| # | Institution | Articles | Citations | R ARank | R CRank |
|---|
| 1 | University of California | 195 | 24,538 | 759 | 331 |
| 2 | Howard Hughes Medical Institute | 129 | 19,505 | 59 | 53 |
| 3 | National Institutes of Health | 71 | 13,558 | 418 | 162 |
| 4 | University of Texas Southwestern Medical Center | 39 | 7,503 | 333 | 91 |
| 5 | National Center for Scientific Research | 66 | 7,462 | 903 | 269 |
| 6 | Scripps Research | 32 | 6,805 | 167 | 74 |
| 7 | University of California, San Diego | 45 | 6,768 | 330 | 147 |
| 8 | University of California, San Francisco | 48 | 5,755 | 288 | 155 |
| 9 | University of Cambridge | 51 | 5,414 | 696 | 327 |
| 10 | University of Virginia | 20 | 4,933 | 781 | 119 |
| 11 | National Institute of Dental and Craniofacial Research | 17 | 4,823 | 85 | 21 |
| 12 | University of North Carolina | 28 | 4,761 | 571 | 172 |
| 13 | Washington University School of Medicine | 28 | 4,546 | 533 | 209 |
| 14 | University of Basel | 24 | 4,233 | 274 | 64 |
| 15 | University College London | 35 | 4,037 | 814 | 343 |
| 16 | University of Pennsylvania | 24 | 3,964 | 1.4K | 469 |
| 17 | Cancer Research UK | 18 | 3,826 | 75 | 37 |
| 18 | Duke University Hospital | 28 | 3,738 | 541 | 218 |
| 19 | University of Washington | 23 | 3,727 | 1.9K | 621 |
| 20 | Ludwig Cancer Research | 19 | 3,719 | 74 | 44 |
| 21 | Massachusetts Institute of Technology | 30 | 3,629 | 731 | 411 |
| 22 | Kyoto University | 9 | 3,512 | 2.7K | 291 |
| 23 | University of Michigan | 19 | 3,251 | 2.3K | 703 |
| 24 | Memorial Sloan Kettering Cancer Center | 22 | 3,229 | 432 | 176 |
| 25 | University of Manchester | 21 | 3,194 | 1.2K | 285 |
| 26 | Whitehead Institute | 13 | 3,125 | 53 | 39 |
| 27 | Columbia University | 17 | 3,114 | 1.7K | 493 |
| 28 | Friedrich Miescher Institute for Biomedical Research | 15 | 3,080 | 29 | 16 |
| 29 | Stanford University | 22 | 2,899 | 1.4K | 687 |
| 30 | University of Toronto | 16 | 2,765 | 3.0K | 788 |
| 31 | Johns Hopkins University | 31 | 2,712 | 794 | 460 |
| 32 | University of Chicago | 22 | 2,650 | 996 | 460 |
| 33 | University of Dundee | 25 | 2,540 | 175 | 96 |
| 34 | University of Geneva | 38 | 2,463 | 164 | 151 |
| 35 | Cornell University | 12 | 2,363 | 2.0K | 555 |
| 36 | Lawrence Berkeley National Laboratory | 7 | 2,334 | 586 | 172 |
| 37 | Albert Einstein College of Medicine | 14 | 2,298 | 575 | 164 |
| 38 | University of Göttingen | 5 | 2,223 | 1.8K | 177 |
| 39 | University of Tokyo | 14 | 2,190 | 2.7K | 664 |
| 40 | Beth Israel Deaconess Medical Center | 7 | 2,189 | 1.1K | 230 |
| 41 | Harvard University | 18 | 2,166 | 1.2K | 660 |
| 42 | Fred Hutchinson Cancer Research Center | 22 | 2,165 | 185 | 126 |
| 43 | The Rockefeller University | 22 | 2,129 | 194 | 164 |
| 44 | University of Edinburgh | 19 | 2,057 | 1.2K | 465 |
| 45 | Massachusetts General Hospital | 19 | 2,050 | 1.1K | 484 |
| 46 | University of Iowa | 14 | 2,021 | 1.3K | 367 |
| 47 | University of Wisconsin–Madison | 19 | 1,999 | 2.0K | 871 |
| 48 | French National Institute of Health and Medical Research | 10 | 1,975 | 945 | 243 |
| 49 | Yale University | 18 | 1,974 | 1.0K | 528 |
| 50 | Eunice Kennedy Shriver National Institute of Child Health and Human Development | 15 | 1,919 | 209 | 115 |
| 51 | Sanford Burnham Prebys Medical Discovery Institute | 8 | 1,888 | 84 | 31 |
| 52 | McGill University | 16 | 1,884 | 1.7K | 638 |
| 53 | Johns Hopkins School of Medicine | 17 | 1,837 | 447 | 219 |
| 54 | Princeton University | 19 | 1,757 | 722 | 466 |
| 55 | University of British Columbia | 12 | 1,722 | 2.8K | 891 |
| 56 | Imperial College London | 13 | 1,709 | 2.1K | 794 |
| 57 | California Institute of Technology | 16 | 1,687 | 681 | 407 |
| 58 | University of Texas at Austin | 14 | 1,684 | 2.4K | 849 |
| 59 | University of Queensland | 13 | 1,680 | 2.1K | 649 |
| 60 | University of North Carolina at Chapel Hill | 16 | 1,663 | 1.0K | 453 |
| 61 | National Jewish Health | 4 | 1,658 | 214 | 51 |
| 62 | Heidelberg University | 12 | 1,653 | 1.3K | 443 |
| 63 | Lunenfeld-Tanenbaum Research Institute | 13 | 1,652 | 75 | 47 |
| 64 | Stony Brook University | 6 | 1,631 | 1.4K | 246 |
| 65 | Osaka University | 17 | 1,614 | 1.2K | 383 |
| 66 | University of Colorado | 15 | 1,584 | 995 | 450 |
| 67 | Cleveland Clinic | 5 | 1,539 | 1.7K | 331 |
| 68 | Utrecht University | 16 | 1,510 | 1.2K | 589 |
| 69 | Pasteur Institute | 23 | 1,474 | 234 | 198 |
| 70 | Virginia Tech | 3 | 1,470 | 3.5K | 310 |
| 71 | University of California, Berkeley | 15 | 1,453 | 668 | 436 |
| 72 | Research Institute of Molecular Pathology | 6 | 1,443 | 30 | 24 |
| 73 | Vanderbilt University Medical Center | 4 | 1,432 | 1.4K | 194 |
| 74 | University of Utah | 15 | 1,424 | 1.2K | 519 |
| 75 | Budapest University of Technology and Economics | 1 | 1,423 | 1.7K | 28 |
| 76 | ETH Zürich | 16 | 1,400 | 944 | 505 |
| 77 | Baylor College of Medicine | 16 | 1,367 | 1.0K | 503 |
| 78 | University of Helsinki | 16 | 1,366 | 1.2K | 570 |
| 79 | Michigan Medicine | 15 | 1,290 | 600 | 325 |
| 80 | University of Würzburg | 6 | 1,272 | 1.3K | 307 |
| 81 | University of Konstanz | 5 | 1,269 | 754 | 135 |
| 82 | John Innes Centre | 10 | 1,252 | 126 | 80 |
| 83 | Weizmann Institute of Science | 13 | 1,249 | 386 | 230 |
| 84 | National Heart, Lung, and Blood Institute | 8 | 1,239 | 310 | 151 |
| 85 | University of Minnesota | 14 | 1,219 | 2.6K | 1.3K |
| 86 | University of Texas MD Anderson Cancer Center | 6 | 1,163 | 1.0K | 295 |
| 87 | University of Arizona | 5 | 1,150 | 3.5K | 746 |
| 88 | University of Münster | 6 | 1,148 | 1.4K | 336 |
| 89 | Weill Cornell Medicine | 10 | 1,137 | 978 | 415 |
| 90 | St. Jude Children's Research Hospital | 6 | 1,104 | 420 | 153 |
| 91 | Austrian Academy of Sciences | 6 | 1,083 | 243 | 74 |
| 92 | PSL University | 14 | 1,075 | 140 | 61 |
| 93 | Northwestern University | 17 | 1,059 | 1.3K | 961 |
| 94 | University of Oxford | 23 | 1,050 | 1.5K | 1.4K |
| 95 | WEHI | 3 | 1,029 | 245 | 70 |
| 96 | Addenbrooke's Hospital | 7 | 1,018 | 543 | 236 |
| 97 | University of Pittsburgh | 11 | 1,013 | 2.3K | 1.1K |
| 98 | Johns Hopkins Hospital | 1 | 1,008 | 1.6K | 131 |
| 99 | Stowers Institute for Medical Research | 7 | 988 | 46 | 28 |
| 100 | Ludwig Maximilian University of Munich | 8 | 987 | 1.9K | 692 |
| 101 | Robert Wood Johnson Medical School | 7 | 981 | 253 | 77 |
| 102 | Tel Aviv University | 7 | 956 | 2.6K | 720 |
| 103 | Uppsala University | 5 | 954 | 2.9K | 782 |
| 104 | Karolinska Institute | 8 | 953 | 1.9K | 858 |
| 105 | Vanderbilt University | 14 | 937 | 881 | 589 |
| 106 | Tokyo Medical and Dental University | 1 | 934 | 2.3K | 207 |
| 107 | Salk Institute for Biological Studies | 14 | 914 | 59 | 79 |
| 108 | University of Lausanne | 8 | 907 | 786 | 342 |
| 109 | University Medical Center Utrecht | 8 | 877 | 929 | 440 |
| 110 | Brandeis University | 7 | 857 | 402 | 174 |
| 111 | Rutgers University | 6 | 834 | 2.8K | 899 |
| 112 | Mayo Clinic | 7 | 818 | 2.1K | 814 |
| 113 | University of Birmingham | 9 | 809 | 2.1K | 863 |
| 114 | University of Illinois at Chicago | 4 | 792 | 2.8K | 586 |
| 115 | London Research Institute | 11 | 789 | 33 | 48 |
| 116 | Universidade Lusófona do Porto | 1 | 760 | 120 | 19 |
| 117 | University of Massachusetts | 5 | 740 | 2.1K | 619 |
| 118 | Hebrew University of Jerusalem | 9 | 729 | 1.3K | 603 |
| 119 | Nagoya University | 7 | 725 | 1.8K | 540 |
| 120 | European Molecular Biology Laboratory | 5 | 718 | 66 | 42 |
| 121 | University of Alberta | 13 | 704 | 2.1K | 1.3K |
| 122 | McGill University Health Centre | 4 | 671 | 948 | 301 |
| 123 | Case Western Reserve University | 7 | 669 | 1.9K | 853 |
| 124 | Oklahoma Medical Research Foundation | 3 | 664 | 189 | 50 |
| 125 | Barnes-Jewish Hospital | 1 | 648 | 262 | 35 |
| 126 | University of Alabama at Birmingham | 4 | 637 | 2.5K | 772 |
| 127 | State University of New York Upstate Medical University | 4 | 618 | 653 | 121 |
| 128 | University of Turku | 3 | 613 | 2.2K | 475 |
| 129 | National Institute of Genetics | 6 | 611 | 142 | 88 |
| 130 | Beatson West of Scotland Cancer Centre | 2 | 603 | 156 | 41 |
| 131 | Hammersmith Hospital | 3 | 598 | 837 | 283 |
| 132 | Children's Hospital Boston | 2 | 595 | 508 | 176 |
| 133 | La Jolla Institute for Allergy and Immunology | 1 | 570 | 176 | 51 |
| 134 | University of Leicester | 3 | 566 | 2.4K | 554 |
| 135 | Lehigh University | 2 | 557 | 1.4K | 239 |
| 136 | Washington State University | 3 | 550 | 2.8K | 749 |
| 137 | MRC London Institute of Medical Sciences | 1 | 528 | 123 | 15 |
| 138 | École polytechnique fédérale de Lausanne | 9 | 525 | 593 | 458 |
| 139 | University of Turin | 2 | 516 | 4.1K | 798 |
| 140 | UCL Institute of Ophthalmology | 5 | 513 | 81 | 47 |
| 141 | Thomas Jefferson University | 4 | 498 | 1.5K | 534 |
| 142 | Georgetown University Medical Center | 1 | 488 | 1.4K | 217 |
| 143 | Oregon Health & Science University | 3 | 480 | 1.7K | 587 |
| 143 | European Institute of Oncology | 6 | 480 | 194 | 148 |
| 143 | Huntsman Cancer Institute | 6 | 480 | 138 | 79 |
| 146 | University of Groningen | 7 | 479 | 2.1K | 1.3K |
| 146 | University of California, Los Angeles | 10 | 479 | 1.3K | 1.2K |
| 148 | Radboud University Nijmegen | 2 | 475 | 3.1K | 788 |
| 149 | University of Rochester | 4 | 474 | 2.3K | 1.0K |
| 149 | Karlsruhe Institute of Technology | 3 | 474 | 2.1K | 671 |
| 151 | NYU Langone Medical Center | 5 | 470 | 654 | 312 |
| 152 | University of Southern California | 5 | 458 | 3.7K | 1.8K |
| 153 | State University of New York | 5 | 451 | 2.7K | 1.1K |
| 154 | Iwate Medical University | 1 | 450 | 860 | 69 |
| 155 | University of Milan | 3 | 449 | 3.8K | 1.2K |
| 156 | University of California, Davis | 5 | 438 | 1.6K | 888 |
| 157 | Emory University | 5 | 437 | 2.3K | 1.1K |
| 158 | National University of Singapore | 12 | 430 | 1.9K | 1.6K |
| 159 | Boston Biomedical Research Institute | 2 | 421 | 78 | 35 |
| 160 | Stanford University Medical Center | 4 | 417 | 638 | 291 |
| 161 | Ottawa Hospital Research Institute | 1 | 415 | 934 | 189 |
| 162 | University of Copenhagen | 8 | 413 | 2.6K | 1.8K |
| 162 | Vollum Institute | 2 | 413 | 103 | 59 |
| 164 | AMOLF | 3 | 411 | 57 | 35 |
| 165 | Buck Institute for Research on Aging | 2 | 409 | 116 | 74 |
| 166 | University of Vienna | 6 | 408 | 2.1K | 1.1K |
| 166 | Radboud University Medical Centre | 1 | 408 | 1.7K | 420 |
| 168 | Biozentrum University of Basel | 3 | 407 | 44 | 33 |
| 169 | University of Sydney | 3 | 403 | 6.0K | 2.2K |
| 170 | Fudan University | 1 | 400 | 5.3K | 1.1K |
| 170 | Wellcome Trust Centre for Cell-Matrix Research | 5 | 400 | 18 | 17 |
| 172 | Yokohama City University | 1 | 397 | 1.3K | 153 |
| 173 | Carnegie Institution for Science | 7 | 395 | 193 | 205 |
| 174 | Leiden University Medical Center | 2 | 380 | 2.1K | 831 |
| 175 | Curie Institute | 2 | 378 | 44 | 20 |
| 175 | Sunnybrook Health Sciences Centre | 1 | 378 | 1.1K | 199 |
| 177 | University of Liverpool | 1 | 376 | 6.2K | 1.2K |
| 178 | Max Planck Institute of Biochemistry | 4 | 372 | 41 | 42 |
| 179 | University of East Anglia | 1 | 369 | 3.5K | 672 |
| 180 | University of Bristol | 11 | 367 | 1.6K | 1.6K |
| 181 | Duke University | 6 | 363 | 2.3K | 1.5K |
| 181 | Nara Institute of Science and Technology | 3 | 363 | 453 | 145 |
| 183 | University of Cologne | 9 | 352 | 1.2K | 982 |
| 183 | Carnegie Mellon University | 4 | 352 | 1.9K | 1.1K |
| 183 | Erasmus University Rotterdam | 3 | 352 | 1.7K | 713 |
| 186 | University of Bergen | 3 | 351 | 2.3K | 902 |
| 186 | University of Tsukuba | 1 | 351 | 4.8K | 838 |
| 186 | École Normale Supérieure | 3 | 351 | 750 | 323 |
| 189 | University of Leeds | 4 | 349 | 3.5K | 1.5K |
| 189 | Florida State University | 5 | 349 | 1.9K | 951 |
| 191 | Ontario Cancer Institute | 1 | 347 | 386 | 125 |
| 191 | University Health Network | 1 | 347 | 2.1K | 570 |
| 193 | National Institute for Physiological Sciences | 1 | 346 | 324 | 75 |
| 194 | Karolinska University Hospital | 2 | 345 | 1.9K | 749 |
| 195 | University of Colorado Denver | 2 | 342 | 1.8K | 483 |
| 196 | Anschutz Medical Campus | 4 | 340 | 941 | 535 |
| 197 | Ghent University | 3 | 336 | 4.1K | 1.7K |
| 198 | University of Glasgow | 9 | 335 | 1.7K | 1.5K |
| 199 | University of New Mexico | 1 | 333 | 4.7K | 852 |
| 200 | University of Georgia | 3 | 328 | 3.8K | 1.5K |
| 201 | University of Freiburg | 4 | 327 | 1.8K | 935 |
| 201 | Louis Pasteur University | 2 | 327 | 851 | 302 |
| 203 | University of Sheffield | 5 | 325 | 2.9K | 1.6K |
| 204 | University of Marburg | 2 | 319 | 2.2K | 725 |
| 205 | Manchester Academic Health Science Centre | 2 | 316 | 487 | 142 |
| 206 | University of Connecticut Health Center | 4 | 315 | 705 | 404 |
| 207 | Aarhus University | 3 | 312 | 3.6K | 1.5K |
| 208 | King's College London | 4 | 310 | 2.9K | 1.5K |
| 209 | University of Seville | 2 | 305 | 3.0K | 615 |
| 210 | University of Hamburg | 2 | 304 | 3.0K | 862 |
| 210 | Indiana University | 3 | 304 | 3.8K | 1.6K |
| 212 | Czech Academy of Sciences | 2 | 301 | 2.6K | 788 |
| 213 | Pennsylvania State University | 3 | 293 | 6.0K | 2.6K |
| 214 | Wistar Institute | 2 | 292 | 299 | 146 |
| 215 | TU Dresden | 7 | 290 | 1.2K | 871 |
| 215 | University of Calgary | 2 | 290 | 5.5K | 1.8K |
| 215 | Kumamoto University | 1 | 290 | 2.5K | 430 |
| 218 | University of Giessen | 2 | 289 | 2.3K | 791 |
| 219 | Roy J. and Lucille A. Carver College of Medicine | 1 | 282 | 687 | 137 |
| 220 | University of Nebraska Medical Center | 2 | 280 | 2.0K | 629 |
| 221 | Ohio State University | 5 | 277 | 4.9K | 2.9K |
| 222 | Dartmouth College | 3 | 275 | 1.3K | 666 |
| 223 | UNC Lineberger Comprehensive Cancer Center | 3 | 274 | 39 | 37 |
| 224 | University of Oslo | 2 | 271 | 4.7K | 1.7K |
| 224 | Pierre and Marie Curie University | 3 | 271 | 1.6K | 822 |
| 226 | Erasmus MC | 6 | 270 | 1.1K | 924 |
| 227 | Instituto Gulbenkian de Ciência | 5 | 268 | 67 | 83 |
| 228 | Royal Netherlands Academy of Arts and Sciences | 3 | 267 | 53 | 57 |
| 229 | University of Wisconsin School of Medicine and Public Health | 2 | 265 | 667 | 299 |
| 230 | Wellcome Trust Centre for Stem Cell Research | 1 | 264 | 11 | 8 |
| 231 | Nebraska Medical Center | 1 | 262 | 575 | 124 |
| 232 | University of Alabama | 2 | 261 | 3.7K | 1.1K |
| 233 | USC Norris Comprehensive Cancer Center | 1 | 260 | 79 | 28 |
| 234 | University of Hyogo | 1 | 258 | 1.2K | 162 |
| 235 | University of New South Wales | 3 | 255 | 5.2K | 2.4K |
| 236 | University of Lisbon | 4 | 252 | 1.9K | 775 |
| 237 | University of Notre Dame | 2 | 248 | 2.7K | 901 |
| 238 | (former) Institut National de Recherche pour l'Agriculture, l'Alimentation et l'Environnement | 2 | 247 | 1.8K | 922 |
| 239 | University of Ottawa | 2 | 241 | 4.8K | 1.7K |
| 240 | Technical University of Madrid | 1 | 236 | 2.4K | 456 |
| 240 | Colorado State University | 3 | 236 | 2.7K | 1.4K |
| 242 | University of Texas System | 3 | 233 | 1.3K | 753 |
| 243 | UC San Diego School of Medicine | 2 | 232 | 398 | 177 |
| 244 | Instituto de Medicina Molecular | 3 | 231 | 158 | 104 |
| 244 | University Medical Center | 4 | 231 | 1.3K | 866 |
| 246 | University of Zurich | 9 | 229 | 1.6K | 1.9K |
| 247 | University of Colorado Boulder | 3 | 225 | 1.6K | 939 |
| 247 | Leonard M. Miller School of Medicine | 2 | 225 | 2.1K | 925 |
| 249 | Brown University | 2 | 221 | 3.6K | 1.6K |
| 250 | University of Ferrara | 1 | 219 | 2.5K | 588 |
| 251 | University of Buenos Aires | 1 | 215 | 4.5K | 891 |
| 251 | University of Chile | 2 | 215 | 3.0K | 918 |
| 253 | National and Kapodistrian University of Athens | 1 | 213 | 5.2K | 1.4K |
| 254 | University of Oregon | 3 | 211 | 1.8K | 997 |
| 255 | Life Sciences Institute | 1 | 210 | 34 | 24 |
| 256 | University of Oulu | 1 | 208 | 3.4K | 998 |
| 257 | Medical University of Innsbruck | 4 | 207 | 698 | 534 |
| 257 | Oregon State University | 1 | 207 | 4.4K | 1.5K |
| 259 | Tokyo Institute of Technology | 3 | 206 | 2.3K | 1.1K |
| 259 | City University of New York | 1 | 206 | 5.1K | 1.1K |
| 261 | Université de Montréal | 3 | 205 | 3.7K | 2.0K |
| 262 | University at Albany | 1 | 201 | 1.6K | 422 |
| 263 | Oslo University Hospital | 1 | 200 | 1.8K | 557 |
| 264 | Queen Mary University of London | 2 | 199 | 1.7K | 661 |
| 265 | Trinity College Dublin | 1 | 193 | 3.4K | 939 |
| 265 | Klinikum Aachen | 1 | 193 | 568 | 132 |
| 265 | Åbo Akademi University | 1 | 193 | 1.3K | 332 |
| 268 | Charles University | 1 | 192 | 4.6K | 1.2K |
| 269 | University of Maryland | 1 | 190 | 8.8K | 3.4K |
| 270 | University of Gothenburg | 2 | 189 | 4.2K | 2.0K |
| 270 | University of Oklahoma | 1 | 189 | 4.4K | 1.2K |
| 272 | University of Illinois at Urbana–Champaign | 2 | 188 | 4.7K | 2.3K |
| 273 | Stockholm University | 2 | 186 | 3.1K | 1.5K |
| 274 | University of Graz | 1 | 182 | 2.6K | 762 |
| 275 | National Institute for Basic Biology, Japan | 1 | 181 | 352 | 162 |
| 276 | University of Naples Federico II | 4 | 178 | 2.4K | 1.6K |
| 277 | Chinese Academy of Sciences | 5 | 176 | 4.7K | 3.6K |
| 278 | Texas Tech University Health Sciences Center | 1 | 174 | 1.5K | 370 |
| 279 | Michigan State University | 3 | 173 | 5.0K | 3.0K |
| 279 | Columbia University Irving Medical Center | 3 | 173 | 1.4K | 847 |
| 281 | Lund University | 2 | 172 | 5.3K | 2.7K |
| 281 | Cardiff University | 2 | 172 | 3.8K | 1.8K |
| 283 | University of Cincinnati | 3 | 171 | 3.0K | 1.7K |
| 284 | Medical Research Council | 3 | 170 | 171 | 192 |
| 285 | Institut Gustave Roussy | 1 | 167 | 614 | 300 |
| 285 | Children's Hospital of Philadelphia | 3 | 167 | 1.2K | 820 |
| 287 | University of Barcelona | 3 | 159 | 3.7K | 2.2K |
| 287 | University of Wyoming | 1 | 159 | 2.6K | 779 |
| 289 | Boston University | 2 | 155 | 4.1K | 2.1K |
| 289 | Monash University | 4 | 155 | 4.3K | 3.2K |
| 291 | North Carolina State University | 1 | 150 | 5.8K | 2.3K |
| 291 | Massey University | 2 | 150 | 2.0K | 946 |
| 293 | Western General Hospital | 2 | 146 | 705 | 484 |
| 294 | Purdue University | 2 | 144 | 5.7K | 3.1K |
| 295 | Peking University | 1 | 142 | 5.5K | 2.3K |
| 296 | The Christie NHS Foundation Trust | 1 | 138 | 451 | 200 |
| 297 | Ehime University | 1 | 137 | 1.8K | 464 |
| 297 | University of Nice Sophia Antipolis | 1 | 137 | 1.2K | 389 |
| 297 | St James's University Hospital | 1 | 137 | 602 | 232 |
| 300 | Iowa State University | 2 | 136 | 4.0K | 2.3K |
| 301 | University of Aberdeen | 1 | 135 | 4.4K | 1.8K |
| 301 | Indiana University Bloomington | 1 | 135 | 1.7K | 576 |
| 303 | Medical University of Vienna | 4 | 134 | 1.5K | 1.3K |
| 304 | Tohoku University | 2 | 132 | 4.0K | 2.1K |
| 305 | ESPCI Paris, PSL University | 1 | 123 | 278 | 137 |
| 306 | Max Planck Society | 2 | 122 | 311 | 240 |
| 306 | James Franck Institute | 1 | 122 | 104 | 81 |
| 308 | Vrije Universiteit Amsterdam | 2 | 120 | 3.5K | 2.2K |
| 308 | Laval University | 1 | 120 | 5.1K | 2.3K |
| 308 | Centre hospitalier universitaire de Québec | 1 | 120 | 293 | 145 |
| 311 | Ajuntament de L’Hospitalet | 1 | 119 | 443 | 187 |
| 312 | Leipzig University | 1 | 118 | 3.9K | 1.7K |
| 312 | University of Neuchâtel | 1 | 118 | 1.1K | 471 |
| 312 | University of Bern | 4 | 118 | 2.5K | 2.3K |
| 315 | Temple Health | 1 | 117 | 107 | 22 |
| 316 | Umeå University | 1 | 116 | 3.9K | 1.8K |
| 317 | Witten/Herdecke University | 1 | 115 | 520 | 209 |
| 317 | University of Victoria | 1 | 115 | 3.7K | 1.5K |
| 319 | Drexel University | 1 | 114 | 3.1K | 1.3K |
| 320 | University of Tromsø – The Arctic University of Norway | 1 | 113 | 2.2K | 969 |
| 321 | University of St Andrews | 1 | 111 | 2.2K | 927 |
| 321 | Nanyang Technological University | 2 | 111 | 3.5K | 2.1K |
| 323 | École Normale Supérieure de Lyon | 2 | 110 | 679 | 459 |
| 324 | Osnabrück University | 3 | 109 | 584 | 507 |
| 324 | University of Siena | 1 | 109 | 3.0K | 1.3K |
| 324 | University of Western Australia | 1 | 109 | 5.4K | 2.7K |
| 327 | German Cancer Research Center | 2 | 104 | 621 | 540 |
| 328 | Pompeu Fabra University | 3 | 102 | 710 | 661 |
| 328 | New York University | 2 | 102 | 4.4K | 2.7K |
| 330 | INFN Sezione di Napoli | 1 | 100 | 62 | 29 |
| 331 | University of Medicine and Dentistry of New Jersey | 1 | 99 | 848 | 496 |
| 332 | Raman Research Institute | 1 | 97 | 215 | 89 |
| 332 | Cancer Research Institute | 2 | 97 | 540 | 400 |
| 334 | Institut Pasteur de Lille | 1 | 96 | 458 | 316 |
| 335 | Xiamen University | 1 | 95 | 3.2K | 1.3K |
| 336 | Saarland University | 2 | 93 | 2.0K | 1.4K |
| 336 | Sorbonne Paris Cité University (group) | 1 | 93 | 1.4K | 679 |
| 338 | University of Missouri | 2 | 92 | 4.9K | 3.2K |
| 338 | Nathan Kline Institute for Psychiatric Research | 1 | 92 | 120 | 102 |
| 338 | VCU Medical Center | 2 | 92 | 971 | 727 |
| 341 | National Human Genome Research Institute | 1 | 90 | 600 | 431 |
| 342 | National Institute on Aging | 2 | 88 | 663 | 723 |
| 343 | University of Stuttgart | 1 | 87 | 2.7K | 1.3K |
| 344 | University Medical Center Groningen | 2 | 86 | 1.7K | 1.4K |
| 345 | Hungarian Academy of Sciences | 1 | 85 | 3.0K | 1.6K |
| 345 | Semmelweis University | 1 | 85 | 2.0K | 1.0K |
| 345 | University of Kentucky | 1 | 85 | 6.3K | 3.2K |
| 345 | Loyola University Chicago | 1 | 85 | 2.2K | 945 |
| 349 | University of Rijeka | 1 | 84 | 883 | 230 |
| 349 | University of York | 2 | 84 | 2.9K | 2.1K |
| 351 | Janelia Research Campus | 2 | 83 | 79 | 107 |
| 352 | University of Bayreuth | 2 | 82 | 1.3K | 980 |
| 353 | Hokkaido University | 1 | 81 | 5.1K | 2.5K |
| 354 | Technical University of Braunschweig | 1 | 80 | 2.0K | 993 |
| 355 | University of Vermont | 1 | 79 | 3.5K | 1.8K |
| 355 | University of California, Santa Cruz | 1 | 79 | 1.1K | 681 |
| 357 | Chinese University of Hong Kong | 1 | 78 | 5.9K | 3.3K |
| 358 | University of Amsterdam | 2 | 77 | 5.3K | 4.0K |
| 358 | Guy's Hospital | 2 | 77 | 781 | 697 |
| 360 | Leiden University | 1 | 74 | 4.3K | 2.4K |
| 360 | Ruhr University Bochum | 1 | 74 | 4.0K | 2.2K |
| 362 | NUI Galway | 1 | 73 | 1.4K | 673 |
| 362 | Children's Hospital Los Angeles | 1 | 73 | 968 | 615 |
| 364 | Harvard Stem Cell Institute | 1 | 72 | 172 | 164 |
| 364 | Merck & Co. | 2 | 72 | 468 | 596 |
| 366 | Osaka Bioscience Institute | 1 | 71 | 123 | 111 |
| 367 | Simon Fraser University | 1 | 70 | 4.4K | 2.4K |
| 367 | National Institute of Neurological Disorders and Stroke | 1 | 70 | 426 | 359 |
| 367 | University of Texas Health Science Center at San Antonio | 1 | 70 | 2.2K | 1.4K |
| 370 | Technical University of Munich | 1 | 69 | 5.8K | 3.5K |
| 370 | University of Toledo | 1 | 69 | 2.4K | 1.2K |
| 370 | Japan Science and Technology Agency | 1 | 69 | 960 | 656 |
| 370 | University of Tennessee | 1 | 69 | 6.2K | 3.4K |
| 374 | Assistance Publique – Hôpitaux de Paris | 1 | 68 | 1.7K | 1.1K |
| 375 | RWTH Aachen University | 1 | 67 | 4.3K | 2.6K |
| 375 | University of Bremen | 1 | 67 | 2.6K | 1.4K |
| 375 | University of Bath | 1 | 67 | 3.6K | 2.1K |
| 375 | National Institute of Allergy and Infectious Diseases | 1 | 67 | 1.1K | 852 |
| 379 | National Centre for Biological Sciences | 2 | 66 | 160 | 155 |
| 380 | McMaster University | 1 | 65 | 6.7K | 4.0K |
| 380 | University of Insubria | 1 | 65 | 1.4K | 889 |
| 380 | Robert Larner College of Medicine | 1 | 65 | 903 | 596 |
| 380 | National Institute on Deafness and Other Communication Disorders | 1 | 65 | 184 | 166 |
| 380 | Mayo Clinic College of Medicine and Science | 1 | 65 | 1.5K | 1.1K |
| 385 | University of Missouri–Kansas City | 3 | 64 | 756 | 974 |
| 386 | University of Colorado Anschutz Medical Campus | 1 | 62 | 1.2K | 686 |
| 387 | Ulsan National Institute of Science and Technology | 1 | 61 | 668 | 350 |
| 387 | National Research Council Canada | 1 | 61 | 1.8K | 1.3K |
| 389 | University of Warwick | 1 | 60 | 5.2K | 3.1K |
| 390 | University of Chinese Academy of Sciences | 2 | 59 | 2.5K | 1.8K |
| 391 | University of Toulouse | 1 | 57 | 2.8K | 1.7K |
| 391 | Helmholtz Zentrum München | 2 | 57 | 655 | 773 |
| 391 | University of Namur | 1 | 57 | 806 | 537 |
| 391 | Kyoto Sangyo University | 1 | 57 | 382 | 172 |
| 391 | National Institute of Diabetes and Digestive and Kidney Diseases | 1 | 57 | 500 | 457 |
| 396 | Grenoble University - Grenoble Alpes University | 1 | 56 | 1.3K | 650 |
| 397 | Université libre de Bruxelles | 2 | 55 | 2.9K | 2.6K |
| 397 | Rice University | 1 | 55 | 3.1K | 2.1K |
| 397 | New York State Department of Health | 1 | 55 | 850 | 685 |
| 400 | Syracuse University | 2 | 54 | 2.4K | 2.1K |
| 400 | Champalimaud Foundation | 1 | 54 | 114 | 83 |
| 402 | Charité | 1 | 51 | 2.0K | 1.4K |
| 402 | Imperial College School of Medicine | 1 | 51 | 742 | 672 |
| 404 | Regeneron Pharmaceuticals | 1 | 50 | 168 | 169 |
| 405 | University of Potsdam | 1 | 49 | 2.0K | 1.3K |
| 405 | University of Jena | 2 | 49 | 2.1K | 2.1K |
| 405 | Pablo de Olavide University | 1 | 49 | 820 | 422 |
| 405 | Illinois State University | 1 | 49 | 1.8K | 1.0K |
| 405 | Georgia Institute of Technology | 1 | 49 | 4.5K | 3.2K |
| 405 | Augusta University | 1 | 49 | 1.4K | 830 |
| 411 | University of Strasbourg | 2 | 48 | 1.6K | 1.5K |
| 411 | Wuhan University | 1 | 48 | 4.0K | 2.6K |
| 413 | University of Connecticut | 1 | 47 | 6.0K | 3.9K |
| 413 | Zhejiang University | 1 | 47 | 6.1K | 4.1K |
| 413 | Ben-Gurion University of the Negev | 1 | 47 | 4.6K | 3.0K |
| 413 | Sinai Health System | 1 | 47 | 243 | 140 |
| 413 | Ben Gurion University of the Negev | 1 | 47 | 1.6K | 1.1K |
| 418 | University of Trieste | 1 | 46 | 2.8K | 1.9K |
| 418 | Churchill Hospital | 1 | 46 | 598 | 507 |
| 418 | John Radcliffe Hospital | 2 | 46 | 1.2K | 1.3K |
| 421 | University of Kaiserslautern | 1 | 45 | 1.1K | 796 |
| 421 | Cancer Research Center | 1 | 45 | 265 | 253 |
| 423 | University of Düsseldorf | 1 | 44 | 1.9K | 1.6K |
| 423 | Garvan Institute of Medical Research | 2 | 44 | 327 | 492 |
| 425 | Katholieke Universiteit Leuven | 1 | 43 | 7.2K | 5.1K |
| 425 | McLean Hospital | 1 | 43 | 606 | 562 |
| 427 | Newcastle University | 1 | 41 | 5.4K | 3.8K |
| 427 | University of Oviedo | 1 | 41 | 2.8K | 1.8K |
| 427 | Institut Jacques Monod | 2 | 41 | 104 | 185 |
| 427 | Université de Paris | 2 | 41 | 1.1K | 916 |
| 431 | Universitätsmedizin Göttingen | 3 | 40 | 333 | 553 |
| 431 | Merrimack Pharmaceuticals (United States) | 1 | 40 | 14 | 24 |
| 431 | Agency for Science, Technology and Research | 2 | 40 | 387 | 569 |
| 434 | West Virginia University | 1 | 39 | 4.3K | 2.9K |
| 434 | Technion – Israel Institute of Technology | 2 | 39 | 2.8K | 2.9K |
| 434 | Middlesex Hospital | 2 | 39 | 271 | 348 |
| 434 | University of Strathclyde | 1 | 39 | 3.8K | 2.7K |
| 434 | Centre d'Immunologie de Marseille-Luminy | 1 | 39 | 47 | 51 |
| 434 | Spanish National Research Council | 2 | 39 | 1.1K | 1.4K |
| 440 | University of Lyon | 1 | 38 | 866 | 745 |
| 440 | Freie Universität Berlin | 1 | 38 | 4.4K | 3.2K |
| 440 | Weill Cornell Medical College in Qatar | 1 | 38 | 63 | 93 |
| 443 | Neurocrine Biosciences | 1 | 37 | 49 | 77 |
| 444 | Vassar College | 1 | 36 | 485 | 314 |
| 444 | University of Montana | 1 | 36 | 1.5K | 1.1K |
| 444 | Albany Medical Center | 1 | 36 | 1.2K | 950 |
| 447 | Russell Berrie Nanotechnology Institute | 1 | 35 | 78 | 94 |
| 448 | Hospital Universitario de La Princesa | 1 | 34 | 467 | 391 |
| 449 | RIKEN | 1 | 33 | 1.5K | 1.2K |
| 449 | Virginia Commonwealth University | 1 | 33 | 4.6K | 3.6K |
| 449 | Indo-Zambia Bank Limited | 1 | 33 | 1 | 5 |
| 452 | Worcester Polytechnic Institute | 1 | 32 | 1.3K | 1.0K |
| 452 | Weldon School of Biomedical Engineering | 1 | 32 | 267 | 301 |
| 452 | University of Nevada, Reno | 1 | 32 | 4.3K | 3.1K |
| 455 | Jena University Hospital | 1 | 30 | 762 | 687 |
| 455 | Center for Infectious Disease Research | 1 | 30 | 110 | 175 |
| 455 | Sorbonne University | 2 | 30 | 1.6K | 1.5K |
| 458 | Temple University | 1 | 29 | 4.4K | 3.4K |
| 459 | University of Tübingen | 1 | 28 | 4.6K | 3.9K |
| 459 | Kyushu University | 1 | 28 | 5.6K | 4.3K |
| 459 | Hong Kong University of Science and Technology | 4 | 28 | 1.2K | 2.2K |
| 462 | University of Padua | 2 | 27 | 5.2K | 5.5K |
| 462 | Okayama University | 1 | 27 | 2.8K | 2.1K |
| 462 | University College Dublin | 1 | 27 | 4.2K | 3.6K |
| 462 | Asan Medical Center | 1 | 27 | 1.1K | 893 |
| 466 | Linköping University | 1 | 26 | 4.0K | 3.6K |
| 467 | University of London | 2 | 25 | 5.7K | 6.2K |
| 467 | Delft University of Technology | 1 | 25 | 3.7K | 3.3K |
| 467 | Kavli Institute of Nanoscience | 1 | 25 | 182 | 209 |
| 470 | University of Texas Health Science Center at Houston | 1 | 22 | 2.4K | 2.3K |
| 471 | Technische Universität Darmstadt | 1 | 21 | 2.4K | 2.3K |
| 471 | UK HealthCare | 1 | 21 | 673 | 723 |
| 471 | University Hospitals of Cleveland | 1 | 21 | 1.3K | 1.4K |
| 474 | University of Hong Kong | 1 | 20 | 6.7K | 6.0K |
| 474 | Collège de France | 1 | 20 | 608 | 778 |
| 474 | Paris Diderot University | 1 | 20 | 2.2K | 2.3K |
| 474 | Birkbeck, University of London | 1 | 20 | 1.4K | 1.5K |
| 478 | Tata Institute of Fundamental Research | 2 | 19 | 719 | 940 |
| 478 | Amgen | 1 | 19 | 661 | 804 |
| 478 | Curtin University | 1 | 19 | 4.3K | 4.2K |
| 478 | Great Ormond Street Hospital | 1 | 19 | 1.4K | 1.5K |
| 478 | Ikerbasque | 1 | 19 | 268 | 389 |
| 478 | University of Toronto Mississauga | 1 | 19 | 673 | 830 |
| 484 | University of Salamanca | 1 | 18 | 2.7K | 2.3K |
| 484 | Université catholique de Louvain | 1 | 18 | 4.5K | 4.5K |
| 484 | University of Exeter | 1 | 18 | 4.4K | 4.2K |
| 484 | Aix-Marseille University | 1 | 18 | 3.0K | 2.9K |
| 484 | St George's, University of London | 1 | 18 | 914 | 1.1K |
| 489 | Tata Institute of Fundamental Research Hyderabad | 1 | 17 | 45 | 53 |
| 489 | King's College | 1 | 17 | 2.5K | 2.7K |
| 491 | Moscow State University | 1 | 16 | 3.8K | 3.1K |
| 491 | Goethe University Frankfurt | 1 | 16 | 3.8K | 3.8K |
| 491 | University of Montpellier | 1 | 16 | 2.4K | 2.5K |
| 494 | Tulane University | 1 | 14 | 3.2K | 3.4K |
| 494 | University of Côte d'Azur | 2 | 14 | 392 | 806 |
| 494 | Sorbonne Universités | 1 | 14 | 1.3K | 1.7K |
| 497 | Wellcome Sanger Institute | 1 | 12 | 433 | 646 |
| 497 | Fujita Health University | 1 | 12 | 932 | 1.0K |
| 499 | University of Fribourg | 1 | 11 | 1.6K | 2.0K |
| 499 | University of São Paulo | 1 | 11 | 9.4K | 8.1K |
| 499 | University of Texas Medical Branch | 1 | 11 | 2.9K | 3.2K |
| 502 | University of Miami | 1 | 10 | 5.3K | 5.8K |
| 502 | University of Paris-Sud | 1 | 10 | 2.1K | 2.7K |
| 504 | Medical University of South Carolina | 1 | 9 | 3.3K | 3.7K |
| 505 | Humboldt University of Berlin | 1 | 8 | 4.7K | 5.4K |
| 505 | Durham University | 1 | 8 | 4.0K | 4.7K |
| 505 | University of Florida | 1 | 8 | 9.6K | 10.0K |
| 508 | University of Kansas | 1 | 7 | 5.7K | 6.6K |
| 508 | Merck KGaA | 3 | 7 | 136 | 612 |
| 508 | Institut de recherches cliniques de Montréal (IRCM) | 1 | 7 | 246 | 415 |
| 511 | University of Bonn | 1 | 6 | 4.5K | 5.5K |
| 511 | Kindai University | 1 | 6 | 2.1K | 2.5K |
| 511 | UCL Medical School | 1 | 6 | 1.2K | 1.8K |
| 511 | University of California, Santa Barbara | 1 | 6 | 2.3K | 3.2K |
| 515 | University of Melbourne | 1 | 4 | 9.1K | 10.7K |
| 515 | ComUE Paris-Saclay University | 1 | 4 | 2.8K | 3.7K |
| 517 | University of Kent | 1 | 3 | 2.9K | 4.2K |
| 518 | Indian Institutes of Science Education and Research | 1 | 2 | 483 | 959 |
| 518 | MRC Laboratory for Molecular Cell Biology | 1 | 2 | 3 | 13 |
| 518 | Newcastle General Hospital | 1 | 2 | 445 | 732 |
| 521 | Barcelona Institute for Science and Technology | 1 | 0 | 133 | 324 |
| 521 | Walloon Excellence in Lifesciences and Biotechnology | 1 | 0 | 16 | 57 |
