| # | Institution | Articles | Citations | R ARank | R CRank |
|---|
| 1 | Stockholm University | 117 | 2,761 | 54 | 148 |
| 2 | University of Copenhagen | 70 | 2,103 | 281 | 518 |
| 3 | Aarhus University | 28 | 525 | 520 | 1.0K |
| 4 | Lund University | 32 | 521 | 631 | 1.3K |
| 5 | University of Zurich | 23 | 511 | 657 | 1.1K |
| 6 | University of Gothenburg | 29 | 503 | 454 | 1.0K |
| 7 | University of Bonn | 4 | 500 | 2.3K | 837 |
| 8 | University of California | 23 | 479 | 5.0K | 5.5K |
| 9 | Scripps Institution of Oceanography | 16 | 463 | 55 | 115 |
| 10 | Osnabrück University | 12 | 443 | 142 | 169 |
| 11 | Natural History Museum | 27 | 436 | 129 | 233 |
| 12 | University of Bologna | 28 | 414 | 607 | 1.3K |
| 13 | University of Saskatchewan | 11 | 336 | 895 | 911 |
| 14 | University of Queensland | 22 | 334 | 1.3K | 2.4K |
| 15 | Swedish Museum of Natural History | 24 | 316 | 40 | 94 |
| 16 | University of Vienna | 12 | 315 | 1.2K | 1.3K |
| 17 | Uppsala University | 23 | 302 | 812 | 1.8K |
| 18 | University of Washington | 13 | 295 | 2.9K | 3.5K |
| 19 | Humboldt University of Berlin | 14 | 291 | 666 | 1.1K |
| 20 | University of Hamburg | 11 | 268 | 764 | 947 |
| 21 | University of Victoria | 12 | 263 | 513 | 802 |
| 22 | University of Tübingen | 10 | 256 | 1.2K | 1.3K |
| 23 | University of Padua | 15 | 255 | 1.3K | 2.0K |
| 24 | Complutense University of Madrid | 20 | 241 | 654 | 1.3K |
| 25 | Moscow State University | 8 | 233 | 1.3K | 827 |
| 26 | University of Buenos Aires | 12 | 231 | 685 | 834 |
| 27 | Russian Academy of Sciences | 18 | 223 | 1.4K | 1.6K |
| 28 | Jagiellonian University | 19 | 220 | 349 | 635 |
| 29 | University of Messina | 16 | 219 | 326 | 676 |
| 29 | Royal Agricultural University | 4 | 219 | 23 | 10 |
| 31 | University of New South Wales | 6 | 210 | 3.4K | 2.7K |
| 32 | University of Calgary | 9 | 203 | 2.1K | 2.3K |
| 33 | University of California, San Diego | 4 | 200 | 2.9K | 2.2K |
| 34 | National Scientific and Technical Research Council | 12 | 197 | 263 | 446 |
| 35 | University of São Paulo | 15 | 191 | 2.4K | 2.9K |
| 36 | University of Innsbruck | 3 | 187 | 1.6K | 926 |
| 37 | Muséum national d'histoire naturelle | 15 | 186 | 112 | 314 |
| 38 | University of Chicago | 3 | 178 | 4.2K | 2.7K |
| 39 | Clemson University | 4 | 176 | 1.6K | 979 |
| 40 | University of Jena | 14 | 172 | 415 | 1.0K |
| 41 | University of Cambridge | 7 | 171 | 3.6K | 3.7K |
| 42 | Katholieke Universiteit Leuven | 11 | 166 | 1.7K | 2.9K |
| 43 | University of Campinas | 10 | 163 | 1.1K | 1.1K |
| 44 | Ludwig Maximilian University of Munich | 9 | 162 | 1.7K | 2.3K |
| 45 | Heidelberg University | 7 | 156 | 2.0K | 2.3K |
| 45 | University of Sydney | 10 | 156 | 2.8K | 3.8K |
| 45 | University of the Witwatersrand | 8 | 156 | 959 | 1.0K |
| 45 | Murdoch University | 14 | 156 | 198 | 625 |
| 49 | Bavarian State Collection of Zoology | 6 | 152 | 12 | 19 |
| 50 | Universidad Nacional de La Plata | 9 | 147 | 338 | 410 |
| 51 | Smithsonian Institution | 8 | 146 | 235 | 346 |
| 52 | American Museum of Natural History | 6 | 139 | 169 | 220 |
| 53 | National Center for Scientific Research | 5 | 135 | 5.2K | 4.2K |
| 54 | Dalhousie University | 5 | 128 | 1.9K | 2.0K |
| 55 | University of Brasília | 9 | 126 | 616 | 707 |
| 56 | Central College | 5 | 125 | 27 | 23 |
| 57 | Claude Bernard University Lyon 1 | 3 | 122 | 1.5K | 1.1K |
| 58 | Newcastle University | 6 | 121 | 1.9K | 2.3K |
| 59 | Swansea University | 6 | 118 | 837 | 1.2K |
| 60 | University of Hong Kong | 6 | 117 | 2.7K | 3.0K |
| 61 | University of Göttingen | 2 | 112 | 3.1K | 2.1K |
| 61 | University of Mysore | 5 | 112 | 246 | 194 |
| 63 | Instituto Butantan | 12 | 111 | 51 | 163 |
| 64 | American University of Beirut | 2 | 108 | 1.6K | 780 |
| 64 | University of Modena and Reggio Emilia | 4 | 108 | 1.3K | 1.3K |
| 64 | University of Siena | 10 | 108 | 584 | 1.3K |
| 67 | Yale University | 1 | 107 | 6.6K | 3.3K |
| 67 | Federal University of Rio Grande do Sul | 4 | 107 | 2.4K | 1.6K |
| 67 | Universidade Estadual Paulista | 13 | 107 | 610 | 1.1K |
| 70 | University of Pittsburgh | 3 | 106 | 5.2K | 4.0K |
| 71 | University of New Hampshire | 3 | 105 | 1.2K | 1.0K |
| 71 | Ghent University | 4 | 105 | 3.5K | 3.2K |
| 73 | Freie Universität Berlin | 4 | 104 | 2.0K | 2.0K |
| 74 | University of Pisa | 7 | 95 | 1.8K | 2.5K |
| 74 | Universidade Federal do Paraná | 6 | 95 | 1.0K | 912 |
| 74 | University of Trieste | 5 | 95 | 967 | 1.2K |
| 77 | University of Milan | 4 | 94 | 3.2K | 3.1K |
| 77 | University of Basel | 6 | 94 | 1.2K | 1.9K |
| 77 | Paris Diderot University | 4 | 94 | 941 | 1.1K |
| 77 | Field Museum of Natural History | 4 | 94 | 160 | 202 |
| 81 | University of Cologne | 5 | 93 | 1.9K | 2.2K |
| 82 | University of Manchester | 4 | 91 | 4.6K | 4.4K |
| 83 | University of Freiburg | 2 | 89 | 2.8K | 2.1K |
| 84 | St George's, University of London | 1 | 87 | 914 | 535 |
| 85 | University of Salford | 3 | 86 | 949 | 777 |
| 86 | University of Silesia in Katowice | 5 | 85 | 551 | 396 |
| 87 | University of Kansas | 4 | 84 | 2.7K | 2.9K |
| 87 | Karolinska Institute | 2 | 84 | 3.6K | 2.9K |
| 89 | Kagawa University | 7 | 83 | 284 | 484 |
| 89 | Federal University of Viçosa | 7 | 83 | 545 | 624 |
| 91 | University of Salzburg | 13 | 82 | 82 | 625 |
| 92 | University of Burgundy | 1 | 81 | 1.5K | 818 |
| 92 | University of Bath | 4 | 81 | 1.6K | 1.9K |
| 92 | University of Palermo | 2 | 81 | 2.1K | 1.5K |
| 95 | Åbo Akademi University | 4 | 80 | 446 | 671 |
| 96 | University of Guelph | 7 | 78 | 1.2K | 2.1K |
| 96 | Umeå University | 8 | 78 | 1.0K | 2.2K |
| 96 | University of Iowa | 2 | 78 | 4.9K | 3.6K |
| 99 | North-West University | 2 | 77 | 850 | 479 |
| 100 | Australian National University | 3 | 76 | 3.3K | 3.1K |
| 101 | University of Bristol | 3 | 75 | 3.9K | 3.7K |
| 102 | Universidade do Estado do Rio de Janeiro | 5 | 73 | 612 | 515 |
| 103 | Kanazawa University | 5 | 71 | 1.3K | 1.4K |
| 104 | University of Western Australia | 4 | 70 | 2.7K | 3.4K |
| 105 | University of Louisiana at Lafayette | 3 | 69 | 425 | 436 |
| 105 | Northern Illinois University | 2 | 69 | 1.5K | 1.1K |
| 105 | Hvidovre Hospital | 1 | 69 | 564 | 410 |
| 108 | University of Wrocław | 4 | 66 | 830 | 734 |
| 108 | Saint Louis University | 4 | 66 | 761 | 1.2K |
| 108 | University of Leicester | 2 | 66 | 3.1K | 2.5K |
| 111 | University of Ottawa | 3 | 65 | 3.8K | 3.6K |
| 112 | University of Rajasthan | 7 | 63 | 152 | 275 |
| 112 | Natural History Museum of Los Angeles County | 2 | 63 | 87 | 84 |
| 114 | McGill University | 2 | 62 | 6.9K | 5.5K |
| 115 | University of Genoa | 3 | 61 | 2.5K | 2.4K |
| 116 | Bielefeld University | 2 | 60 | 1.7K | 1.4K |
| 116 | Dartmouth College | 3 | 60 | 1.3K | 1.7K |
| 116 | University of Calabria | 4 | 60 | 826 | 1.2K |
| 116 | Royal Ontario Museum | 2 | 60 | 123 | 119 |
| 120 | Harvard University | 3 | 59 | 4.4K | 4.8K |
| 120 | University of Málaga | 8 | 59 | 526 | 1.3K |
| 122 | Dicle University | 4 | 56 | 257 | 351 |
| 122 | Estonian University of Life Sciences | 1 | 56 | 415 | 275 |
| 124 | University of Oslo | 5 | 55 | 2.8K | 4.0K |
| 124 | Southeastern Louisiana University | 5 | 55 | 62 | 169 |
| 126 | Norwegian University of Science and Technology | 3 | 54 | 2.5K | 2.9K |
| 127 | University of Toronto | 7 | 53 | 5.3K | 7.6K |
| 127 | Tallinn University of Technology | 4 | 53 | 311 | 563 |
| 127 | Industrial University of Santander | 3 | 53 | 248 | 219 |
| 130 | Tel Aviv University | 3 | 52 | 4.4K | 4.5K |
| 130 | Chinese Academy of Sciences | 5 | 52 | 4.7K | 5.3K |
| 130 | Wittenberg University | 3 | 52 | 35 | 58 |
| 133 | University of New England | 7 | 51 | 375 | 1.3K |
| 133 | University of Girona | 2 | 51 | 787 | 840 |
| 135 | University of Tokyo | 5 | 50 | 4.8K | 5.5K |
| 135 | University of Rostock | 5 | 50 | 783 | 1.6K |
| 135 | Roskilde University | 1 | 50 | 915 | 566 |
| 135 | Osmania University | 3 | 50 | 425 | 440 |
| 139 | National Academy of Sciences | 1 | 49 | 1.2K | 515 |
| 139 | University of Alberta | 4 | 49 | 5.0K | 5.9K |
| 139 | University of Granada | 4 | 49 | 2.2K | 2.7K |
| 139 | University of Maine | 3 | 49 | 957 | 1.3K |
| 143 | Seoul National University | 1 | 48 | 5.8K | 4.0K |
| 143 | University of London | 3 | 48 | 4.6K | 5.0K |
| 143 | University of Barcelona | 4 | 48 | 3.1K | 3.8K |
| 143 | National Autonomous University of Mexico | 10 | 48 | 1.3K | 2.9K |
| 143 | Czech Academy of Sciences | 3 | 48 | 2.1K | 2.2K |
| 143 | Rhodes University | 3 | 48 | 593 | 690 |
| 143 | Florida State University | 1 | 48 | 4.5K | 3.0K |
| 143 | Pontifícia Universidade Católica de Minas Gerais | 3 | 48 | 156 | 133 |
| 143 | Pierre and Marie Curie University | 3 | 48 | 1.6K | 2.2K |
| 152 | Université libre de Bruxelles | 3 | 47 | 2.2K | 2.8K |
| 153 | Saint Petersburg State University | 5 | 46 | 479 | 547 |
| 153 | University of Cincinnati | 2 | 46 | 3.8K | 3.5K |
| 153 | Royal Veterinary and Agricultural University | 1 | 46 | 498 | 485 |
| 156 | University of Hawaii | 3 | 45 | 1.5K | 2.2K |
| 157 | Immanuel Kant Baltic Federal University | 2 | 44 | 195 | 73 |
| 157 | University of Florence | 3 | 44 | 3.1K | 3.5K |
| 159 | Cairo University | 5 | 43 | 1.9K | 2.5K |
| 159 | University of Wales | 2 | 43 | 1.6K | 1.8K |
| 159 | Royal Veterinary College | 2 | 43 | 339 | 433 |
| 162 | Niigata University | 2 | 42 | 1.7K | 1.4K |
| 162 | Ruhr University Bochum | 1 | 42 | 4.0K | 2.9K |
| 162 | University of Cantabria | 2 | 42 | 506 | 636 |
| 162 | Keio University | 3 | 42 | 2.2K | 2.3K |
| 162 | University of Massachusetts | 1 | 42 | 5.6K | 4.1K |
| 162 | Ochanomizu University | 3 | 42 | 318 | 403 |
| 162 | Saitama University | 3 | 42 | 638 | 664 |
| 162 | Universidad Autónoma del Estado de Hidalgo | 4 | 42 | 151 | 210 |
| 170 | Minia University | 5 | 41 | 268 | 514 |
| 171 | Hebrew University of Jerusalem | 1 | 40 | 5.1K | 3.6K |
| 171 | Utrecht University | 2 | 40 | 4.7K | 4.8K |
| 171 | Macquarie University | 5 | 40 | 1.5K | 3.0K |
| 171 | Huazhong Agricultural University | 2 | 40 | 1.1K | 1.2K |
| 171 | Universidad de la República | 2 | 40 | 1.1K | 952 |
| 176 | Wrocław University of Environmental and Life Sciences | 6 | 39 | 159 | 340 |
| 176 | Bernardo O'Higgins University | 2 | 39 | 148 | 154 |
| 176 | National University of Mar del Plata | 6 | 39 | 124 | 406 |
| 176 | University of Tennessee | 1 | 39 | 6.2K | 4.4K |
| 180 | University of Bergen | 8 | 38 | 1.1K | 3.2K |
| 180 | University of South Florida | 3 | 38 | 3.3K | 4.1K |
| 182 | University of Antwerp | 3 | 37 | 2.0K | 3.0K |
| 182 | University College London | 2 | 37 | 6.5K | 6.2K |
| 182 | Sapienza University of Rome | 1 | 37 | 6.2K | 4.5K |
| 182 | University of Southern Denmark | 2 | 37 | 2.2K | 2.5K |
| 182 | Instituto Biológico | 2 | 37 | 216 | 256 |
| 187 | University of Adelaide | 7 | 36 | 1.9K | 4.4K |
| 187 | Leipzig University | 3 | 36 | 2.2K | 2.9K |
| 187 | University of León | 3 | 36 | 527 | 809 |
| 187 | Beijing Forestry University | 3 | 36 | 564 | 730 |
| 191 | RWTH Aachen University | 1 | 35 | 4.3K | 3.4K |
| 191 | Washington State University | 3 | 35 | 2.8K | 3.8K |
| 191 | University of Alcalá | 3 | 35 | 1.1K | 1.6K |
| 191 | University of Cádiz | 3 | 35 | 683 | 986 |
| 191 | Bharathidasan University | 2 | 35 | 474 | 517 |
| 196 | University of Valencia | 2 | 34 | 3.5K | 3.3K |
| 196 | University of Maryland | 1 | 34 | 8.8K | 6.7K |
| 196 | Kyushu University | 3 | 34 | 3.8K | 4.0K |
| 196 | Smithsonian Tropical Research Institute | 1 | 34 | 390 | 388 |
| 196 | Natural History Museum Vienna | 1 | 34 | 92 | 68 |
| 201 | University of Latvia | 3 | 33 | 402 | 648 |
| 201 | Harbor Branch Oceanographic Institute | 2 | 33 | 68 | 127 |
| 201 | Shahid Bahonar University of Kerman | 6 | 33 | 249 | 728 |
| 201 | Weifang University | 1 | 33 | 208 | 92 |
| 205 | University of St Andrews | 2 | 32 | 1.5K | 1.8K |
| 205 | University of North Carolina | 2 | 32 | 4.9K | 5.3K |
| 205 | Academy of Natural Sciences of Drexel University | 1 | 32 | 145 | 154 |
| 208 | University of Pretoria | 5 | 31 | 1.3K | 2.5K |
| 208 | University of Allahabad | 4 | 31 | 275 | 572 |
| 210 | School for Advanced Studies in the Social Sciences | 1 | 30 | 279 | 263 |
| 210 | University of Bern | 1 | 30 | 4.9K | 4.1K |
| 210 | California State University | 2 | 30 | 4.0K | 3.8K |
| 210 | University of Alaska Fairbanks | 1 | 30 | 952 | 878 |
| 210 | Obafemi Awolowo University | 2 | 30 | 815 | 744 |
| 215 | University of Halle-Wittenberg | 1 | 29 | 1.1K | 1.0K |
| 215 | Mercer University | 2 | 29 | 213 | 347 |
| 217 | King's College London | 3 | 28 | 3.5K | 4.8K |
| 217 | University of Regensburg | 2 | 28 | 2.0K | 2.5K |
| 217 | Stellenbosch University | 1 | 28 | 3.5K | 2.8K |
| 217 | University of Porto | 3 | 28 | 1.8K | 2.9K |
| 217 | University of Reading | 2 | 28 | 2.6K | 3.1K |
| 217 | Bundelkhand University | 2 | 28 | 32 | 106 |
| 217 | Federal Rural University of Rio de Janeiro | 4 | 28 | 342 | 484 |
| 217 | Royal Holloway, University of London | 2 | 28 | 1.1K | 1.5K |
| 217 | University of Mons-Hainaut | 2 | 28 | 171 | 314 |
| 217 | École Normale Supérieure | 1 | 28 | 1.5K | 1.4K |
| 217 | Université de Paris | 1 | 28 | 1.7K | 1.1K |
| 228 | Monash University | 2 | 27 | 6.2K | 6.5K |
| 228 | University of Sheffield | 1 | 27 | 6.6K | 5.4K |
| 228 | University of the Ryukyus | 2 | 27 | 1.2K | 1.3K |
| 228 | Nanjing Normal University | 2 | 27 | 1.3K | 1.4K |
| 228 | Northwest University | 1 | 27 | 1.6K | 1.2K |
| 228 | Fayoum University | 1 | 27 | 620 | 415 |
| 228 | Northeastern University | 1 | 27 | 5.0K | 4.1K |
| 235 | Ohio University | 1 | 26 | 2.5K | 2.1K |
| 235 | University of Richmond | 1 | 26 | 692 | 580 |
| 235 | University of Santiago de Compostela | 3 | 26 | 1.2K | 2.1K |
| 235 | The Nippon Dental University | 1 | 26 | 278 | 242 |
| 235 | Toho University | 1 | 26 | 1.3K | 963 |
| 240 | University of Leeds | 1 | 25 | 6.9K | 5.7K |
| 240 | Technical University of Denmark | 1 | 25 | 3.4K | 3.2K |
| 240 | Central Marine Fisheries Research Institute | 2 | 25 | 48 | 77 |
| 240 | University of Rzeszow | 1 | 25 | 794 | 457 |
| 244 | Leiden University | 2 | 24 | 3.1K | 3.7K |
| 244 | Vikram University | 5 | 24 | 42 | 115 |
| 244 | King's College | 2 | 24 | 1.6K | 2.4K |
| 247 | Nagoya City University | 1 | 23 | 1.1K | 981 |
| 247 | Naturalis Biodiversity Center | 2 | 23 | 134 | 199 |
| 247 | Kaliningrad State Technical University | 2 | 23 | 41 | 15 |
| 247 | Bangor University | 3 | 23 | 862 | 1.7K |
| 247 | Ehime University | 1 | 23 | 1.8K | 1.4K |
| 247 | Rothamsted Research | 1 | 23 | 646 | 714 |
| 247 | Limnological Institute | 2 | 23 | 107 | 167 |
| 254 | University of Tsukuba | 1 | 22 | 4.8K | 3.9K |
| 254 | University of Liège | 1 | 22 | 3.9K | 3.6K |
| 254 | Ca' Foscari University of Venice | 3 | 22 | 281 | 732 |
| 254 | Virginia Commonwealth University | 1 | 22 | 4.6K | 4.1K |
| 254 | National University of Tucumán | 1 | 22 | 388 | 367 |
| 254 | Carnegie Museum of Natural History | 2 | 22 | 61 | 114 |
| 254 | Museum of Paleontology Egidio Feruglio | 1 | 22 | 33 | 43 |
| 261 | University of Glasgow | 2 | 21 | 4.8K | 5.7K |
| 261 | Wageningen University & Research | 1 | 21 | 3.1K | 3.2K |
| 261 | Freiberg University of Mining and Technology | 1 | 21 | 825 | 795 |
| 261 | University of KwaZulu-Natal | 1 | 21 | 2.7K | 2.3K |
| 261 | Queen's University Belfast | 2 | 21 | 2.7K | 3.6K |
| 261 | Austral University of Chile | 1 | 21 | 1.0K | 915 |
| 261 | Bulgarian Academy of Sciences | 4 | 21 | 1.1K | 1.9K |
| 261 | Universidade Federal do Ceará | 3 | 21 | 1.4K | 1.8K |
| 261 | Universidad Nacional del Comahue | 1 | 21 | 378 | 350 |
| 261 | Universidade Estadual de Maringá | 2 | 21 | 1.2K | 1.1K |
| 261 | Mato Grosso do Sul State University | 1 | 21 | 265 | 162 |
| 272 | University of Konstanz | 1 | 20 | 2.2K | 2.2K |
| 272 | Tohoku University | 2 | 20 | 4.0K | 4.4K |
| 272 | Beni-Suef University | 2 | 20 | 579 | 775 |
| 272 | Universidade Federal de São Carlos | 1 | 20 | 2.5K | 1.8K |
| 272 | Universidad Autónoma del Estado de México | 5 | 20 | 137 | 426 |
| 277 | University of Canterbury | 2 | 19 | 2.0K | 2.9K |
| 277 | National University of Singapore | 2 | 19 | 5.9K | 6.8K |
| 277 | University of Oregon | 2 | 19 | 2.3K | 3.1K |
| 277 | Station biologique de Roscoff | 2 | 19 | 87 | 223 |
| 277 | University of Kerala | 3 | 19 | 275 | 583 |
| 277 | Tuscia University | 2 | 19 | 574 | 1.0K |
| 277 | United States Forest Service | 1 | 19 | 836 | 909 |
| 277 | Mississippi State University | 1 | 19 | 2.8K | 2.6K |
| 277 | Erciyes University | 1 | 19 | 1.8K | 1.5K |
| 277 | Bozok University | 1 | 19 | 408 | 355 |
| 277 | Ondokuz Mayıs University | 1 | 19 | 1.5K | 1.2K |
| 277 | Federal University of Espírito Santo | 4 | 19 | 687 | 1.2K |
| 289 | University of Helsinki | 1 | 18 | 6.6K | 6.0K |
| 289 | Cornell University | 4 | 18 | 4.1K | 7.1K |
| 289 | Osaka University | 3 | 18 | 3.4K | 4.6K |
| 289 | Ocean University of China | 1 | 18 | 1.8K | 1.8K |
| 289 | University of Murcia | 2 | 18 | 2.0K | 2.6K |
| 289 | Villanova University | 1 | 18 | 1.3K | 1.3K |
| 289 | Henderson State University | 1 | 18 | 35 | 36 |
| 289 | University of Johannesburg | 2 | 18 | 1.3K | 1.7K |
| 289 | National University of the Northeast | 1 | 18 | 277 | 219 |
| 289 | Virginia Institute of Marine Science | 5 | 18 | 118 | 358 |
| 289 | Universidad de Navarra | 1 | 18 | 2.0K | 1.9K |
| 289 | Oklahoma State University | 1 | 18 | 3.6K | 3.5K |
| 301 | Goethe University Frankfurt | 1 | 17 | 3.8K | 3.7K |
| 301 | Hiroshima University | 1 | 17 | 4.0K | 3.5K |
| 301 | Kyoto University | 4 | 17 | 4.2K | 6.2K |
| 301 | National and Kapodistrian University of Athens | 1 | 17 | 5.2K | 4.8K |
| 301 | University of Arkansas | 1 | 17 | 3.5K | 3.2K |
| 301 | Shimane University | 2 | 17 | 975 | 1.3K |
| 301 | University of New Mexico | 1 | 17 | 4.7K | 4.6K |
| 301 | University of Catania | 2 | 17 | 2.5K | 3.6K |
| 301 | Saint Anselm College | 2 | 17 | 29 | 71 |
| 310 | University of Utah | 1 | 16 | 6.8K | 6.5K |
| 310 | Universiti Brunei Darussalam | 1 | 16 | 575 | 643 |
| 310 | Polish Academy of Sciences | 1 | 16 | 4.5K | 4.2K |
| 310 | University of Naples Federico II | 3 | 16 | 2.9K | 4.9K |
| 310 | Hokkaido University | 2 | 16 | 4.0K | 4.5K |
| 310 | Colorado State University | 2 | 16 | 3.5K | 4.8K |
| 310 | Shiraz University | 8 | 16 | 473 | 2.0K |
| 310 | Federal Rural University of Pernambuco | 2 | 16 | 651 | 724 |
| 310 | Western Australian Museum | 1 | 16 | 117 | 153 |
| 319 | University of Michigan | 2 | 15 | 8.6K | 9.8K |
| 319 | National Centre for Biological Sciences | 1 | 15 | 245 | 355 |
| 319 | University of Alabama at Birmingham | 1 | 15 | 4.8K | 4.8K |
| 319 | Visva-Bharati University | 1 | 15 | 428 | 454 |
| 319 | Federal University of Pernambuco | 1 | 15 | 3.2K | 2.4K |
| 319 | Saint Mary's College | 1 | 15 | 93 | 103 |
| 319 | Australian Museum | 1 | 15 | 202 | 251 |
| 319 | University of Veterinary and Pharmaceutical Sciences Brno | 2 | 15 | 109 | 252 |
| 327 | Stanford University | 1 | 14 | 8.6K | 8.5K |
| 327 | University of Greifswald | 3 | 14 | 782 | 1.9K |
| 327 | University of Tartu | 1 | 14 | 2.5K | 2.7K |
| 327 | University of Vigo | 2 | 14 | 1.1K | 1.8K |
| 327 | Stony Brook University | 1 | 14 | 4.3K | 4.6K |
| 327 | Federal University of São Paulo | 3 | 14 | 1.7K | 2.8K |
| 327 | Université de Nantes | 1 | 14 | 1.5K | 1.8K |
| 327 | Poznań University of Life Sciences | 1 | 14 | 630 | 708 |
| 327 | Nagpur University | 5 | 14 | 43 | 184 |
| 336 | Lesya Ukrainka Volyn National University | 1 | 13 | 126 | 53 |
| 336 | Brown University | 1 | 13 | 4.8K | 5.1K |
| 336 | University of Kiel | 1 | 13 | 2.1K | 2.6K |
| 336 | University of Münster | 3 | 13 | 2.3K | 4.3K |
| 336 | University of Amsterdam | 1 | 13 | 6.8K | 6.9K |
| 336 | Université catholique de Louvain | 1 | 13 | 4.5K | 4.9K |
| 336 | Arkansas State University | 1 | 13 | 415 | 526 |
| 336 | Technical University of Braunschweig | 1 | 13 | 2.0K | 2.2K |
| 336 | University of the West Indies | 1 | 13 | 1.7K | 1.8K |
| 336 | Barry University | 1 | 13 | 192 | 309 |
| 336 | Florida International University | 1 | 13 | 3.6K | 3.8K |
| 336 | Smithsonian Conservation Biology Institute | 1 | 13 | 186 | 241 |
| 336 | Presidency University | 1 | 13 | 600 | 619 |
| 336 | Vidyasagar University | 1 | 13 | 521 | 579 |
| 336 | Indian Institute of Chemical Technology | 1 | 13 | 445 | 581 |
| 336 | Staatliches Museum für Naturkunde Stuttgart | 2 | 13 | 45 | 114 |
| 336 | National Zoological Park | 1 | 13 | 208 | 301 |
| 353 | Kasetsart University | 5 | 12 | 464 | 1.6K |
| 353 | Gujarat University | 3 | 12 | 154 | 444 |
| 353 | Université Cheikh Anta Diop | 2 | 12 | 227 | 561 |
| 353 | Saint Mary's University | 2 | 12 | 381 | 860 |
| 353 | Ohio Northern University | 1 | 12 | 146 | 217 |
| 353 | University of Northern Iowa | 1 | 12 | 773 | 934 |
| 353 | Mount Saint Vincent University | 2 | 12 | 139 | 374 |
| 353 | Museum für Naturkunde | 1 | 12 | 18 | 46 |
| 361 | Virginia Tech | 2 | 11 | 4.4K | 6.1K |
| 361 | University of British Columbia | 2 | 11 | 7.9K | 9.9K |
| 361 | George Washington University | 1 | 11 | 4.5K | 4.8K |
| 361 | University of Miami | 1 | 11 | 5.3K | 5.7K |
| 361 | Baylor University | 1 | 11 | 2.0K | 2.3K |
| 361 | University of Wisconsin–Madison | 2 | 11 | 7.9K | 9.9K |
| 361 | Yokohama City University | 1 | 11 | 1.3K | 1.5K |
| 361 | Prince of Songkla University | 1 | 11 | 1.7K | 1.9K |
| 361 | University of Plymouth | 2 | 11 | 1.5K | 2.8K |
| 361 | Hasselt University | 2 | 11 | 715 | 1.6K |
| 361 | Universidade Federal de Ouro Preto | 2 | 11 | 457 | 862 |
| 361 | Universidade Católica de Brasília | 1 | 11 | 381 | 483 |
| 361 | University of Vale do Rio dos Sinos | 1 | 11 | 682 | 537 |
| 361 | National Museums Scotland | 3 | 11 | 33 | 119 |
| 361 | Virginia–Maryland College of Veterinary Medicine | 2 | 11 | 217 | 462 |
| 376 | Warsaw University of Life Sciences | 3 | 10 | 310 | 866 |
| 376 | University of Oxford | 1 | 10 | 9.7K | 9.7K |
| 376 | American University | 1 | 10 | 1.6K | 1.9K |
| 376 | Stazione Zoologica Anton Dohrn | 2 | 10 | 162 | 417 |
| 376 | Technion – Israel Institute of Technology | 3 | 10 | 2.2K | 4.3K |
| 376 | Daegu University | 1 | 10 | 542 | 776 |
| 376 | Masaryk University | 1 | 10 | 2.5K | 2.8K |
| 376 | University of Calcutta | 3 | 10 | 1.0K | 2.0K |
| 376 | University of Colorado | 1 | 10 | 6.2K | 6.8K |
| 376 | St. Lawrence University | 1 | 10 | 219 | 284 |
| 376 | Nicolaus Copernicus University in Toruń | 2 | 10 | 1.2K | 1.8K |
| 376 | Weber State University | 2 | 10 | 157 | 459 |
| 376 | Jadavpur University | 2 | 10 | 1.5K | 2.3K |
| 376 | University of Portsmouth | 2 | 10 | 1.3K | 2.7K |
| 376 | National Council for Scientific and Technological Development | 1 | 10 | 175 | 257 |
| 376 | Millikin University | 1 | 10 | 63 | 110 |
| 376 | Federal University of Mato Grosso | 1 | 10 | 1.1K | 883 |
| 376 | California State University, Fullerton | 1 | 10 | 364 | 612 |
| 376 | Oswaldo Cruz Foundation | 1 | 10 | 1.6K | 1.8K |
| 395 | University of Ljubljana | 1 | 9 | 4.1K | 4.5K |
| 395 | Autonomous University of Barcelona | 1 | 9 | 5.0K | 5.5K |
| 395 | University of Iceland | 2 | 9 | 1.2K | 2.5K |
| 395 | University of Belgrade | 2 | 9 | 3.2K | 4.6K |
| 395 | University of Tromsø – The Arctic University of Norway | 1 | 9 | 2.2K | 2.9K |
| 395 | Panjab University | 1 | 9 | 1.7K | 1.9K |
| 395 | University of Perugia | 1 | 9 | 3.4K | 4.1K |
| 395 | University of Toyama | 1 | 9 | 2.2K | 2.4K |
| 395 | DePauw University | 1 | 9 | 174 | 249 |
| 395 | Chinese Academy of Fishery Sciences | 1 | 9 | 474 | 639 |
| 395 | Federal University of Alfenas | 1 | 9 | 530 | 521 |
| 395 | Queensland Museum | 3 | 9 | 50 | 184 |
| 407 | Federal University of Minas Gerais | 3 | 8 | 2.4K | 4.4K |
| 407 | Yarmouk University | 2 | 8 | 430 | 1.0K |
| 407 | Ohio State University | 3 | 8 | 6.4K | 9.9K |
| 407 | University of Duisburg-Essen | 1 | 8 | 3.4K | 4.2K |
| 407 | Federal University of Rio Grande | 1 | 8 | 881 | 845 |
| 407 | Universidade Federal de Goiás | 1 | 8 | 2.2K | 2.0K |
| 407 | Université de Bamenda | 1 | 8 | 366 | 585 |
| 407 | Marine Biological Association of the United Kingdom | 1 | 8 | 126 | 244 |
| 407 | Sparsholt College | 1 | 8 | 6 | 14 |
| 407 | Universitat de Lleida | 1 | 8 | 482 | 740 |
| 417 | Columbia University | 1 | 7 | 8.9K | 9.6K |
| 417 | Barnard College | 1 | 7 | 462 | 682 |
| 417 | Pasteur Institute | 1 | 7 | 1.7K | 2.2K |
| 417 | James Cook University | 1 | 7 | 2.3K | 3.3K |
| 417 | Banaras Hindu University | 2 | 7 | 1.9K | 3.1K |
| 417 | Mie University | 1 | 7 | 1.5K | 1.8K |
| 417 | Paris Descartes University | 1 | 7 | 2.4K | 3.1K |
| 417 | University of Patras | 1 | 7 | 3.0K | 3.8K |
| 417 | Hólar University College | 1 | 7 | 33 | 67 |
| 417 | Maria Curie-Skłodowska University | 3 | 7 | 523 | 1.2K |
| 417 | Palacký University Olomouc | 1 | 7 | 1.5K | 2.0K |
| 417 | Indian Statistical Institute | 1 | 7 | 1.1K | 1.4K |
| 417 | Joetsu University of Education | 1 | 7 | 141 | 178 |
| 417 | Universidade Federal de Santa Maria | 1 | 7 | 2.5K | 2.2K |
| 417 | Universidade Federal do Pará | 1 | 7 | 1.7K | 1.8K |
| 417 | Framingham State University | 1 | 7 | 85 | 147 |
| 417 | Norwegian Contractors | 1 | 7 | 2 | 4 |
| 417 | Plovdiv University | 2 | 7 | 104 | 282 |
| 417 | Paleontological Institute, Russian Academy of Sciences | 1 | 7 | 35 | 64 |
| 417 | California State University Fresno | 1 | 7 | 137 | 306 |
| 437 | University of Limpopo | 1 | 6 | 550 | 783 |
| 437 | Jacobs University Bremen | 1 | 6 | 686 | 1.2K |
| 437 | Instituto Antártico Chileno | 1 | 6 | 21 | 52 |
| 437 | Zhejiang University | 1 | 6 | 6.1K | 6.8K |
| 437 | University of Giessen | 2 | 6 | 2.3K | 4.1K |
| 437 | Loyola University | 1 | 6 | 983 | 1.5K |
| 437 | University of Khartoum | 3 | 6 | 311 | 1.2K |
| 437 | National University of Colombia | 2 | 6 | 1.5K | 2.8K |
| 437 | Lafayette College | 1 | 6 | 518 | 804 |
| 437 | Sichuan University | 1 | 6 | 4.9K | 5.4K |
| 447 | Gunma University | 1 | 5 | 1.7K | 2.1K |
| 447 | Razi University | 1 | 5 | 994 | 1.4K |
| 447 | University of Split | 1 | 5 | 909 | 1.4K |
| 447 | University of Nairobi | 1 | 5 | 1.4K | 1.9K |
| 447 | University of Texas at Arlington | 1 | 5 | 2.4K | 3.4K |
| 447 | Pontifical Xavierian University | 2 | 5 | 591 | 1.5K |
| 447 | University of the Free State | 1 | 5 | 1.5K | 2.0K |
| 447 | Norwegian University of Life Sciences | 1 | 5 | 1.2K | 1.9K |
| 447 | University of Warmia and Mazury in Olsztyn | 2 | 5 | 788 | 1.4K |
| 447 | King Saud University | 3 | 5 | 2.6K | 5.6K |
| 447 | Memorial University of Newfoundland | 1 | 5 | 3.3K | 4.5K |
| 447 | Jena University Hospital | 1 | 5 | 762 | 1.3K |
| 447 | Savitribai Phule Pune University | 1 | 5 | 1.1K | 1.7K |
| 447 | Qufu Normal University | 1 | 5 | 842 | 1.1K |
| 447 | National Institute of Oceanography | 1 | 5 | 346 | 586 |
| 447 | Mar Ivanios College | 1 | 5 | 76 | 119 |
| 447 | Southwest Forestry University | 1 | 5 | 325 | 543 |
| 447 | University of Findlay | 3 | 5 | 21 | 143 |
| 447 | University of Guilan | 1 | 5 | 1.2K | 1.6K |
| 447 | University of Indianapolis | 3 | 5 | 46 | 351 |
| 447 | Yibin University | 1 | 5 | 180 | 287 |
| 447 | Academia Sinica | 1 | 5 | 3.5K | 4.3K |
| 447 | University of Virginia | 1 | 5 | 6.4K | 7.6K |
| 447 | Universidad del Bío-Bío | 1 | 5 | 350 | 519 |
| 471 | National Sun Yat-sen University | 1 | 4 | 1.8K | 2.7K |
| 471 | University of Warsaw | 1 | 4 | 3.3K | 3.9K |
| 471 | Rajamangala University of Technology | 1 | 4 | 158 | 310 |
| 471 | Infineon Technologies | 1 | 4 | 198 | 302 |
| 471 | University of Turin | 1 | 4 | 5.5K | 6.9K |
| 471 | Southeast Missouri State University | 1 | 4 | 258 | 544 |
| 471 | Arizona State University | 1 | 4 | 6.6K | 8.0K |
| 471 | Chulalongkorn University | 1 | 4 | 3.1K | 4.3K |
| 471 | University of Debrecen | 1 | 4 | 2.2K | 3.0K |
| 471 | San Diego State University | 1 | 4 | 3.2K | 4.5K |
| 471 | Instituto Antártico Argentino | 1 | 4 | 62 | 145 |
| 471 | University of South Carolina | 1 | 4 | 5.4K | 7.0K |
| 471 | Tokyo Metropolitan University | 1 | 4 | 1.9K | 2.5K |
| 471 | Missouri State University | 1 | 4 | 945 | 1.5K |
| 471 | Gazi University | 2 | 4 | 1.9K | 3.5K |
| 471 | NOVA University Lisbon | 1 | 4 | 2.6K | 3.7K |
| 471 | University of Natal | 1 | 4 | 894 | 1.4K |
| 471 | Federal University of ABC | 1 | 4 | 969 | 1.4K |
| 471 | Universidade Federal de Mato Grosso do Sul | 1 | 4 | 1.1K | 1.1K |
| 471 | Instituto Lauro de Souza Lima | 1 | 4 | 34 | 70 |
| 471 | National Institute for Basic Biology, Japan | 1 | 4 | 352 | 545 |
| 471 | University of Durban-Westville | 1 | 4 | 190 | 328 |
| 471 | Museu da Lourinhã | 1 | 4 | 1 | 12 |
| 494 | New York University | 1 | 3 | 6.1K | 7.7K |
| 494 | University of Birmingham | 1 | 3 | 7.2K | 9.0K |
| 494 | Alzahra University | 2 | 3 | 304 | 952 |
| 494 | University of the Western Cape | 1 | 3 | 1.4K | 2.2K |
| 494 | Federal University of Rio de Janeiro | 1 | 3 | 5.2K | 5.9K |
| 494 | University of Oldenburg | 1 | 3 | 1.1K | 1.8K |
| 494 | European Molecular Biology Laboratory | 1 | 3 | 189 | 372 |
| 494 | Atatürk University | 1 | 3 | 1.9K | 2.7K |
| 494 | Austin Peay State University | 1 | 3 | 169 | 341 |
| 494 | Slovak Academy of Sciences | 1 | 3 | 2.2K | 3.2K |
| 494 | University of Akron | 1 | 3 | 2.0K | 3.1K |
| 494 | University of Valparaíso | 1 | 3 | 603 | 1.1K |
| 494 | University of Campania "Luigi Vanvitelli" | 1 | 3 | 2.1K | 3.3K |
| 494 | Taizhou University | 1 | 3 | 467 | 802 |
| 494 | University of Chinese Academy of Sciences | 1 | 3 | 3.2K | 4.1K |
| 494 | Quanzhou Normal University | 1 | 3 | 193 | 362 |
| 494 | Shenyang Normal University | 1 | 3 | 290 | 511 |
| 494 | Virginia Wesleyan College | 1 | 3 | 25 | 59 |
| 494 | Henan Normal University | 1 | 3 | 968 | 1.4K |
| 494 | Zoological Survey of India | 1 | 3 | 111 | 165 |
| 514 | University of Massachusetts Lowell | 1 | 2 | 1.3K | 2.3K |
| 514 | Technical University of Munich | 1 | 2 | 5.8K | 7.7K |
| 514 | Karnatak University | 2 | 2 | 357 | 951 |
| 514 | Oregon State University | 1 | 2 | 4.4K | 6.4K |
| 514 | University of Cape Town | 1 | 2 | 4.4K | 6.3K |
| 514 | Chiang Mai University | 1 | 2 | 2.3K | 3.7K |
| 514 | University of Otago | 1 | 2 | 4.6K | 6.5K |
| 514 | Lithuanian Academy of Sciences | 1 | 2 | 57 | 120 |
| 514 | Murmansk Marine Biological Institute | 1 | 2 | 43 | 74 |
| 514 | Utkal University | 1 | 2 | 353 | 717 |
| 514 | Karadeniz Technical University | 1 | 2 | 1.6K | 2.6K |
| 514 | Munzur University | 1 | 2 | 206 | 441 |
| 514 | Sylhet Agricultural University | 1 | 2 | 116 | 222 |
| 514 | Toronto Western Hospital | 1 | 2 | 692 | 1.2K |
| 514 | Marine and Freshwater Research Institute | 1 | 2 | 153 | 305 |
| 514 | Sydney Medical School | 1 | 2 | 94 | 324 |
| 514 | Mohanlal Sukhadia University, Udaipur | 1 | 2 | 190 | 386 |
| 514 | University of California, San Francisco | 1 | 2 | 3.7K | 5.0K |
| 532 | National Marine Fisheries Service | 1 | 1 | 457 | 770 |
| 532 | Middle East Technical University | 1 | 1 | 2.5K | 4.1K |
| 532 | Boğaziçi University | 1 | 1 | 1.4K | 2.6K |
| 532 | University of Ibadan | 2 | 1 | 1.6K | 3.9K |
| 532 | Naresuan University | 1 | 1 | 610 | 1.3K |
| 532 | National Polytechnic Institute | 1 | 1 | 2.3K | 3.6K |
| 532 | Sultan Qaboos University | 1 | 1 | 1.7K | 3.2K |
| 532 | University of Alicante | 1 | 1 | 2.2K | 3.5K |
| 532 | Aichi Gakuin University | 1 | 1 | 268 | 561 |
| 532 | Istanbul University-Cerrahpaşa | 1 | 1 | 680 | 1.4K |
| 532 | Meerut College | 1 | 1 | 50 | 118 |
| 532 | South Valley University | 1 | 1 | 693 | 1.4K |
| 532 | Tokyo Dental College | 1 | 1 | 422 | 713 |
| 532 | Universidad Juárez Autónoma de Tabasco | 1 | 1 | 298 | 632 |
| 532 | Universidade Paranaense | 1 | 1 | 133 | 240 |
| 532 | Khadir Mohideen College | 1 | 1 | 13 | 46 |
| 532 | universidad técnica Latinoamericana | 1 | 1 | 195 | 308 |
| 532 | University of Newcastle | 1 | 1 | 5.0K | 7.5K |
| 532 | Cotton University | 1 | 1 | 80 | 207 |
| 551 | Nagasaki University | 2 | 0 | 1.4K | 3.2K |
| 551 | University of Notre Dame | 1 | 0 | 3.9K | 6.4K |
| 551 | Tokai University | 1 | 0 | 1.7K | 2.9K |
| 551 | California Lutheran University | 1 | 0 | 88 | 328 |
| 551 | Carleton University | 1 | 0 | 3.5K | 5.9K |
| 551 | University of Nice Sophia Antipolis | 1 | 0 | 1.2K | 2.3K |
| 551 | Namık Kemal University | 1 | 0 | 508 | 1.3K |
| 551 | Kyoto Sangyo University | 1 | 0 | 382 | 823 |
| 551 | Weill Cornell Medicine | 2 | 0 | 2.5K | 4.8K |
| 551 | Mansoura University | 1 | 0 | 2.4K | 4.1K |
| 551 | Deen Dayal Upadhyay Gorakhpur University | 1 | 0 | 193 | 336 |
| 551 | Dr. Hari Singh Gour University | 1 | 0 | 57 | 131 |
| 551 | Jichi Medical University | 1 | 0 | 1.1K | 1.9K |
| 551 | Federal University of Juiz de Fora | 1 | 0 | 1.6K | 2.6K |
