für Naturkunde Stuttgart | 2 | 53 | 46 | 51 |
| 149 | Akdeniz University | 3 | 52 | 704 | 742 |
| 150 | Hasselt University | 4 | 51 | 374 | 787 |
| 150 | Indonesian Institute of Sciences | 2 | 51 | 214 | 126 |
| 152 | Pennsylvania State University | 1 | 50 | 9.3K | 6.1K |
| 152 | George Mason University | 2 | 50 | 2.4K | 2.1K |
| 152 | University of Peradeniya | 2 | 50 | 497 | 447 |
| 152 | Australian National Wildlife Collection | 2 | 50 | 20 | 26 |
| 156 | Ewha Womans University | 2 | 49 | 1.5K | 1.4K |
| 156 | San José State University | 1 | 49 | 1.5K | 766 |
| 156 | University of Patras | 4 | 49 | 1.2K | 1.8K |
| 159 | Saint Louis University | 3 | 48 | 1.1K | 1.5K |
| 159 | University of the Ryukyus | 2 | 48 | 1.2K | 947 |
| 159 | Virginia Commonwealth University | 2 | 48 | 3.4K | 3.1K |
| 162 | Xi'an Jiaotong University | 1 | 47 | 3.6K | 2.3K |
| 162 | Swedish University of Agricultural Sciences | 3 | 47 | 1.1K | 1.5K |
| 162 | University of Hyogo | 2 | 47 | 808 | 609 |
| 162 | Institut Agronomique et Veterinaire Hassan II | 2 | 47 | 63 | 73 |
| 166 | University of Aveiro | 2 | 46 | 1.8K | 1.7K |
| 166 | University of Bologna | 2 | 46 | 5.2K | 4.5K |
| 166 | University of Kiel | 3 | 46 | 1.0K | 1.7K |
| 166 | University of Genoa | 3 | 46 | 2.5K | 2.8K |
| 166 | Vrije Universiteit Brussel | 3 | 46 | 1.7K | 2.3K |
| 166 | Consiglio Nazionale delle Ricerche | 2 | 46 | 1.3K | 1.5K |
| 172 | University of Pisa | 3 | 45 | 3.2K | 3.6K |
| 172 | Marche Polytechnic University | 2 | 45 | 1.7K | 1.8K |
| 172 | Museu Paraense Emílio Goeldi | 7 | 45 | 53 | 144 |
| 175 | University of Concepción | 3 | 44 | 935 | 1.0K |
| 175 | University of Waikato | 1 | 44 | 1.8K | 1.2K |
| 175 | Masaryk University | 4 | 44 | 965 | 1.5K |
| 175 | Ohio University | 2 | 44 | 1.7K | 1.6K |
| 175 | University of Maine | 2 | 44 | 1.3K | 1.4K |
| 175 | South Australian Museum | 1 | 44 | 139 | 106 |
| 175 | Florida Museum of Natural History | 5 | 44 | 102 | 225 |
| 182 | The University of Tampa | 1 | 43 | 293 | 179 |
| 182 | Federal University of Rio Grande do Sul | 4 | 43 | 2.4K | 2.7K |
| 182 | King Saud University | 4 | 43 | 2.1K | 2.9K |
| 185 | Bielefeld University | 4 | 42 | 1.0K | 1.7K |
| 185 | University of Cambridge | 1 | 42 | 9.0K | 6.4K |
| 185 | Pierre and Marie Curie University | 2 | 42 | 2.0K | 2.3K |
| 185 | Prince of Songkla University | 2 | 42 | 1.1K | 954 |
| 189 | Scripps Institution of Oceanography | 3 | 41 | 234 | 413 |
| 189 | Royal Belgian Institute of Natural Sciences | 4 | 41 | 82 | 191 |
| 189 | National Museums Northern Ireland | 1 | 41 | 11 | 11 |
| 192 | Oberlin College | 1 | 40 | 502 | 341 |
| 192 | University of Campinas | 3 | 40 | 2.7K | 2.7K |
| 192 | Catholic University of the North | 2 | 40 | 338 | 362 |
| 192 | Natural History Museum Vienna | 1 | 40 | 92 | 56 |
| 192 | Northeastern University | 5 | 40 | 2.0K | 3.5K |
| 192 | Monash University Malaysia | 1 | 40 | 584 | 394 |
| 198 | University of Szeged | 1 | 39 | 2.3K | 1.6K |
| 198 | University of León | 1 | 39 | 1.2K | 784 |
| 198 | Czechoslovak Academy of Sciences | 2 | 39 | 726 | 628 |
| 198 | National Food Chain Safety Office | 1 | 39 | 68 | 39 |
| 198 | Korea Polar Research Institute | 2 | 39 | 173 | 200 |
| 198 | Universidad Politécnica de Valencia | 1 | 39 | 1.1K | 920 |
| 204 | Boston University | 1 | 38 | 5.6K | 4.1K |
| 204 | University of Alabama | 2 | 38 | 3.7K | 3.6K |
| 204 | University of Delhi | 1 | 38 | 3.1K | 1.8K |
| 204 | University of the Basque Country | 2 | 38 | 1.9K | 1.9K |
| 204 | Canadian Association for Suicide Prevention | 1 | 38 | 45 | 36 |
| 204 | Universidade do Porto | 4 | 38 | 909 | 1.7K |
| 210 | University of Nottingham | 3 | 37 | 4.1K | 5.1K |
| 210 | Marine Science Institute | 5 | 37 | 59 | 201 |
| 212 | Yale University | 1 | 36 | 6.6K | 5.1K |
| 212 | University of Göttingen | 3 | 36 | 2.5K | 3.4K |
| 212 | Nanjing University | 1 | 36 | 3.7K | 2.8K |
| 212 | China University of Geosciences | 1 | 36 | 1.6K | 1.2K |
| 212 | University of Calabria | 2 | 36 | 1.4K | 1.5K |
| 212 | Maharishi International University | 2 | 36 | 31 | 62 |
| 218 | Michigan State University | 3 | 35 | 5.0K | 5.9K |
| 218 | University of Turin | 2 | 35 | 4.1K | 4.1K |
| 218 | University of Maryland | 2 | 35 | 6.9K | 6.6K |
| 218 | National Academy of Sciences of Ukraine | 1 | 35 | 2.0K | 1.3K |
| 222 | University of Bordeaux | 1 | 34 | 2.6K | 1.9K |
| 222 | University of Łódź | 2 | 34 | 1.2K | 956 |
| 222 | University of the Western Cape | 3 | 34 | 608 | 926 |
| 222 | University of Murcia | 3 | 34 | 1.5K | 2.0K |
| 222 | Universidade Federal do Amazonas | 2 | 34 | 551 | 405 |
| 222 | Universidad Nacional Mayor de San Marcos | 2 | 34 | 361 | 231 |
| 228 | Hohai University | 1 | 33 | 1.6K | 1.1K |
| 228 | Far Eastern Federal University | 2 | 33 | 349 | 237 |
| 228 | North Carolina Central University | 1 | 33 | 412 | 308 |
| 228 | Federal University of Western Pará | 1 | 33 | 229 | 80 |
| 232 | University of Utah | 2 | 32 | 5.1K | 5.3K |
| 232 | University of Pavia | 2 | 32 | 3.0K | 3.2K |
| 232 | Chulalongkorn University | 3 | 32 | 1.6K | 2.2K |
| 232 | Bulgarian Academy of Sciences | 2 | 32 | 1.7K | 1.6K |
| 232 | Finnish Museum of Natural History | 2 | 32 | 75 | 124 |
| 237 | University of Graz | 2 | 31 | 1.7K | 2.1K |
| 237 | University of Rennes 1 | 1 | 31 | 1.8K | 1.6K |
| 237 | Boğaziçi University | 1 | 31 | 1.4K | 1.1K |
| 237 | University of the Balearic Islands | 3 | 31 | 534 | 957 |
| 241 | University of Basel | 1 | 30 | 3.7K | 3.2K |
| 241 | Pontifical Catholic University of Chile | 1 | 30 | 3.1K | 2.3K |
| 241 | Ege University | 2 | 30 | 1.7K | 1.7K |
| 241 | University of Insubria | 2 | 30 | 949 | 1.3K |
| 241 | Weber State University | 2 | 30 | 157 | 232 |
| 241 | University of Salford | 2 | 30 | 1.3K | 1.5K |
| 241 | Huntsman Marine Science Centre | 1 | 30 | 11 | 13 |
| 241 | N.Z. Oceanographic Institute | 1 | 30 | 37 | 34 |
| 241 | National Museum of Natural Science | 2 | 30 | 38 | 74 |
| 250 | University of Belgrade | 1 | 29 | 4.4K | 3.1K |
| 250 | University of Molise | 1 | 29 | 594 | 623 |
| 250 | PSL University | 1 | 29 | 1.3K | 1.1K |
| 250 | Beijing Normal University | 2 | 29 | 2.2K | 2.3K |
| 250 | University of Tennessee at Chattanooga | 2 | 29 | 197 | 305 |
| 250 | Sorbonne Universités | 3 | 29 | 601 | 1.3K |
| 250 | Geological Survey of Sweden | 1 | 29 | 56 | 63 |
| 257 | National Taiwan University | 2 | 28 | 4.2K | 4.4K |
| 257 | Federal University of Minas Gerais | 2 | 28 | 3.2K | 3.0K |
| 257 | Université de Montréal | 1 | 28 | 6.2K | 5.1K |
| 257 | Chungnam National University | 3 | 28 | 1.3K | 1.8K |
| 257 | Burdur Mehmet Akif Ersoy University | 1 | 28 | 272 | 143 |
| 257 | University of Cádiz | 2 | 28 | 961 | 1.1K |
| 263 | Cornell University | 1 | 27 | 7.6K | 6.4K |
| 263 | TU Dresden | 1 | 27 | 3.9K | 3.3K |
| 263 | York University | 1 | 27 | 4.5K | 3.6K |
| 263 | Indian Institute of Science | 2 | 27 | 2.2K | 2.3K |
| 263 | University of South Dakota | 1 | 27 | 1.0K | 931 |
| 263 | Academy of Natural Sciences of Drexel University | 2 | 27 | 94 | 169 |
| 269 | University of Padua | 4 | 26 | 3.6K | 5.6K |
| 269 | Ferdowsi University of Mashhad | 2 | 26 | 1.4K | 1.5K |
| 269 | Florida State University | 3 | 26 | 2.7K | 3.8K |
| 269 | University of Cape Town | 2 | 26 | 3.1K | 3.6K |
| 269 | University of Puget Sound | 1 | 26 | 287 | 263 |
| 269 | Umeå University | 4 | 26 | 1.8K | 3.6K |
| 269 | Linyi University | 1 | 26 | 499 | 355 |
| 269 | University of Magdalena | 1 | 26 | 194 | 60 |
| 277 | University of Washington | 2 | 25 | 8.1K | 8.5K |
| 277 | University of Florence | 1 | 25 | 5.1K | 4.4K |
| 277 | Montpellier 2 University | 1 | 25 | 654 | 795 |
| 277 | University of Alaska Anchorage | 2 | 25 | 217 | 406 |
| 277 | Estonian Academy of Sciences | 1 | 25 | 148 | 131 |
| 277 | Hakim Sabzevari University | 2 | 25 | 232 | 294 |
| 277 | Shanghai Ocean University | 1 | 25 | 652 | 590 |
| 277 | Federal University of Espírito Santo | 2 | 25 | 1.2K | 1.0K |
| 285 | Bangor University | 1 | 24 | 1.7K | 1.7K |
| 285 | Nelson Mandela University | 1 | 24 | 685 | 499 |
| 285 | University of the Sunshine Coast | 1 | 24 | 1.0K | 980 |
| 285 | Autonomous University of Baja California Sur | 2 | 24 | 82 | 101 |
| 285 | Ben Gurion University of the Negev | 2 | 24 | 965 | 1.6K |
| 290 | Ohio Wesleyan University | 1 | 23 | 155 | 134 |
| 290 | Pontifícia Universidade Católica do Rio Grande do Sul | 2 | 23 | 754 | 838 |
| 290 | University of Colorado Boulder | 1 | 23 | 3.2K | 3.1K |
| 290 | Instituto Biológico | 1 | 23 | 332 | 327 |
| 290 | Oswaldo Cruz Foundation | 2 | 23 | 1.1K | 1.4K |
| 290 | Field Museum of Natural History | 2 | 23 | 246 | 378 |
| 296 | Jagiellonian University | 1 | 22 | 3.4K | 2.6K |
| 296 | Universidade Federal Fluminense | 1 | 22 | 2.9K | 1.7K |
| 296 | Southwest University | 1 | 22 | 2.2K | 1.9K |
| 296 | Zoological Museum of Moscow University | 2 | 22 | 10 | 22 |
| 296 | Nanchang University | 2 | 22 | 1.4K | 1.7K |
| 296 | Federal University of Rio Grande do Norte | 3 | 22 | 1.3K | 1.5K |
| 296 | Huaihai Institute of Technology | 2 | 22 | 220 | 302 |
| 303 | University of Milan | 1 | 21 | 5.9K | 5.3K |
| 303 | University of Central Florida | 1 | 21 | 3.8K | 3.4K |
| 303 | Humboldt University of Berlin | 1 | 21 | 4.7K | 4.2K |
| 303 | University of the Aegean | 1 | 21 | 1.0K | 921 |
| 303 | University of Trier | 1 | 21 | 866 | 875 |
| 303 | University of North Dakota | 1 | 21 | 1.5K | 1.6K |
| 303 | Instituto Butantan | 2 | 21 | 314 | 514 |
| 303 | Southern Cross University | 1 | 21 | 936 | 999 |
| 303 | University of Akron | 1 | 21 | 2.0K | 1.9K |
| 303 | University of Exeter | 1 | 21 | 4.4K | 4.0K |
| 303 | University of Guyana | 1 | 21 | 53 | 42 |
| 303 | Pontifical Xavierian University | 1 | 21 | 1.0K | 753 |
| 303 | Idaho State University | 1 | 21 | 962 | 961 |
| 303 | Northern State University | 1 | 21 | 44 | 30 |
| 303 | North Carolina Museum of Natural Sciences | 1 | 21 | 83 | 96 |
| 303 | University of Nottingham Malaysia Campus | 1 | 21 | 171 | 273 |
| 303 | Buffalo State College | 1 | 21 | 296 | 292 |
| 303 | Federal University of Lavras | 1 | 21 | 1.1K | 797 |
| 303 | Universidade Federal da Bahia | 1 | 21 | 2.5K | 1.6K |
| 303 | University of Maryland, Baltimore County | 1 | 21 | 1.1K | 1.3K |
| 303 | National Museum Wales | 2 | 21 | 54 | 89 |
| 303 | Universidad Veritas | 1 | 21 | 3 | 1 |
| 325 | University of Nairobi | 1 | 20 | 1.4K | 1.1K |
| 325 | Oregon State University | 2 | 20 | 3.2K | 4.3K |
| 325 | Nanjing Normal University | 1 | 20 | 1.9K | 1.6K |
| 325 | Università degli Studi Roma Tre | 1 | 20 | 776 | 726 |
| 325 | Sokoine University of Agriculture | 1 | 20 | 431 | 414 |
| 330 | University of Aberdeen | 1 | 19 | 4.4K | 4.2K |
| 330 | University of Tromsø – The Arctic University of Norway | 1 | 19 | 2.2K | 2.4K |
| 330 | Gangneung–Wonju National University | 1 | 19 | 594 | 536 |
| 330 | Aristotle University of Thessaloniki | 3 | 19 | 3.1K | 4.6K |
| 330 | Universidad Nacional de La Plata | 1 | 19 | 2.0K | 1.6K |
| 330 | Norwegian Polar Institute | 1 | 19 | 191 | 245 |
| 330 | University of Massachusetts | 1 | 19 | 5.6K | 5.3K |
| 330 | University of Malaya | 1 | 19 | 4.2K | 3.7K |
| 330 | University of Algarve | 2 | 19 | 711 | 1.2K |
| 330 | University of Hawaii at Manoa | 1 | 19 | 2.3K | 2.5K |
| 330 | Pokhara University | 2 | 19 | 59 | 82 |
| 330 | Smithsonian Marine Station at Fort Pierce | 1 | 19 | 70 | 94 |
| 330 | Fram Centre | 1 | 19 | 70 | 116 |
| 330 | Uni Research | 1 | 19 | 91 | 176 |
| 344 | Utrecht University | 1 | 18 | 6.2K | 6.0K |
| 344 | University of Chile | 1 | 18 | 4.2K | 3.7K |
| 344 | University of Melbourne | 2 | 18 | 7.1K | 8.2K |
| 344 | Kyoto University | 2 | 18 | 5.6K | 6.1K |
| 344 | Adam Mickiewicz University in Poznań | 2 | 18 | 1.3K | 1.6K |
| 344 | University of La Laguna | 2 | 18 | 1.2K | 1.7K |
| 344 | University of Oklahoma | 1 | 18 | 4.4K | 4.1K |
| 344 | West University of Timișoara | 1 | 18 | 414 | 331 |
| 344 | Aix-Marseille University | 1 | 18 | 3.0K | 2.9K |
| 344 | Andrés Bello University | 1 | 18 | 820 | 765 |
| 344 | yasouj university | 2 | 18 | 264 | 373 |
| 344 | Universidade Estadual de Londrina | 1 | 18 | 1.2K | 837 |
| 356 | Autonomous University of Barcelona | 1 | 17 | 5.0K | 4.6K |
| 356 | University of Pennsylvania | 1 | 17 | 8.8K | 8.0K |
| 356 | Sun Yat-sen University | 2 | 17 | 4.1K | 4.8K |
| 356 | University of Hawaii | 2 | 17 | 2.0K | 3.2K |
| 356 | Saarland University | 1 | 17 | 2.8K | 2.8K |
| 356 | City University of New York | 1 | 17 | 5.1K | 4.5K |
| 356 | Sichuan University | 1 | 17 | 4.9K | 4.4K |
| 356 | Islamic Azad University | 1 | 17 | 5.7K | 4.5K |
| 356 | National Museums of Kenya | 1 | 17 | 184 | 229 |
| 356 | Federal University of São Paulo | 1 | 17 | 3.1K | 2.6K |
| 356 | College of Staten Island | 1 | 17 | 210 | 308 |
| 356 | Kunming Institute of Zoology | 1 | 17 | 154 | 232 |
| 356 | Qinghai University | 1 | 17 | 467 | 398 |
| 356 | Taif University | 1 | 17 | 986 | 903 |
| 356 | Maasai Mara University | 1 | 17 | 20 | 26 |
| 356 | Hamad Bin Khalifa University | 1 | 17 | 371 | 447 |
| 356 | Yibin University | 1 | 17 | 180 | 157 |
| 356 | Sorbonne University | 3 | 17 | 1.2K | 1.9K |
| 356 | Université de Nantes | 2 | 17 | 954 | 1.7K |
| 375 | Australian Antarctic Division | 1 | 16 | 177 | 253 |
| 375 | Utah Valley University | 1 | 16 | 226 | 262 |
| 375 | University of Salento | 1 | 16 | 1.6K | 1.8K |
| 375 | Monterey Bay Aquarium Research Institute | 1 | 16 | 160 | 226 |
| 375 | Royal Veterinary and Agricultural University | 1 | 16 | 508 | 667 |
| 380 | Brown University | 1 | 15 | 4.8K | 4.9K |
| 380 | Bavarian State Collection of Zoology | 1 | 15 | 67 | 84 |
| 380 | James Madison University | 1 | 15 | 1.1K | 1.2K |
| 380 | Museums Victoria | 1 | 15 | 55 | 78 |
| 380 | Smithsonian Tropical Research Institute | 1 | 15 | 391 | 508 |
| 380 | South China Normal University | 1 | 15 | 1.6K | 1.6K |
| 380 | Universidade Federal de São Carlos | 1 | 15 | 2.5K | 2.0K |
| 380 | The Education University of Hong Kong | 1 | 15 | 753 | 811 |
| 380 | Leshan Normal University | 1 | 15 | 91 | 100 |
| 380 | Claude Bernard University Lyon 1 | 1 | 15 | 2.6K | 2.9K |
| 380 | Academia Sinica | 1 | 15 | 3.5K | 3.5K |
| 391 | University of Warsaw | 1 | 14 | 3.3K | 2.8K |
| 391 | University of Greifswald | 1 | 14 | 1.6K | 1.9K |
| 391 | Autonomous University of Madrid | 2 | 14 | 2.8K | 4.0K |
| 391 | Aarhus University | 1 | 14 | 6.1K | 6.2K |
| 391 | Shinshu University | 1 | 14 | 2.0K | 1.9K |
| 391 | Old Dominion University | 1 | 14 | 2.3K | 2.5K |
| 391 | Universiti Malaysia Terengganu | 1 | 14 | 546 | 564 |
| 391 | National Council for Scientific and Technological Development | 1 | 14 | 177 | 215 |
| 391 | Syiah Kuala University | 1 | 14 | 624 | 344 |
| 391 | Lewis & Clark College | 1 | 14 | 90 | 154 |
| 391 | Alberta Biodiversity Monitoring Institute | 1 | 14 | 13 | 24 |
| 391 | Spanish National Research Council | 1 | 14 | 1.6K | 2.0K |
| 391 | Universidade de Santiago de Compostela | 1 | 14 | 1.4K | 1.5K |
| 404 | University of Antwerp | 1 | 13 | 3.9K | 4.2K |
| 404 | Imperial College London | 1 | 13 | 7.3K | 7.5K |
| 404 | Cairo University | 1 | 13 | 4.7K | 4.2K |
| 404 | University of Perugia | 1 | 13 | 3.4K | 3.8K |
| 404 | Kyushu University | 1 | 13 | 5.6K | 5.3K |
| 404 | National Polytechnic Institute | 2 | 13 | 1.6K | 2.1K |
| 404 | National University of Córdoba | 2 | 13 | 1.2K | 1.7K |
| 404 | Hofstra University | 1 | 13 | 864 | 991 |
| 404 | Bournemouth University | 1 | 13 | 1.2K | 1.4K |
| 404 | Beijing Forestry University | 1 | 13 | 1.0K | 1.1K |
| 404 | University of Tabuk | 1 | 13 | 498 | 530 |
| 404 | Tianjin Normal University | 1 | 13 | 641 | 685 |
| 404 | Museo Nacional de Ciencias Naturales | 1 | 13 | 296 | 391 |
| 417 | National Tsing Hua University | 1 | 12 | 2.4K | 2.5K |
| 417 | California Institute of Technology | 1 | 12 | 3.0K | 3.4K |
| 417 | Istanbul University | 1 | 12 | 3.5K | 3.5K |
| 417 | University of Liverpool | 1 | 12 | 6.3K | 6.3K |
| 417 | Chinese Culture University | 1 | 12 | 533 | 723 |
| 417 | California State University | 2 | 12 | 4.0K | 5.3K |
| 417 | Czech University of Life Sciences Prague | 1 | 12 | 600 | 682 |
| 417 | University of Cagliari | 1 | 12 | 2.8K | 3.3K |
| 417 | Federal University of Pernambuco | 1 | 12 | 3.2K | 2.6K |
| 417 | Andhra University | 1 | 12 | 887 | 883 |
| 417 | University of French Guiana | 1 | 12 | 48 | 88 |
| 417 | National Yang Ming Chiao Tung University | 1 | 12 | 851 | 705 |
| 417 | Kitakyushu Museum of Natural History and Human History | 1 | 12 | 27 | 38 |
| 430 | University of New Hampshire | 1 | 11 | 2.4K | 2.9K |
| 430 | University of Duisburg-Essen | 1 | 11 | 3.4K | 3.9K |
| 430 | University of East Anglia | 1 | 11 | 3.5K | 4.0K |
| 430 | University of Nice Sophia Antipolis | 1 | 11 | 1.2K | 1.6K |
| 430 | University of Messina | 1 | 11 | 2.9K | 3.3K |
| 430 | Eastern Connecticut State University | 1 | 11 | 142 | 218 |
| 430 | Marshall University | 1 | 11 | 683 | 979 |
| 430 | University of Cyprus | 1 | 11 | 1.5K | 1.8K |
| 430 | Centre for Agriculture and Biosciences International | 1 | 11 | 97 | 162 |
| 430 | University of California, Davis | 1 | 11 | 4.2K | 4.9K |
| 440 | University College London | 1 | 10 | 8.4K | 8.6K |
| 440 | University of Jaén | 2 | 10 | 951 | 1.8K |
| 440 | Queen Saovabha Memorial Institute | 1 | 10 | 43 | 68 |
| 440 | National Cheng Kung University | 1 | 10 | 3.7K | 4.1K |
| 440 | Yokohama National University | 1 | 10 | 1.4K | 1.6K |
| 440 | University of Western Australia | 2 | 10 | 4.0K | 6.1K |
| 440 | University of Ostrava | 1 | 10 | 427 | 553 |
| 440 | Hamilton College | 1 | 10 | 379 | 537 |
| 440 | University of Lorraine | 1 | 10 | 2.2K | 2.5K |
| 440 | Indiana University | 1 | 10 | 6.5K | 6.8K |
| 440 | Federal Rural University of Rio de Janeiro | 1 | 10 | 1.0K | 832 |
| 440 | Agriculture and Agriculture-Food Canada | 1 | 10 | 587 | 811 |
| 440 | Université de Perpignan | 2 | 10 | 256 | 594 |
| 453 | University of Jena | 1 | 9 | 3.0K | 3.6K |
| 453 | University of Malta | 1 | 9 | 967 | 1.3K |
| 453 | University of Illinois at Urbana–Champaign | 1 | 9 | 6.2K | 6.9K |
| 453 | University of Bristol | 1 | 9 | 6.4K | 7.1K |
| 453 | Université libre de Bruxelles | 1 | 9 | 4.0K | 4.7K |
| 453 | Drexel University | 1 | 9 | 3.1K | 3.9K |
| 453 | Southeastern Louisiana University | 3 | 9 | 118 | 504 |
| 453 | Tunghai University | 1 | 9 | 721 | 1.0K |
| 453 | Baylor University | 1 | 9 | 2.0K | 2.4K |
| 453 | University of Texas at Arlington | 2 | 9 | 1.5K | 3.0K |
| 453 | University of Putra Malaysia | 1 | 9 | 2.7K | 3.2K |
| 453 | University of Palermo | 1 | 9 | 3.2K | 3.9K |
| 453 | Sidi Mohamed Ben Abdellah University | 1 | 9 | 357 | 430 |
| 453 | National University of the Northeast | 1 | 9 | 277 | 327 |
| 453 | National University of Salta | 1 | 9 | 190 | 287 |
| 453 | Utica College | 1 | 9 | 99 | 134 |
| 453 | Huzhou University | 1 | 9 | 527 | 657 |
| 453 | Universidad Austral | 1 | 9 | 239 | 376 |
| 453 | University of Durban-Westville | 1 | 9 | 193 | 259 |
| 453 | University of Agriculture | 1 | 9 | 1.6K | 1.8K |
| 473 | University of Marburg | 1 | 8 | 3.1K | 3.8K |
| 473 | Sapienza University of Rome | 1 | 8 | 6.2K | 6.8K |
| 473 | University of A Coruña | 1 | 8 | 1.4K | 1.7K |
| 473 | Razi University | 1 | 8 | 997 | 1.3K |
| 473 | University of South Florida | 1 | 8 | 5.8K | 6.5K |
| 473 | Sonoma State University | 1 | 8 | 323 | 552 |
| 473 | University of Montpellier | 1 | 8 | 2.4K | 3.1K |
| 473 | Howard University | 1 | 8 | 1.7K | 2.2K |
| 473 | Colgate University | 1 | 8 | 639 | 846 |
| 473 | Edith Cowan University | 1 | 8 | 1.6K | 2.3K |
| 473 | University of Houston | 1 | 8 | 4.5K | 5.2K |
| 473 | IFREMER | 1 | 8 | 378 | 589 |
| 473 | University of Karachi | 3 | 8 | 414 | 1.2K |
| 473 | University of Windsor | 1 | 8 | 2.2K | 2.9K |
| 473 | Qufu Normal University | 1 | 8 | 844 | 1.0K |
| 473 | Ningbo University | 1 | 8 | 1.7K | 2.0K |
| 473 | Northeast Normal University | 1 | 8 | 1.3K | 1.6K |
| 473 | University of Abomey-Calavi | 1 | 8 | 451 | 624 |
| 473 | University of Minnesota Supercomputing Institute | 1 | 8 | 38 | 75 |
| 473 | Sichuan Agricultural University | 1 | 8 | 1.0K | 1.3K |
| 473 | Instituto Nacional de Ciencias Agrícolas | 1 | 8 | 195 | 284 |
| 473 | University of Nevada, Reno | 1 | 8 | 4.3K | 5.0K |
| 495 | Saint Petersburg State University | 1 | 7 | 1.5K | 1.5K |
| 495 | University of Münster | 1 | 7 | 4.0K | 4.9K |
| 495 | University of Salamanca | 1 | 7 | 2.7K | 3.1K |
| 495 | University of Lausanne | 1 | 7 | 3.3K | 4.3K |
| 495 | University of Groningen | 1 | 7 | 5.9K | 6.8K |
| 495 | Smithsonian Conservation Biology Institute | 1 | 7 | 186 | 287 |
| 495 | Tulane University | 1 | 7 | 3.2K | 4.1K |
| 495 | Louisiana State University | 3 | 7 | 3.0K | 6.3K |
| 495 | Bandung Institute of Technology | 1 | 7 | 737 | 900 |
| 495 | National Biodiversity Institute | 1 | 7 | 33 | 80 |
| 495 | Erciyes University | 1 | 7 | 1.8K | 2.2K |
| 495 | University of Caldas | 1 | 7 | 287 | 305 |
| 495 | Silesian Museum | 2 | 7 | 7 | 20 |
| 495 | National Zoological Park | 1 | 7 | 209 | 333 |
| 495 | University of Allahabad | 1 | 7 | 747 | 1.0K |
| 510 | Flinders University | 1 | 6 | 3.2K | 4.4K |
| 510 | University of Southampton | 1 | 6 | 6.1K | 7.3K |
| 510 | Stazione Zoologica Anton Dohrn | 1 | 6 | 263 | 464 |
| 510 | Monash University | 1 | 6 | 8.0K | 9.0K |
| 510 | Swiss Ornithological Institute Sempach | 1 | 6 | 11 | 19 |
| 510 | University of Vigo | 1 | 6 | 1.7K | 2.4K |
| 510 | Technical University of Braunschweig | 1 | 6 | 2.0K | 2.6K |
| 510 | University of Waterloo | 1 | 6 | 5.6K | 6.6K |
| 510 | University of New Brunswick | 1 | 6 | 2.1K | 2.8K |
| 510 | Xunta de Galicia | 1 | 6 | 56 | 141 |
| 510 | Wistar Institute | 1 | 6 | 426 | 722 |
| 510 | National Oceanography Centre | 1 | 6 | 270 | 410 |
| 510 | University of the French Antilles | 1 | 6 | 248 | 473 |
| 510 | Hessisches Landesmuseum Darmstadt | 1 | 6 | 26 | 53 |
| 510 | National Parks Board | 1 | 6 | 73 | 156 |
| 525 | Stanford University | 1 | 5 | 8.6K | 10.1K |
| 525 | University of Limpopo | 1 | 5 | 553 | 822 |
| 525 | Hiroshima University | 1 | 5 | 4.0K | 4.7K |
| 525 | Kasetsart University | 2 | 5 | 980 | 2.0K |
| 525 | University of Sfax | 1 | 5 | 1.5K | 2.0K |
| 525 | University of New South Wales | 1 | 5 | 8.2K | 9.5K |
| 525 | Katholieke Universiteit Leuven | 1 | 5 | 7.2K | 8.6K |
| 525 | Asahikawa Medical University | 1 | 5 | 505 | 741 |
| 525 | University of Castile-La Mancha | 1 | 5 | 1.3K | 1.8K |
| 525 | University of Liège | 1 | 5 | 3.9K | 5.1K |
| 525 | Portland State University | 1 | 5 | 2.0K | 3.0K |
| 525 | Wrocław University of Environmental and Life Sciences | 1 | 5 | 632 | 940 |
| 525 | National University of Jujuy | 1 | 5 | 94 | 162 |
| 525 | South China Agricultural University | 1 | 5 | 1.5K | 2.0K |
| 525 | University of Alaska Museum of the North | 1 | 5 | 61 | 118 |
| 525 | Xinzhou Teachers University | 1 | 5 | 47 | 85 |
| 541 | Central Michigan University | 1 | 4 | 1.4K | 2.3K |
| 541 | Universidad Nacional del Litoral | 1 | 4 | 561 | 875 |
| 541 | Tel Aviv University | 2 | 4 | 5.4K | 8.3K |
| 541 | University at Buffalo | 1 | 4 | 5.8K | 7.2K |
| 541 | University of Konstanz | 1 | 4 | 2.2K | 3.1K |
| 541 | University of New Mexico | 1 | 4 | 4.7K | 6.3K |
| 541 | Addis Ababa University | 1 | 4 | 1.4K | 2.2K |
| 541 | Brawijaya University | 1 | 4 | 1.1K | 933 |
| 541 | University of Reading | 1 | 4 | 3.7K | 5.0K |
| 541 | Indiana University of Pennsylvania | 1 | 4 | 516 | 930 |
| 541 | King Juan Carlos University | 1 | 4 | 1.7K | 2.4K |
| 541 | Universidade Federal de Santa Maria | 1 | 4 | 2.5K | 2.5K |
| 541 | Bahir Dar University | 1 | 4 | 434 | 747 |
| 541 | Edinboro University of Pennsylvania | 1 | 4 | 92 | 197 |
| 541 | Federal University of Paraíba | 1 | 4 | 2.0K | 2.2K |
| 541 | Universidade de Taubaté | 1 | 4 | 197 | 295 |
| 541 | University of Ilorin | 1 | 4 | 986 | 1.4K |
| 558 | Miguel Hernández University of Elche | 1 | 3 | 1.3K | 2.2K |
| 558 | Universidad Autónoma de Nuevo León | 1 | 3 | 1.4K | 2.2K |
| 558 | Alfred Wegener Institute for Polar and Marine Research | 1 | 3 | 409 | 646 |
| 558 | Syracuse University | 1 | 3 | 3.4K | 4.8K |
| 558 | Utah State University | 1 | 3 | 2.9K | 4.4K |
| 558 | Dali University | 1 | 3 | 196 | 423 |
| 558 | Limnological Institute | 1 | 3 | 187 | 307 |
| 565 | University of Delaware | 1 | 2 | 4.2K | 6.0K |
| 565 | East Carolina University | 1 | 2 | 2.7K | 4.3K |
| 565 | University of Agder | 1 | 2 | 741 | 1.5K |
| 565 | Savitribai Phule Pune University | 1 | 2 | 1.1K | 2.0K |
| 565 | University of Nebraska State Museum | 1 | 2 | 46 | 89 |
| 565 | Centre d'Écologie Fonctionnelle et Évolutive | 1 | 2 | 164 | 316 |
| 571 | Banaras Hindu University | 1 | 1 | 2.6K | 3.9K |
| 571 | University of Port Elizabeth | 1 | 1 | 198 | 350 |
| 571 | Universidad Central | 1 | 1 | 178 | 326 |
| 571 | Instituto Multidisciplinario de Biología Vegetal | 1 | 1 | 78 | 168 |
| 575 | Ludwig Maximilian University of Munich | 1 | 0 | 5.7K | 8.3K |
| 575 | University of London | 1 | 0 | 7.7K | 10.9K |
| 575 | Universidad Michoacana de San Nicolás de Hidalgo | 1 | 0 | 796 | 1.6K |
| 575 | Nicolaus Copernicus University in Toruń | 1 | 0 | 1.8K | 3.0K |
| 575 | University of Uyo | 1 | 0 | 335 | 847 |
