| 1 | University of California | 425 | 32.5K | 325 | 298 |
| 2 | University of California, Davis | 61 | 8.4K | 129 | 35 |
| 3 | University of Oxford | 88 | 8.0K | 490 | 321 |
| 4 | Indiana University | 94 | 7.7K | 246 | 150 |
| 5 | Cornell University | 113 | 7.7K | 226 | 234 |
| 6 | Harvard University | 66 | 7.6K | 1456 | 750 |
| 7 | Duke University | 83 | 7.4K | 263 | 175 |
| 8 | University of Texas at Austin | 82 | 7.3K | 553 | 330 |
| 9 | University of British Columbia | 85 | 7.3K | 380 | 225 |
| 10 | University of Arizona | 79 | 6.9K | 235 | 134 |
| 11 | University of Minnesota | 65 | 6.8K | 679 | 315 |
| 12 | University of Washington | 45 | 6.2K | 1199 | 487 |
| 13 | Princeton University | 74 | 6.2K | 298 | 249 |
| 14 | University of Edinburgh | 57 | 6.1K | 422 | 193 |
| 15 | Uppsala University | 41 | 6.1K | 524 | 138 |
| 16 | Madras Christian College | 61 | 5.8K | 404 | 175 |
| 17 | Imperial College London | 59 | 5.8K | 724 | 343 |
| 18 | University of North Carolina | 51 | 5.4K | 718 | 339 |
| 19 | University of North Carolina at Chapel Hill | 45 | 5.2K | 798 | 353 |
| 20 | McGill University | 46 | 5.1K | 726 | 281 |
| 21 | Marie Curie | 44 | 5.1K | 323 | 136 |
| 22 | Stanford University | 59 | 5.1K | 960 | 678 |
| 23 | University of Florida | 70 | 4.6K | 505 | 325 |
| 24 | University of Queensland | 56 | 4.5K | 596 | 316 |
| 25 | University of Michigan | 65 | 4.5K | 876 | 637 |
| 26 | University of Toronto | 99 | 4.4K | 588 | 625 |
| 27 | University of Wisconsin–Madison | 41 | 4.0K | 1167 | 548 |
| 28 | University of Chicago | 77 | 3.7K | 386 | 461 |
| 29 | University of Tennessee | 34 | 3.7K | 461 | 162 |
| 30 | Florida State University | 58 | 3.6K | 144 | 120 |
| 31 | Yale University | 49 | 3.6K | 675 | 565 |
| 32 | University of Helsinki | 59 | 3.6K | 384 | 305 |
| 33 | University of Sheffield | 49 | 3.5K | 350 | 212 |
| 34 | Arizona State University | 37 | 3.3K | 432 | 216 |
| 35 | University of Groningen | 36 | 3.3K | 561 | 286 |
| 36 | Michigan State University | 53 | 3.3K | 397 | 299 |
| 37 | Lund University | 51 | 3.2K | 459 | 361 |
| 38 | Colorado State University | 22 | 3.0K | 553 | 176 |
| 39 | University of Maryland | 35 | 2.9K | 1088 | 581 |
| 40 | Spanish National Research Council | 37 | 2.8K | 166 | 100 |
| 41 | Washington State University | 31 | 2.7K | 345 | 172 |
| 42 | Indiana State University | 13 | 2.7K | 46 | 3 |
| 42 | University of Montana | 47 | 2.7K | 34 | 46 |
| 44 | University of Zurich | 53 | 2.7K | 230 | 265 |
| 45 | University of New Mexico | 37 | 2.6K | 242 | 159 |
| 46 | University of Amsterdam | 40 | 2.6K | 628 | 493 |
| 47 | University of New South Wales | 37 | 2.6K | 820 | 479 |
| 48 | Wageningen University & Research | 31 | 2.6K | 442 | 305 |
| 49 | Natural Environment Research Council | 23 | 2.5K | 32 | 23 |
| 50 | Stony Brook University | 24 | 2.5K | 430 | 209 |
| 51 | University of Missouri | 34 | 2.5K | 678 | 300 |
| 52 | Carolina University | 7 | 2.4K | 844 | 28 |
| 53 | University of Melbourne | 33 | 2.3K | 1189 | 716 |
| 54 | Simon Fraser University | 36 | 2.3K | 158 | 102 |
| 55 | Dartmouth College | 26 | 2.3K | 157 | 117 |
| 56 | Utrecht University | 21 | 2.3K | 1090 | 517 |
| 57 | National Center for Ecological Analysis and Synthesis | 13 | 2.2K | 13 | 13 |
| 58 | University of Georgia | 33 | 2.2K | 603 | 333 |
| 59 | Korea University of Science and Technology | 36 | 2.2K | 3165 | 1449 |
| 60 | Santa Fe Institute | 22 | 2.1K | 32 | 31 |
| 61 | Australian National University | 27 | 2.1K | 621 | 351 |
| 62 | ECOLOG | 41 | 2.1K | 203 | 106 |
| 63 | University of Cambridge | 37 | 2.0K | 1041 | 927 |
| 64 | North Carolina State University | 20 | 2.0K | 909 | 359 |
| 65 | University of Virginia | 35 | 1.9K | 492 | 452 |
| 66 | Pennsylvania State University | 23 | 1.9K | 1527 | 765 |
| 67 | University of Colorado | 30 | 1.9K | 801 | 605 |
| 68 | Coastal Carolina University | 2 | 1.9K | 161 | 1 |
| 69 | Lone Star College System | 22 | 1.9K | 1191 | 640 |
| 70 | Montpellier 2 University | 15 | 1.8K | 268 | 112 |
| 71 | King's Buildings | 14 | 1.8K | 144 | 69 |
| 72 | University of Utah | 24 | 1.8K | 1031 | 566 |
| 73 | University of Oslo | 30 | 1.8K | 678 | 462 |
| 74 | University of Lausanne | 37 | 1.8K | 158 | 197 |
| 75 | Museum of Vertebrate Zoology | 17 | 1.8K | 26 | 22 |
| 76 | Arnold Arboretum | 4 | 1.7K | 32 | 6 |
| 77 | Biology Institute of Shandong Academy of Sciences | 27 | 1.7K | 414 | 202 |
| 78 | United States Geological Survey | 28 | 1.7K | 510 | 331 |
| 79 | University of Connecticut | 21 | 1.6K | 716 | 371 |
| 80 | Smithsonian Tropical Research Institute | 27 | 1.6K | 47 | 48 |
| 81 | Institute for Biodiversity and Ecosystem Dynamics | 31 | 1.6K | 11 | 26 |
| 82 | Department of Mathematical Sciences | 26 | 1.6K | 449 | 160 |
| 83 | University of Liverpool | 27 | 1.6K | 709 | 460 |
| 84 | University of Idaho | 24 | 1.6K | 140 | 91 |
| 85 | Bio-Medical Science (South Korean) | 12 | 1.5K | 3156 | 1205 |
| 86 | University of Pittsburgh | 24 | 1.5K | 1422 | 1017 |
| 87 | Phillips Exeter Academy | 31 | 1.5K | 383 | 356 |
| 88 | University of Cape Town | 11 | 1.5K | 1044 | 274 |
| 89 | National Evolutionary Synthesis Center | 4 | 1.4K | 17 | 7 |
| 90 | University of Calgary | 20 | 1.4K | 1120 | 612 |
| 91 | University of Bristol | 31 | 1.4K | 678 | 686 |
| 92 | Utah State University | 24 | 1.4K | 203 | 140 |
| 93 | James Cook University | 23 | 1.4K | 186 | 133 |
| 94 | University of Guelph | 26 | 1.4K | 386 | 297 |
| 95 | Eastern Michigan University | 2 | 1.4K | 559 | 2 |
| 96 | University of Alberta | 27 | 1.4K | 1237 | 913 |
| 97 | Oregon State University | 20 | 1.3K | 642 | 413 |
| 98 | United States Forest Service | 13 | 1.3K | 402 | 201 |
| 99 | University of Reading | 11 | 1.3K | 795 | 283 |
| 100 | University of Massachusetts | 18 | 1.3K | 1014 | 661 |
| 101 | Stockholm University | 22 | 1.3K | 543 | 436 |
| 102 | Texas A&M University | 22 | 1.3K | 1426 | 878 |
| 103 | Brown University | 19 | 1.3K | 883 | 591 |
| 104 | University of Basel | 16 | 1.2K | 730 | 428 |
| 105 | Leiden University | 22 | 1.2K | 940 | 777 |
| 106 | University of Jyväskylä | 20 | 1.2K | 201 | 112 |
| 107 | Stellenbosch University | 8 | 1.1K | 1075 | 189 |
| 108 | National Institute for Mathematical and Biological Synthesis | 6 | 1.1K | 10 | 2 |
| 109 | Ohio State University | 20 | 1.1K | 2723 | 1721 |
| 110 | Monash University | 33 | 1.1K | 977 | 1124 |
| 111 | Max Planck Society | 9 | 1.1K | 1839 | 944 |
| 112 | Louisiana State University | 18 | 1.1K | 869 | 500 |
| 113 | Max Planck Institute for Ornithology | 16 | 1.1K | 29 | 21 |
| 114 | Rice University | 32 | 1.1K | 211 | 353 |
| 115 | Auburn University | 19 | 1.1K | 445 | 214 |
| 116 | University of Oklahoma | 14 | 1.1K | 944 | 426 |
| 117 | University of Alabama | 5 | 1.1K | 2126 | 332 |
| 118 | University of Kansas | 25 | 1.1K | 407 | 388 |
| 119 | University of Ottawa | 18 | 1.1K | 1007 | 609 |
| 120 | Université de Sherbrooke | 9 | 1.1K | 693 | 165 |
| 121 | University of Sydney | 25 | 1.0K | 1701 | 1441 |
| 121 | University of Miami | 15 | 1.0K | 1100 | 613 |
| 123 | Rocky Mountain Research Station | 5 | 1.0K | 146 | 53 |
| 124 | University College London | 19 | 1.0K | 1925 | 1446 |
| 125 | University of Fribourg | 10 | 1.0K | 225 | 121 |
| 126 | Columbia University | 14 | 1.0K | 2715 | 1658 |
| 127 | Umeå University | 24 | 996 | 381 | 446 |
| 128 | Statistics New Zealand | 16 | 991 | 1265 | 675 |
| 129 | Miami University | 5 | 974 | 791 | 104 |
| 130 | Scripps Institution of Oceanography | 6 | 964 | 414 | 191 |
| 131 | Ludwig Maximilian University of Munich | 9 | 929 | 2933 | 1189 |
| 132 | Zoological Society of London | 12 | 927 | 75 | 48 |
| 133 | University of Copenhagen | 7 | 908 | 3280 | 1275 |
| 133 | Claude Bernard University Lyon 1 | 14 | 908 | 956 | 526 |
| 135 | University of Kentucky | 12 | 906 | 1741 | 759 |
| 136 | Virginia Tech | 16 | 896 | 1102 | 690 |
| 137 | Halifax | 12 | 891 | 1243 | 630 |
| 138 | University of Bern | 11 | 890 | 1514 | 788 |
| 139 | Finnish Museum of Natural History | 10 | 887 | 42 | 19 |
| 140 | Morton Arboretum | 4 | 870 | 37 | 3 |
| 141 | Iowa State University | 19 | 861 | 798 | 730 |
| 142 | University of Aberdeen | 21 | 858 | 477 | 549 |
| 143 | University of Sussex | 19 | 857 | 402 | 446 |
| 144 | University of Leeds | 14 | 851 | 1499 | 918 |
| 145 | Barnard College | 2 | 835 | 433 | 22 |
| 146 | University of Turku | 10 | 833 | 1028 | 465 |
| 147 | Smithsonian Environmental Research Center | 3 | 823 | 407 | 98 |
| 148 | Bernardo O'Higgins University | 15 | 818 | 5296 | 1750 |
| 149 | Aarhus University | 14 | 816 | 1345 | 920 |
| 150 | University of Bergen | 10 | 813 | 1132 | 591 |
| 151 | University of Montpellier | 20 | 807 | 525 | 443 |
| 152 | Max Planck Institute for Evolutionary Anthropology | 6 | 804 | 116 | 72 |
| 153 | University of Oregon | 9 | 801 | 819 | 421 |
| 154 | Wesleyan University | 4 | 798 | 424 | 65 |
| 155 | ETH Zürich | 15 | 792 | 1032 | 922 |
| 156 | Deakin University | 11 | 787 | 983 | 441 |
| 157 | University of Illinois at Urbana–Champaign | 14 | 781 | 1316 | 976 |
| 158 | Case Western Reserve University | 22 | 773 | 943 | 999 |
| 159 | Katholieke Universiteit Leuven | 11 | 772 | 2411 | 1382 |
| 160 | Georgia Institute of Technology | 14 | 769 | 1120 | 838 |
| 161 | Idaho State University | 1 | 746 | 1083 | 20 |
| 162 | Tel Aviv University | 18 | 738 | 1412 | 1164 |
| 163 | University of East Anglia | 9 | 735 | 867 | 476 |
| 164 | University of California, San Diego | 6 | 734 | 2303 | 1002 |
| 165 | University of Pretoria | 10 | 733 | 880 | 278 |
| 166 | University of South Florida | 6 | 727 | 2402 | 736 |
| 167 | Nicholas School of the Environment | 6 | 713 | 178 | 101 |
| 168 | Laval University | 10 | 701 | 1532 | 792 |
| 169 | University of Western Australia | 10 | 694 | 1835 | 1057 |
| 170 | Museo Nacional de Ciencias Naturales | 11 | 693 | 98 | 63 |
| 171 | Helmholtz Centre for Environmental Research | 9 | 690 | 209 | 148 |
| 172 | Northern Arizona University | 9 | 686 | 239 | 139 |
| 173 | University of Basilicata | 6 | 676 | 177 | 51 |
| 173 | University of Wyoming | 12 | 676 | 379 | 262 |
| 175 | University of Otago | 14 | 667 | 845 | 660 |
| 176 | Institute for Advanced Study | 7 | 659 | 981 | 408 |
| 176 | Harvey Mudd College | 2 | 659 | 174 | 14 |
| 178 | University of Vermont | 9 | 655 | 723 | 437 |
| 178 | Commonwealth Scientific and Industrial Research Organisation | 8 | 655 | 128 | 86 |
| 180 | Ben-Gurion University of the Negev | 13 | 654 | 752 | 452 |
| 181 | Kyoto University | 21 | 653 | 1791 | 1474 |
| 182 | Carleton University | 16 | 650 | 337 | 297 |
| 183 | University of London | 9 | 637 | 3214 | 1640 |
| 184 | University of Granada | 8 | 630 | 1560 | 535 |
| 185 | Hebrew University of Jerusalem | 9 | 629 | 1586 | 854 |
| 186 | International Institute for Applied Systems Analysis | 12 | 626 | 84 | 103 |
| 187 | Indian Institute of Science | 9 | 625 | 999 | 425 |
| 188 | Penryn Campus | 18 | 623 | 29 | 52 |
| 189 | University of Stirling | 13 | 619 | 267 | 252 |
| 190 | San Francisco State University | 7 | 617 | 247 | 112 |
| 191 | University of Glasgow | 13 | 613 | 1495 | 1127 |
| 191 | National Autonomous University of Mexico | 16 | 613 | 1116 | 643 |
| 191 | Rocky Mountain Biological Laboratory | 11 | 613 | 19 | 22 |
| 194 | Peter Wall Institute for Advanced Studies | 4 | 596 | 1 | 1 |
| 195 | University of Texas at Arlington | 5 | 595 | 880 | 220 |
| 195 | Portland State University | 5 | 595 | 604 | 163 |
| 197 | Swedish University of Agricultural Sciences | 10 | 592 | 644 | 488 |
| 198 | State University of New York | 14 | 591 | 1683 | 1426 |
| 199 | University of Waterloo | 4 | 584 | 3062 | 957 |
| 199 | Patuxent Wildlife Research Center | 3 | 584 | 305 | 86 |
| 201 | National Marine Fisheries Service | 10 | 579 | 179 | 135 |
| 202 | University of Barcelona | 5 | 574 | 3522 | 1291 |
| 202 | Marine Science Institute | 9 | 574 | 152 | 124 |
| 204 | Boston University | 9 | 567 | 2186 | 1477 |
| 205 | Eawag | 10 | 565 | 84 | 95 |
| 206 | Binghamton University | 6 | 560 | 504 | 192 |
| 207 | University of Gothenburg | 9 | 556 | 1868 | 1223 |
| 208 | Drake University | 1 | 553 | 407 | 17 |
| 208 | Truman State University | 1 | 553 | 198 | 6 |
| 210 | Czech Academy of Sciences | 11 | 551 | 1159 | 711 |
| 210 | Massey University | 4 | 551 | 1466 | 391 |
| 212 | Siena College | 1 | 548 | 194 | 11 |
| 213 | Centre for Ecology & Hydrology | 9 | 542 | 194 | 182 |
| 214 | New Mexico State University | 4 | 540 | 866 | 216 |
| 215 | Purdue University | 8 | 537 | 3171 | 1783 |
| 216 | Queen Mary University of London | 5 | 527 | 1855 | 711 |
| 217 | Royal Holloway University of London | 3 | 523 | 276 | 95 |
| 218 | Microbial Ecology | 7 | 521 | 394 | 295 |
| 219 | Cornell Lab of Ornithology | 15 | 514 | 33 | 42 |
| 220 | University of Southampton | 4 | 498 | 3410 | 1293 |
| 220 | Emory University | 12 | 498 | 1960 | 1648 |
| 222 | Georgia Southern University | 5 | 492 | 310 | 64 |
| 222 | Southern University | 5 | 492 | 1356 | 386 |
| 224 | Smithsonian Institution | 11 | 490 | 234 | 199 |
| 225 | University of Würzburg | 13 | 483 | 862 | 836 |
| 226 | Howard Hughes Medical Institute | 4 | 482 | 863 | 599 |
| 226 | Kenyon College | 3 | 482 | 81 | 30 |
| 228 | Pontifical Catholic University of Chile | 10 | 476 | 591 | 330 |
| 229 | Allan Wilson Centre | 2 | 474 | 56 | 25 |
| 230 | East Carolina University | 5 | 473 | 944 | 324 |
| 231 | University of Vienna | 9 | 468 | 2951 | 1864 |
| 231 | Medway School of Science | 15 | 468 | 2530 | 1865 |
| 233 | autonomous university | 2 | 466 | 2008 | 250 |
| 234 | Rutgers University | 12 | 465 | 2323 | 1954 |
| 235 | Swansea University | 3 | 463 | 1923 | 484 |
| 235 | Union College | 4 | 463 | 152 | 27 |
| 237 | Victoria University of Wellington | 7 | 452 | 629 | 289 |
| 238 | University of São Paulo | 17 | 448 | 2879 | 2179 |
| 239 | University of Wisconsin–Milwaukee | 5 | 444 | 1032 | 384 |
| 240 | University of Oulu | 6 | 440 | 1362 | 686 |
| 241 | Department of Conservation | 8 | 439 | 201 | 133 |
| 242 | United States Department of Agriculture | 4 | 436 | 1064 | 577 |
| 243 | Massachusetts Institute of Technology | 6 | 434 | 2741 | 1965 |
| 243 | University of Jena | 4 | 434 | 1496 | 532 |
| 243 | King Saud University | 1 | 434 | 5466 | 876 |
| 246 | Vanderbilt University | 5 | 431 | 3432 | 1780 |
| 247 | Lincoln University | 5 | 428 | 327 | 140 |
| 247 | Technical University of Denmark | 10 | 428 | 1280 | 1112 |
| 249 | Okinawa Prefecture | 7 | 425 | 626 | 273 |
| 250 | McMaster University | 8 | 424 | 2421 | 1585 |
| 251 | Vrije Universiteit Amsterdam | 8 | 421 | 2840 | 2089 |
| 252 | Dakota State University | 8 | 418 | 670 | 451 |
| 253 | Kobe University | 5 | 417 | 1980 | 664 |
| 254 | University of Hawaii at Manoa | 9 | 415 | 327 | 333 |
| 255 | Field Museum of Natural History | 7 | 413 | 124 | 98 |
| 256 | University of KwaZulu-Natal | 6 | 411 | 1100 | 431 |
| 257 | Douglas | 4 | 409 | 433 | 143 |
| 258 | University of Natal | 1 | 405 | 932 | 110 |
| 259 | Agricultural Research Service | 3 | 404 | 1793 | 924 |
| 260 | Smithsonian Conservation Biology Institute | 6 | 402 | 89 | 54 |
| 261 | Berry College | 3 | 395 | 68 | 9 |
| 262 | Kyushu University | 12 | 394 | 1934 | 1356 |
| 263 | Oklahoma State University | 9 | 393 | 778 | 576 |
| 264 | National Zoological Park | 5 | 392 | 108 | 74 |
| 265 | Jacob Blaustein Institutes for Desert Research | 3 | 385 | 204 | 82 |
| 266 | Royal Botanic Gardens | 4 | 384 | 248 | 127 |
| 267 | Universidade Estadual Paulista | 7 | 383 | 1923 | 624 |
| 268 | Swiss Institute of Bioinformatics | 7 | 382 | 139 | 153 |
| 269 | University of Antwerp | 5 | 381 | 1897 | 981 |
| 270 | University of Auckland | 10 | 380 | 1656 | 1507 |
| 271 | Scott Polar Research Institute | 1 | 379 | 97 | 26 |
| 271 | Met Office | 1 | 379 | 402 | 121 |
| 273 | Concordia University | 3 | 377 | 2002 | 559 |
| 274 | Western Sydney University | 1 | 376 | 1978 | 311 |
| 275 | Flinders University | 7 | 372 | 1115 | 695 |
| 276 | Brigham Young University | 6 | 371 | 1019 | 629 |
| 277 | University of Prince Edward Island | 1 | 370 | 699 | 52 |
| 278 | Université du Québec à Rimouski | 7 | 369 | 110 | 68 |
| 279 | Hopkins Marine Station | 9 | 365 | 54 | 66 |
| 280 | Syracuse University | 6 | 363 | 1104 | 634 |
| 281 | National University of Singapore | 3 | 359 | 5516 | 2268 |
| 282 | University of Tartu | 5 | 353 | 1004 | 489 |
| 283 | University of Louisiana at Lafayette | 7 | 350 | 177 | 95 |
| 284 | Illinois State University | 9 | 346 | 335 | 274 |
| 284 | Norwegian University of Life Sciences | 1 | 346 | 1522 | 357 |
| 284 | Bioforsk | 1 | 346 | 152 | 41 |
| 287 | College of Charleston | 5 | 343 | 164 | 77 |
| 288 | Queen's University Belfast | 4 | 342 | 2188 | 951 |
| 288 | University of Mississippi | 4 | 342 | 990 | 377 |
| 290 | Bucknell University | 4 | 341 | 209 | 62 |
| 291 | Charles University | 8 | 339 | 1905 | 1223 |
| 292 | North Dakota State University | 6 | 338 | 620 | 378 |
| 292 | Chicago Botanic Garden | 6 | 338 | 20 | 21 |
| 294 | Bio21 Institute | 4 | 336 | 112 | 83 |
| 295 | California State University | 7 | 335 | 1531 | 841 |
| 296 | Tohoku University | 10 | 331 | 2295 | 1670 |
| 297 | University of Naples Federico II | 6 | 326 | 3047 | 1903 |
| 298 | University of Michigan Herbarium | 1 | 322 | 27 | 5 |
| 299 | Newcastle University | 4 | 318 | 3116 | 1590 |
| 299 | University of Wales | 5 | 318 | 1183 | 809 |
| 301 | Chinese Academy of Sciences | 7 | 316 | 4862 | 3369 |
| 302 | Wilfrid Laurier University | 4 | 314 | 715 | 320 |
| 303 | Florida Museum of Natural History | 8 | 309 | 89 | 81 |
| 304 | British Antarctic Survey | 6 | 305 | 222 | 186 |
| 305 | University of Maine | 5 | 303 | 684 | 440 |
| 306 | University of Roehampton | 2 | 300 | 309 | 46 |
| 307 | Queensland Brain Institute | 2 | 295 | 278 | 136 |
| 307 | National Oceanography Centre, Southampton | 1 | 295 | 144 | 55 |
| 309 | Victoria University | 5 | 294 | 1552 | 785 |
| 309 | National Research Foundation | 3 | 294 | 132 | 88 |
| 309 | Department of Science and Technology | 3 | 294 | 779 | 241 |
| 312 | University of Bayreuth | 4 | 292 | 668 | 384 |
| 313 | University of Arkansas | 9 | 291 | 678 | 662 |
| 314 | Université de Lyon | 9 | 289 | 711 | 737 |
| 315 | Leibniz Association | 7 | 286 | 865 | 738 |
| 316 | Natural History Museum | 7 | 285 | 641 | 482 |
| 317 | University of Tokyo | 12 | 284 | 3531 | 3066 |
| 317 | Hokkaido University | 6 | 284 | 2834 | 1659 |
| 319 | California Institute of Technology | 4 | 283 | 1803 | 1299 |
| 320 | King Juan Carlos University | 4 | 282 | 839 | 294 |
| 321 | University of Tübingen | 5 | 281 | 1924 | 1297 |
| 322 | University of Adelaide | 7 | 280 | 2221 | 1820 |
| 323 | University of Toronto Mississauga | 5 | 279 | 221 | 165 |
| 324 | Agropolis International | 4 | 276 | 126 | 100 |
| 325 | Macaulay Institute | 5 | 274 | 112 | 118 |
| 326 | Clemson University | 4 | 273 | 1807 | 867 |
| 327 | University of York | 9 | 272 | 1081 | 1179 |
| 327 | Washington University School of Medicine | 4 | 272 | 2434 | 1612 |
| 329 | Canadian Institute for Advanced Research | 2 | 271 | 181 | 131 |
| 330 | University of Campinas | 5 | 270 | 3305 | 1435 |
| 331 | Memorial Sloan Kettering Cancer Center | 3 | 269 | 1664 | 960 |
| 332 | Pontificia Universidad Católica del Ecuador | 4 | 265 | 158 | 36 |
| 333 | University of Southern California | 5 | 264 | 4229 | 2878 |
| 334 | Ohio University | 9 | 254 | 451 | 584 |
| 335 | Max Planck Institute for Evolutionary Biology | 9 | 252 | 28 | 59 |
| 336 | Evolution Group | 10 | 250 | 16 | 60 |
| 337 | St. John's College | 4 | 248 | 67 | 30 |
| 338 | University of Delaware | 2 | 246 | 3340 | 1328 |
| 339 | Johannes Gutenberg University Mainz | 3 | 245 | 1928 | 977 |
| 340 | Georgetown University | 3 | 243 | 2399 | 1058 |
| 340 | Woods Hole Oceanographic Institution | 8 | 243 | 238 | 321 |
| 342 | Polytechnic University of Milan | 6 | 241 | 1504 | 1032 |
| 343 | Royal Veterinary and Agricultural University | 1 | 240 | 732 | 328 |
| 344 | Bennett College | 2 | 239 | 328 | 101 |
| 345 | University of Konstanz | 7 | 237 | 674 | 723 |
| 346 | National Institute of Water and Atmospheric Research | 4 | 236 | 235 | 193 |
| 347 | University of the Ryukyus | 2 | 235 | 1366 | 315 |
| 348 | Carnegie Institution for Science | 1 | 232 | 794 | 357 |
| 349 | Museums Victoria | 5 | 229 | 46 | 38 |
| 349 | International Atomic Energy Agency | 2 | 229 | 386 | 153 |
| 351 | University of Birmingham | 3 | 228 | 4997 | 2579 |
| 352 | Saga University | 1 | 227 | 1737 | 265 |
| 352 | University of Saskatchewan | 5 | 227 | 2053 | 1453 |
| 354 | Russian Academy of Sciences | 2 | 226 | 6146 | 2205 |
| 355 | Tufts University | 5 | 225 | 2158 | 1747 |
| 355 | University of Regina | 5 | 225 | 465 | 301 |
| 355 | Daniel B. Warnell School of Forestry and Natural Resources | 2 | 225 | 242 | 106 |
| 358 | Durham University | 7 | 222 | 1298 | 1215 |
| 359 | Galton Laboratory | 4 | 221 | 50 | 55 |
| 360 | Schiller International University | 3 | 219 | 599 | 392 |
| 360 | Juniata College | 1 | 219 | 117 | 21 |
| 362 | James Hutton Institute | 2 | 218 | 437 | 235 |
| 363 | Max Planck Institute for Demographic Research | 2 | 216 | 114 | 59 |
| 364 | The Nature Conservancy | 4 | 215 | 210 | 167 |
| 365 | Linnaeus University | 12 | 212 | 467 | 812 |
| 365 | Centre for Australian National Biodiversity Research | 1 | 212 | 55 | 24 |
| 367 | University of Neuchâtel | 7 | 211 | 231 | 329 |
| 368 | Drexel University | 3 | 210 | 2400 | 1325 |
| 369 | University of Pennsylvania | 7 | 207 | 4473 | 4042 |
| 369 | Ministry of Foreign Affairs of the Republic of Indonesia | 2 | 207 | 124 | 4 |
| 371 | University of Massachusetts Amherst | 4 | 206 | 1032 | 755 |
| 372 | University of Maryland, Baltimore County | 1 | 204 | 1163 | 319 |
| 373 | Audubon | 3 | 203 | 120 | 70 |
| 374 | La Trobe University | 5 | 201 | 1731 | 1340 |
| 375 | Illinois State Museum | 3 | 199 | 27 | 14 |
| 376 | Purdue University Northwest | 1 | 198 | 312 | 36 |
| 377 | University of Pavia | 3 | 197 | 2584 | 1415 |
| 378 | University of Padua | 3 | 196 | 4655 | 2614 |
| 378 | Paris Diderot University | 4 | 196 | 2229 | 1643 |
| 380 | New York Botanical Garden | 1 | 193 | 208 | 82 |
| 380 | AgroParisTech, Paris-Saclay University | 2 | 193 | 736 | 377 |
| 382 | Université de Montréal | 3 | 191 | 4285 | 2436 |
| 383 | Okinawa Institute of Science and Technology | 5 | 190 | 165 | 158 |
| 384 | University of South Carolina | 4 | 188 | 4080 | 2812 |
| 384 | University of Victoria | 6 | 188 | 1202 | 1211 |
| 384 | Burke Museum of Natural History and Culture | 5 | 188 | 44 | 55 |
| 387 | Texas Tech University | 6 | 187 | 1349 | 1193 |
| 387 | University of Lethbridge | 4 | 187 | 406 | 310 |
| 389 | Wuhan University | 4 | 186 | 3341 | 2039 |
| 390 | German Centre for Integrative Biodiversity Research | 5 | 185 | 116 | 137 |
| 390 | Universidade do Porto | 4 | 185 | 1514 | 984 |
| 392 | University of Illinois at Chicago | 8 | 184 | 1925 | 2331 |
| 392 | University of South Bohemia in České Budějovice | 3 | 184 | 505 | 307 |
| 392 | African Institute for Mathematical Sciences | 1 | 184 | 156 | 24 |
| 395 | Johns Hopkins University | 4 | 183 | 5016 | 3665 |
| 396 | University of Liège | 3 | 182 | 2313 | 1398 |
| 397 | Department of Forestry | 5 | 181 | 671 | 497 |
| 398 | University of Milan | 2 | 180 | 5384 | 2679 |
| 398 | Muséum national d'histoire naturelle | 6 | 180 | 442 | 465 |
| 398 | Slovak Academy of Sciences | 2 | 180 | 1900 | 776 |
| 401 | University of Paris-Sud | 2 | 179 | 2752 | 1577 |
| 401 | Durrell Wildlife Conservation Trust | 1 | 179 | 62 | 21 |
| 401 | Mauritian Wildlife Foundation | 1 | 179 | 23 | 6 |
| 404 | San Diego State University | 3 | 176 | 1959 | 1147 |
| 405 | Mammal Research Institute of the Polish Academy of Sciences | 5 | 175 | 103 | 95 |
| 406 | Industrial Research Limited | 3 | 174 | 2064 | 1214 |
| 407 | Institut de chimie | 2 | 173 | 975 | 540 |
| 408 | National Museums of Kenya | 3 | 171 | 84 | 67 |
| 409 | Gonzaga University | 4 | 170 | 108 | 56 |
| 410 | College of the Atlantic | 1 | 169 | 48 | 19 |
| 411 | Boise State University | 2 | 167 | 848 | 300 |
| 411 | Cancer Research Center | 2 | 167 | 2680 | 1571 |
| 413 | University of Nicosia | 3 | 166 | 290 | 171 |
| 413 | Missouri Botanical Garden | 2 | 166 | 149 | 98 |
| 413 | Generalitat Valenciana | 1 | 166 | 178 | 44 |
| 416 | University of Münster | 4 | 165 | 2369 | 1800 |
| 416 | Florida International University | 5 | 165 | 1335 | 1182 |
| 418 | Polytechnic University of Turin | 2 | 164 | 2086 | 928 |
| 418 | Middle East Technical University | 1 | 164 | 2760 | 782 |
| 418 | University of California, Los Angeles | 1 | 164 | 5041 | 2195 |
| 418 | National Estuarine Research Reserve Association | 1 | 164 | 84 | 29 |
| 422 | University of Debrecen | 2 | 163 | 1733 | 661 |
| 423 | University of Nevada, Reno | 8 | 161 | 1057 | 1514 |
| 424 | University of Hawaii | 4 | 160 | 1420 | 1300 |
| 425 | Buck Institute for Research on Aging | 1 | 159 | 260 | 169 |
| 426 | National Science Foundation | 4 | 157 | 262 | 283 |
| 426 | University of Havana | 1 | 157 | 1012 | 172 |
| 426 | University of Leicester | 4 | 157 | 2276 | 1760 |
| 429 | University of Canterbury | 6 | 156 | 883 | 1020 |
| 430 | Bangor University | 3 | 155 | 968 | 700 |
| 431 | University of St Andrews | 9 | 153 | 498 | 853 |
| 432 | National Centre for Biological Sciences | 4 | 152 | 125 | 127 |
| 432 | Grinnell College | 3 | 152 | 652 | 564 |
| 432 | DeLorme | 1 | 152 | 47 | 15 |
| 435 | Williams College | 6 | 151 | 142 | 237 |
| 436 | The Roslin Institute | 1 | 150 | 608 | 288 |
| 436 | PanThera Biopharma | 1 | 150 | 79 | 36 |
| 438 | Swarthmore College | 6 | 148 | 96 | 171 |
| 438 | Georgia Tech-CNRS Laboratory | 2 | 148 | 16 | 19 |
| 440 | Appalachian State University | 2 | 146 | 775 | 278 |
| 440 | Institute for Advanced Study Berlin | 3 | 146 | 20 | 31 |
| 442 | Fram Centre | 3 | 145 | 93 | 90 |
| 443 | University of Central Florida | 3 | 144 | 2301 | 1424 |
| 443 | Universidad Metropolitana | 4 | 144 | 3119 | 1797 |
| 443 | Institute for Chemistry and Biology of the Marine Environment | 7 | 144 | 45 | 95 |
| 443 | Oregon Department of Fish and Wildlife | 1 | 144 | 86 | 41 |
| 447 | American University | 1 | 142 | 4304 | 997 |
| 447 | Instituto Gulbenkian de Ciência | 3 | 142 | 151 | 170 |
| 449 | University of the Balearic Islands | 2 | 141 | 997 | 458 |
| 450 | Humboldt University of Berlin | 5 | 140 | 4551 | 3842 |
| 450 | University of Potsdam | 5 | 140 | 676 | 817 |
| 450 | University of Nottingham | 4 | 140 | 4078 | 3371 |
| 450 | University of Dundee | 1 | 140 | 3334 | 1453 |
| 450 | Smithsonian Migratory Bird Center | 4 | 140 | 31 | 43 |
| 450 | Schoeps | 1 | 140 | 1 | 1 |
| 450 | Region of Southern Denmark | 2 | 140 | 3029 | 1715 |
| 457 | American Museum of Natural History | 8 | 138 | 157 | 262 |
| 457 | University of Winnipeg | 2 | 138 | 486 | 200 |
| 457 | Bühler (Switzerland) | 1 | 138 | 102 | 36 |
| 460 | University of Regensburg | 5 | 136 | 1340 | 1374 |
| 460 | Xishuangbanna Tropical Botanical Garden | 3 | 136 | 215 | 179 |
| 462 | QIMR Berghofer Medical Research Institute | 1 | 135 | 1077 | 541 |
| 462 | Royal Holloway, University of London | 3 | 135 | 942 | 737 |
| 464 | Marine Biological Laboratory | 4 | 134 | 318 | 401 |
| 464 | Memorial University of Newfoundland | 5 | 134 | 1284 | 1271 |
| 464 | Bernice P. Bishop Museum | 3 | 134 | 48 | 32 |
| 467 | University of Akron | 2 | 133 | 1380 | 692 |
| 467 | University of Windsor | 4 | 133 | 939 | 722 |
| 469 | Academia Sinica | 6 | 132 | 1580 | 1617 |
| 470 | University of Chile | 1 | 131 | 5022 | 1751 |
| 470 | University of Burgundy | 4 | 131 | 1137 | 1014 |
| 470 | Masaryk University | 6 | 131 | 924 | 953 |
| 473 | Ishikawa Prefectural University | 2 | 130 | 110 | 64 |
| 473 | Florida Medical Entomology Laboratory | 3 | 130 | 38 | 37 |
| 475 | University of Chester | 1 | 129 | 539 | 121 |
| 475 | Charles Darwin University | 4 | 129 | 417 | 409 |
| 475 | University Centre in Svalbard | 3 | 129 | 112 | 106 |
| 475 | Vickers | 2 | 129 | 54 | 22 |
| 479 | Jagiellonian University | 5 | 128 | 1968 | 1576 |
| 479 | Pacific Lutheran University | 3 | 128 | 53 | 51 |
| 479 | South University | 2 | 128 | 4590 | 2668 |
| 482 | Marshall University | 3 | 127 | 508 | 346 |
| 482 | Georgetown College | 2 | 127 | 8 | 6 |
| 484 | Karlsruhe Institute of Technology | 1 | 125 | 3864 | 1778 |
| 484 | Montana State University | 3 | 125 | 1191 | 954 |
| 484 | Valdosta State University | 3 | 125 | 101 | 44 |
| 484 | Centre Régional d’Innovation et de Transferts Technologiques des Industries du Bois | 4 | 125 | 37 | 48 |
| 488 | University of Göttingen | 3 | 124 | 2910 | 2264 |
| 488 | Bar-Ilan University | 4 | 124 | 1608 | 1325 |
| 488 | Colgate University | 2 | 124 | 362 | 184 |
| 488 | National Polytechnic Institute | 3 | 124 | 1695 | 879 |
| 492 | Akvaplan-niva | 2 | 123 | 75 | 56 |
| 493 | Franklin & Marshall College | 2 | 122 | 275 | 154 |
| 493 | Uni Research | 2 | 122 | 164 | 152 |
| 493 | Metropolitan College | 2 | 122 | 160 | 32 |
| 496 | University of Houston | 4 | 121 | 2370 | 2061 |
| 496 | Duke University Hospital | 1 | 121 | 3602 | 2094 |
| 498 | University of Manitoba | 3 | 119 | 3518 | 2607 |
| 498 | Southern Connecticut State University | 3 | 119 | 91 | 69 |
| 498 | Mathematical Biosciences Institute | 2 | 119 | 34 | 36 |
| 501 | NOAA | 5 | 118 | 345 | 428 |
| 501 | University of Bath | 5 | 118 | 1593 | 1895 |
| 501 | Australian Museum | 4 | 118 | 211 | 190 |
| 504 | Aberystwyth University | 2 | 117 | 977 | 587 |
| 504 | Sorbonne University | 5 | 117 | 1705 | 1341 |
| 506 | Reed College | 3 | 115 | 132 | 152 |
| 506 | Environment and Climate Change Canada | 2 | 115 | 458 | 401 |
| 508 | Heriot-Watt University | 3 | 114 | 1345 | 1024 |
| 508 | Morrison Institute for Public Policy | 2 | 114 | 6 | 6 |
| 508 | University of Wales, Newport | 3 | 114 | 78 | 63 |
| 511 | Universidad de la República | 4 | 113 | 760 | 563 |
| 512 | Pierre and Marie Curie University | 3 | 112 | 1749 | 1633 |
| 512 | University of Hull | 3 | 112 | 1488 | 1141 |
| 512 | Applied Biomathematics | 2 | 112 | 13 | 23 |
| 512 | Deep Springs College | 1 | 112 | 3 | 6 |
| 516 | Universidad de Antioquia | 2 | 111 | 1529 | 581 |
| 517 | DePauw University | 1 | 110 | 187 | 46 |
| 518 | University of Oviedo | 4 | 109 | 1455 | 1256 |
| 519 | University of Tasmania | 7 | 108 | 1053 | 1694 |
| 519 | Helsinki University of Technology | 2 | 108 | 858 | 698 |
| 519 | New College of Florida | 1 | 108 | 122 | 36 |
| 519 | University of Puerto Rico | 4 | 108 | 891 | 913 |
| 523 | University of Tromsø – The Arctic University of Norway | 3 | 107 | 1539 | 1335 |
| 523 | Ghent University | 6 | 107 | 3285 | 3860 |
| 525 | Konrad Lorenz Institute for Evolution and Cognition Research | 1 | 106 | 81 | 56 |
| 525 | Bradley University | 1 | 106 | 436 | 137 |
| 525 | Southern Federal University | 1 | 106 | 1244 | 296 |
| 525 | Laurentian University | 3 | 106 | 470 | 395 |
| 529 | University of Southern Maine | 2 | 105 | 204 | 137 |
| 529 | Norwegian Institute of Marine Research | 3 | 105 | 290 | 304 |
| 529 | Toho University | 4 | 105 | 884 | 685 |
| 529 | Conservation International | 1 | 105 | 213 | 146 |
| 529 | International Union for Conservation of Nature | 1 | 105 | 136 | 81 |
| 534 | Westfield College | 1 | 104 | 819 | 378 |
| 535 | RWTH Aachen University | 1 | 103 | 4561 | 2214 |
| 535 | Furman University | 1 | 103 | 367 | 115 |
| 535 | Shirshov Institute of Oceanology | 1 | 103 | 413 | 163 |
| 538 | Bryn Mawr College | 2 | 102 | 327 | 230 |
| 538 | Graduate University for Advanced Studies | 5 | 102 | 308 | 403 |
| 538 | Zoological Society of San Diego | 2 | 102 | 84 | 84 |
| 541 | University of Arkansas at Little Rock | 1 | 101 | 648 | 212 |
| 541 | Universidade Federal do Paraná | 2 | 101 | 3055 | 1088 |
| 543 | University of the Witwatersrand | 1 | 100 | 5007 | 1967 |
| 544 | GEOMAR Helmholtz Centre for Ocean Research Kiel | 4 | 99 | 243 | 308 |
| 544 | University of Vigo | 2 | 99 | 1371 | 831 |
| 544 | University of North Carolina Wilmington | 2 | 99 | 608 | 377 |
| 547 | Bard College | 1 | 98 | 276 | 91 |
| 547 | Pace University | 1 | 98 | 650 | 216 |
| 547 | Senckenberg Biodiversity and Climate Research Centre | 1 | 98 | 164 | 101 |
| 547 | University of Rhode Island | 5 | 98 | 842 | 1264 |
| 547 | The Graduate University for Advanced Studies | 4 | 98 | 273 | 278 |
| 552 | Agroscope | 1 | 97 | 555 | 337 |
| 552 | National University of Colombia | 1 | 97 | 3241 | 837 |
| 554 | Michigan Technological University | 1 | 96 | 1784 | 816 |
| 554 | University of Hamburg | 3 | 96 | 1868 | 1806 |
| 554 | Kaohsiung Medical University | 1 | 96 | 2106 | 976 |
| 554 | Osnabrück University | 1 | 96 | 1010 | 482 |
| 554 | University of Colombo | 1 | 96 | 872 | 243 |
| 554 | Connecticut College | 1 | 96 | 331 | 111 |
| 554 | The Earth Institute | 1 | 96 | 435 | 297 |
| 554 | National Ecological Observatory Network | 1 | 96 | 67 | 40 |
| 554 | Field Ornithology Group of Sri Lanka | 1 | 96 | 1 | 2 |
| 554 | Nature Conservation Foundation | 1 | 96 | 84 | 39 |
| 564 | Environmental Change Institute | 1 | 95 | 324 | 223 |
| 564 | Museum für Naturkunde | 3 | 95 | 174 | 165 |
| 566 | South China Botanical Garden, Chinese Academy of Sciences | 2 | 94 | 324 | 277 |
| 567 | University of Oldenburg | 6 | 93 | 460 | 799 |
| 567 | Universidad Autónoma del Estado de Hidalgo | 2 | 93 | 506 | 163 |
| 567 | Babcock University | 1 | 93 | 576 | 223 |
| 570 | Max Planck Institute for Chemical Ecology | 3 | 92 | 138 | 186 |
| 570 | Griffith University | 1 | 92 | 5367 | 2708 |
| 570 | Karlstad University | 2 | 92 | 548 | 365 |
| 573 | University of Hong Kong | 3 | 91 | 7113 | 5568 |
| 573 | John Innes Centre | 3 | 91 | 310 | 390 |
| 573 | National and Kapodistrian University of Athens | 1 | 91 | 7793 | 3903 |
| 573 | Naturalis Biodiversity Center | 1 | 91 | 409 | 215 |
| 573 | University of Rennes 1 | 3 | 91 | 1394 | 1313 |
| 573 | Comcast | 1 | 91 | 301 | 72 |
| 573 | University of Washington Friday Harbor Laboratories | 2 | 91 | 93 | 92 |
| 580 | Department of Earth Sciences | 4 | 90 | 1210 | 1219 |
| 581 | Seoul National University | 2 | 89 | 5965 | 4070 |
| 581 | University of Peradeniya | 2 | 89 | 640 | 386 |
| 581 | Department of Public Health | 1 | 89 | 7606 | 3811 |
| 584 | Minneapolis Community and Technical College | 1 | 88 | 4 | 2 |
| 585 | College of William & Mary | 3 | 86 | 916 | 884 |
| 585 | Museu Paraense Emílio Goeldi | 1 | 86 | 287 | 114 |
| 587 | United States Fish and Wildlife Service | 2 | 85 | 398 | 300 |
| 588 | Goethe University Frankfurt | 1 | 84 | 4411 | 2286 |
| 588 | University of Central Arkansas | 2 | 84 | 219 | 110 |
| 590 | University of Bonn | 2 | 83 | 3899 | 2912 |
| 590 | Wisconsin Department of Natural Resources | 1 | 83 | 147 | 98 |
| 590 | Bamfield Marine Sciences Centre | 2 | 83 | 34 | 35 |
| 590 | Federal University of Uberlândia | 2 | 83 | 1676 | 551 |
| 594 | California Polytechnic State University | 1 | 82 | 1101 | 436 |
| 594 | Trinity University | 1 | 82 | 641 | 236 |
| 594 | Hamilton College | 1 | 82 | 385 | 170 |
| 597 | Evogene | 1 | 81 | 26 | 15 |
| 597 | Shandong Airlines | 2 | 81 | 3373 | 2204 |
| 597 | South Florida Water Management District | 1 | 81 | 101 | 79 |
| 600 | Bielefeld University | 5 | 80 | 704 | 1177 |
| 600 | South Dakota State University | 2 | 80 | 834 | 674 |
| 602 | University of Niš | 1 | 79 | 892 | 254 |
| 602 | University of Porto | 1 | 79 | 4285 | 2098 |
| 602 | Gettysburg College | 1 | 79 | 347 | 143 |
| 602 | Emmanuel College | 3 | 79 | 62 | 67 |
| 606 | Free University | 1 | 78 | 3415 | 2025 |
| 606 | Auburn University at Montgomery | 2 | 78 | 120 | 84 |
| 606 | Université du Québec à Montréal | 6 | 78 | 682 | 1289 |
| 606 | Goodwin College | 1 | 78 | 811 | 552 |
| 610 | University of Toulouse | 5 | 77 | 1760 | 2282 |
| 610 | Beijing Normal University | 5 | 77 | 1600 | 1858 |
| 610 | University at Albany | 1 | 77 | 2263 | 1273 |
| 613 | Hangzhou Normal University | 1 | 76 | 1641 | 758 |
| 613 | Institute for Marine and Antarctic Studies | 4 | 76 | 160 | 213 |
| 615 | University of Indonesia | 1 | 75 | 4428 | 680 |
| 615 | Bandung Institute of Technology | 1 | 75 | 1953 | 448 |
| 615 | IFREMER | 3 | 75 | 573 | 653 |
| 615 | Taiwan Forestry Research Institute | 1 | 75 | 131 | 69 |
| 619 | Kansas State University | 3 | 74 | 2091 | 2033 |
| 619 | Sapienza University of Rome | 1 | 74 | 8161 | 4673 |
| 621 | Hamline University | 1 | 73 | 148 | 63 |
| 621 | Federal University of Rio Grande do Norte | 1 | 73 | 3121 | 957 |
| 623 | Atomic Energy Research Establishment | 1 | 72 | 1235 | 761 |
| 623 | Western University | 3 | 72 | 2048 | 1793 |
| 623 | Rhodes College | 1 | 72 | 251 | 105 |
| 623 | Ricardo-AEA | 1 | 72 | 186 | 122 |
| 623 | Marine Scotland | 3 | 72 | 106 | 151 |
| 623 | Hollings Marine Laboratory | 1 | 72 | 104 | 88 |
| 623 | Great Lakes Science Center | 1 | 72 | 99 | 71 |
| 623 | Woodland Trust | 1 | 72 | 13 | 12 |
| 631 | Ulster University | 2 | 71 | 1973 | 1612 |
| 631 | Virginia Institute of Marine Science | 2 | 71 | 243 | 235 |
| 633 | University of Colorado Boulder | 5 | 70 | 1159 | 2075 |
| 633 | Kumamoto University | 1 | 70 | 2968 | 1545 |
| 633 | Universidad Nacional del Comahue | 3 | 70 | 279 | 283 |
| 633 | National Scientific and Technical Research Council | 3 | 70 | 4660 | 3211 |
| 637 | University of Alaska Fairbanks | 4 | 69 | 435 | 643 |
| 637 | Virginia Commonwealth University | 1 | 69 | 5112 | 3027 |
| 637 | National Oceanography Centre | 2 | 69 | 326 | 349 |
| 637 | University of the Pacific | 1 | 69 | 730 | 372 |
| 641 | Fisheries and Oceans Canada | 4 | 68 | 332 | 445 |
| 642 | Wayne State University | 3 | 67 | 3623 | 3542 |
| 642 | University of Denver | 1 | 67 | 2221 | 1044 |
| 642 | Queensland University of Technology | 2 | 67 | 3561 | 3047 |
| 642 | Australian Research Council | 4 | 67 | 277 | 568 |
| 642 | Hartpury College | 1 | 67 | 57 | 19 |
| 642 | Schneider Children's Medical Center of Israel | 1 | 67 | 1353 | 954 |
| 642 | Västerbotten County | 1 | 67 | 4 | 4 |
| 642 | University of the West of England | 1 | 67 | 1990 | 1046 |
| 650 | Polish Academy of Sciences | 3 | 66 | 3521 | 3129 |
| 650 | University of Castile-La Mancha | 2 | 66 | 1001 | 711 |
| 650 | University of Kent | 1 | 66 | 3149 | 1719 |
| 650 | Vassar College | 1 | 66 | 665 | 379 |
| 650 | Universidade Federal de Goiás | 4 | 66 | 1229 | 891 |
| 655 | University of Salzburg | 1 | 65 | 1828 | 1023 |
| 655 | Austral University of Chile | 3 | 65 | 634 | 608 |
| 655 | Ecolab | 2 | 65 | 154 | 196 |
| 655 | Rekola | 1 | 65 | 2 | 3 |
| 659 | University of Pannonia | 2 | 64 | 465 | 375 |
| 659 | University of Parma | 2 | 64 | 2212 | 1926 |
| 659 | China West Normal University | 1 | 64 | 687 | 314 |
| 659 | Moredun Research Institute | 2 | 64 | 169 | 185 |
| 659 | list of women's colleges | 2 | 64 | 1874 | 1289 |
| 659 | Senckenberg Research Institute and Natural History Museum Frankfurt/M | 2 | 64 | 41 | 61 |
| 665 | Finnish Environment Institute | 2 | 63 | 291 | 331 |
| 666 | Mississippi State University | 1 | 62 | 3158 | 1737 |
| 666 | Bogor Agricultural University | 1 | 62 | 638 | 275 |
| 668 | Technical University of Munich | 1 | 61 | 2378 | 1550 |
| 668 | TU Dresden | 1 | 61 | 5012 | 3068 |
| 668 | National Institutes of Health | 2 | 61 | 4127 | 3763 |
| 668 | Nicolaus Copernicus University in Toruń | 1 | 61 | 2287 | 1000 |
| 668 | Monterey Bay Aquarium Research Institute | 2 | 61 | 131 | 166 |
| 668 | Wildlife Institute of India | 1 | 61 | 184 | 88 |
| 668 | John E. Fogarty International Center | 2 | 61 | 110 | 172 |
| 675 | Technische Universität Darmstadt | 3 | 60 | 1605 | 1657 |
| 675 | George Washington University | 2 | 60 | 4125 | 3383 |
| 675 | City University of New York | 2 | 60 | 3973 | 3079 |
| 678 | Amherst College | 2 | 59 | 304 | 331 |
| 678 | Ryerson University | 1 | 59 | 2295 | 1297 |
| 678 | Oberlin College | 1 | 59 | 517 | 290 |
| 678 | Brazilian Agricultural Research Corporation | 3 | 59 | 1182 | 1117 |
| 678 | University of Iowa | 3 | 59 | 4427 | 4491 |
| 683 | Gibson | 2 | 58 | 136 | 207 |
| 683 | Hungarian Academy of Sciences | 2 | 58 | 2380 | 2176 |
| 683 | Wake Forest University | 4 | 58 | 1038 | 1743 |
| 683 | Université libre de Bruxelles | 4 | 58 | 2116 | 2798 |
| 683 | University of Haifa | 4 | 58 | 1529 | 2010 |
| 683 | Vancouver Island University | 1 | 58 | 172 | 86 |
| 689 | University of Aveiro | 1 | 57 | 3338 | 1946 |
| 689 | Rothamsted Research | 3 | 57 | 449 | 630 |
| 689 | Fondazione Eni Enrico Mattei | 1 | 57 | 93 | 89 |
| 689 | Oswaldo Cruz Foundation | 1 | 57 | 2321 | 1223 |
| 693 | Tata Institute of Fundamental Research | 3 | 56 | 615 | 626 |
| 693 | Correos | 1 | 56 | 573 | 471 |
| 693 | University of Costa Rica | 4 | 56 | 486 | 709 |
| 693 | Ronin Institute | 1 | 56 | 99 | 53 |
| 697 | Shaanxi Normal University | 1 | 55 | 1795 | 1058 |
| 697 | Norwegian Computing Center | 1 | 55 | 134 | 99 |
| 699 | University of Nebraska–Lincoln | 8 | 54 | 891 | 2453 |
| 699 | Manaaki Whenua – Landcare Research | 4 | 54 | 238 | 413 |
| 699 | Lancaster Environment Centre | 1 | 54 | 582 | 523 |
| 699 | University of Alaska Museum of the North | 1 | 54 | 87 | 65 |
| 699 | Natural Resources Research Institute | 1 | 54 | 102 | 95 |
| 704 | University of Freiburg | 3 | 53 | 2834 | 3209 |
| 704 | Technion – Israel Institute of Technology | 2 | 53 | 3860 | 3440 |
| 704 | King Mongkut's Institute of Technology Ladkrabang | 1 | 53 | 779 | 355 |
| 704 | Xiamen University | 1 | 53 | 4154 | 2578 |
| 704 | University of Ontario Institute of Technology | 1 | 53 | 655 | 422 |
| 704 | Ministry of Natural Resources and Wildlife | 1 | 53 | 45 | 56 |
| 704 | Yantai University | 1 | 53 | 1131 | 659 |
| 704 | CORBIDI | 2 | 53 | 19 | 13 |
| 704 | University of Minnesota Twin Cities | 1 | 53 | 367 | 231 |
| 713 | Valparaiso University | 1 | 52 | 269 | 129 |
| 713 | Sustainability Institute | 3 | 52 | 224 | 406 |
| 715 | Nanjing University | 1 | 51 | 5828 | 3917 |
| 715 | University of the West Indies | 2 | 51 | 1521 | 1158 |
| 715 | Towson University | 3 | 51 | 403 | 601 |
| 715 | University of Notre Dame Australia | 3 | 51 | 2750 | 3123 |
| 719 | University of Warsaw | 2 | 50 | 3119 | 2143 |
| 719 | Austrian Academy of Sciences | 1 | 50 | 1378 | 1059 |
| 719 | Estonian University of Life Sciences | 1 | 50 | 503 | 379 |
| 719 | Queens College | 1 | 50 | 1433 | 881 |
| 719 | Natura Cosméticos S.A. | 1 | 50 | 282 | 205 |
| 724 | Trinity College Dublin | 2 | 49 | 2810 | 2736 |
| 724 | Fudan University | 3 | 49 | 4337 | 4221 |
| 724 | University of Turin | 2 | 49 | 4075 | 3638 |
| 724 | California Academy of Sciences | 1 | 49 | 284 | 200 |
| 724 | Institut National de Recherche en Informatique et en Automatique | 1 | 49 | 71 | 81 |
| 724 | University of Plymouth | 2 | 49 | 1826 | 1904 |
| 724 | Hartwick College | 2 | 49 | 40 | 56 |
| 724 | Contraception Research and Development | 1 | 49 | 370 | 310 |
| 732 | University of New England | 3 | 48 | 1308 | 1820 |
| 732 | Tulane University | 4 | 48 | 2076 | 3023 |
| 732 | Edith Cowan University | 2 | 48 | 1323 | 1533 |
| 732 | Thomas Jefferson University | 1 | 48 | 4608 | 3160 |
| 732 | Archbold Biological Station | 1 | 48 | 97 | 76 |
| 732 | Student Conservation Association | 1 | 48 | 2 | 2 |
| 732 | Albertina | 1 | 48 | 198 | 172 |
| 732 | Museu de Ciências Naturais da Fundação Zoobotânica do Rio Grande do Sul | 2 | 48 | 132 | 143 |
| 740 | University of Extremadura | 1 | 47 | 2406 | 1381 |
| 740 | The Rowett Institute | 1 | 47 | 466 | 488 |
| 742 | University of Maryland Center for Environmental Science | 1 | 46 | 316 | 288 |
| 742 | Great Lakes Institute of Management | 2 | 46 | 486 | 621 |
| 744 | University of Bremen | 2 | 45 | 2210 | 2164 |
| 744 | Tokyo Gakugei University | 1 | 45 | 496 | 231 |
| 744 | Zurich University of Applied Sciences/ZHAW | 2 | 45 | 345 | 471 |
| 744 | seznam.cz | 2 | 45 | 29 | 33 |
| 748 | Boston College | 1 | 44 | 2669 | 1922 |
| 748 | Harman International Industries | 1 | 44 | 53 | 30 |
| 748 | Marine Institute of Memorial University of Newfoundland | 1 | 44 | 428 | 329 |
| 751 | Otsuka Pharmaceutical | 3 | 43 | 490 | 757 |
| 751 | Aalborg University | 1 | 43 | 3617 | 2443 |
| 751 | Radboud University Nijmegen | 2 | 43 | 4333 | 4292 |
| 751 | Yokohama National University | 1 | 43 | 1588 | 941 |
| 751 | Royal University | 2 | 43 | 246 | 264 |
| 751 | Regional Government of Castile-La Mancha | 1 | 43 | 28 | 28 |
| 757 | University of Northern Iowa | 1 | 42 | 845 | 554 |
| 757 | Great Basin College | 1 | 42 | 1 | 2 |
| 757 | Pasteur Institute of Montevideo | 1 | 42 | 146 | 143 |
| 757 | University of Lynchburg | 1 | 42 | 99 | 70 |
| 761 | Regis University | 1 | 41 | 172 | 139 |
| 761 | University of Essex | 1 | 41 | 2687 | 1918 |
| 761 | Paris Descartes University | 1 | 41 | 3149 | 2570 |
| 761 | Missouri State University | 1 | 41 | 1557 | 931 |
| 765 | Bowdoin College | 3 | 40 | 167 | 369 |
| 765 | University of Colorado Denver | 1 | 40 | 3037 | 2563 |
| 765 | La Jolla Institute for Allergy and Immunology | 2 | 40 | 151 | 269 |
| 765 | Western Center | 1 | 40 | 47 | 48 |
| 765 | UMR BIOdiversity, GEnes & Communities | 3 | 40 | 91 | 176 |
| 770 | Fordham University | 1 | 39 | 1818 | 1203 |
| 770 | University at Buffalo | 1 | 39 | 6274 | 4589 |
| 770 | University of Portsmouth | 1 | 39 | 2540 | 1912 |
| 770 | Joint Institute for Marine and Atmospheric Research | 2 | 39 | 43 | 81 |
| 770 | Mathematical Institute, University of Oxford | 1 | 39 | 1505 | 1082 |
| 770 | Toulouse III University - Paul-Sabatier | 3 | 39 | 303 | 729 |
| 776 | Texas State University | 2 | 38 | 1553 | 1570 |
| 776 | Hólar University College | 2 | 38 | 22 | 30 |
| 776 | University of Strathclyde | 1 | 38 | 4319 | 3253 |
| 776 | Population Research Institute | 1 | 38 | 132 | 152 |
| 776 | Universidad de Santiago de Chile | 1 | 38 | 1382 | 853 |
| 781 | Stetson University | 1 | 37 | 329 | 219 |
| 781 | John Radcliffe Hospital | 1 | 37 | 1889 | 1706 |
| 781 | interdisciplinarity | 2 | 37 | 8350 | 7189 |
| 781 | Carl Von Ossietzky Universität Oldenburg | 1 | 37 | 453 | 265 |
| 785 | University of Jaén | 1 | 36 | 1592 | 1098 |
| 785 | University of Warwick | 3 | 36 | 3355 | 4359 |
| 785 | University of Lisbon | 3 | 36 | 4011 | 4536 |
| 785 | University College Cork | 3 | 36 | 1625 | 2551 |
| 785 | University of Canberra | 1 | 36 | 1618 | 1309 |
| 785 | Tarleton State University | 1 | 36 | 249 | 161 |
| 785 | Te Pūnaha Matatini | 1 | 36 | 35 | 35 |
| 785 | National Heart, Lung, and Blood Institute | 1 | 36 | 548 | 491 |
| 785 | Oeko Institut | 1 | 36 | 235 | 248 |
| 794 | Florida Atlantic University | 2 | 35 | 1858 | 2017 |
| 794 | Universidad de Sonora | 1 | 35 | 920 | 481 |
| 794 | University of New Brunswick | 1 | 35 | 2313 | 1758 |
| 794 | Atlantic University | 2 | 35 | 1900 | 2047 |
| 794 | Universidade Federal de Sergipe | 2 | 35 | 1361 | 847 |
| 794 | Nicholls State University | 1 | 35 | 151 | 121 |
| 794 | Skidaway Institute of Oceanography | 2 | 35 | 82 | 141 |
| 794 | Agroecology | 2 | 35 | 836 | 907 |
| 802 | University of Ljubljana | 1 | 34 | 4648 | 3349 |
| 802 | Alfred Wegener Institute for Polar and Marine Research | 1 | 34 | 625 | 614 |
| 802 | University of Hohenheim | 2 | 34 | 1166 | 1492 |
| 802 | Aix-Marseille University | 1 | 34 | 4602 | 3468 |
| 802 | French Institute of Pondicherry | 1 | 34 | 52 | 61 |
| 802 | Novia University of Applied Sciences | 2 | 34 | 32 | 54 |
| 808 | University of San Diego | 1 | 33 | 1035 | 844 |
| 808 | Ruhr University Bochum | 2 | 33 | 3649 | 3725 |
| 808 | China University of Geosciences | 1 | 33 | 2127 | 1634 |
| 808 | University of Wollongong | 2 | 33 | 2972 | 3416 |
| 808 | South Australian Research and Development Institute | 3 | 33 | 108 | 275 |
| 808 | CSIRO Oceans and Atmosphere | 2 | 33 | 235 | 378 |
| 808 | Edmund Mach Foundation | 1 | 33 | 406 | 449 |
| 815 | Kanagawa University | 1 | 32 | 740 | 428 |
| 815 | Université du Québec | 1 | 32 | 1556 | 1464 |
| 815 | Leibniz University Hannover | 1 | 32 | 1817 | 1537 |
| 815 | Keio University | 1 | 32 | 4806 | 3446 |
| 815 | University of Valle | 1 | 32 | 1532 | 866 |
| 815 | Anschutz Medical Campus | 1 | 32 | 2373 | 2157 |
| 815 | Arcadia University | 1 | 32 | 203 | 168 |
| 815 | Universidad Icesi | 1 | 32 | 378 | 190 |
| 823 | University of Missouri–St. Louis | 2 | 31 | 55 | 184 |
| 823 | Indian Institutes of Science Education and Research | 2 | 31 | 688 | 798 |
| 823 | Universidade Federal de Mato Grosso do Sul | 1 | 31 | 1692 | 711 |
| 826 | Sun Yat-sen University | 1 | 30 | 7089 | 5501 |
| 826 | Federal University of Rio de Janeiro | 2 | 30 | 5387 | 4398 |
| 826 | Vrije Universiteit Brussel | 1 | 30 | 4039 | 3372 |
| 826 | Providence College | 1 | 30 | 345 | 253 |
| 826 | University of Zadar | 1 | 30 | 260 | 113 |
| 826 | Prairie Research Institute | 2 | 30 | 140 | 224 |
| 826 | Forest Research | 1 | 30 | 774 | 709 |
| 833 | Tokyo Institute of Technology | 1 | 29 | 3730 | 2912 |
| 833 | University of Memphis | 2 | 29 | 1819 | 2350 |
| 833 | Macdonald Campus | 1 | 29 | 575 | 600 |
| 833 | University of Puerto Rico, Río Piedras Campus | 1 | 29 | 48 | 54 |
| 837 | James Madison University | 1 | 28 | 1192 | 956 |
| 837 | Genoscope | 1 | 28 | 167 | 217 |
| 837 | Fujita Health University | 1 | 28 | 1409 | 1122 |
| 837 | Louisiana State University Agricultural Center | 1 | 28 | 553 | 553 |
| 837 | Madison University | 1 | 28 | 1207 | 968 |
| 837 | Mary Baldwin University | 1 | 28 | 27 | 36 |
| 837 | Stonehill College | 1 | 28 | 89 | 88 |
| 837 | University of the French Antilles | 3 | 28 | 229 | 445 |
| 837 | Lincoln Park Zoo | 2 | 28 | 48 | 93 |
| 837 | Queensland Government | 1 | 28 | 131 | 179 |
| 847 | University of Bordeaux | 2 | 27 | 2893 | 3281 |
| 847 | National Taiwan University | 2 | 27 | 5291 | 5469 |
| 847 | Murray State University | 2 | 27 | 236 | 404 |
| 847 | University of the Sunshine Coast | 2 | 27 | 862 | 1269 |
| 847 | Universidade Federal de São Carlos | 1 | 27 | 3331 | 2022 |
| 847 | University of Veterinary Medicine Hanover, Foundation | 1 | 27 | 544 | 564 |
| 853 | Bennington College | 1 | 26 | 56 | 57 |
| 853 | University of San Francisco | 1 | 26 | 957 | 948 |
| 853 | New Jersey Institute of Technology | 2 | 26 | 1118 | 1504 |
| 853 | Sweet Briar College | 1 | 26 | 49 | 65 |
| 853 | Computing Research Association | 1 | 26 | 3 | 10 |
| 853 | Centre for Life | 1 | 26 | 1569 | 1597 |
| 859 | Fraunhofer Society | 1 | 25 | 2007 | 1903 |
| 859 | University of Giessen | 1 | 25 | 3775 | 3397 |
| 859 | University of Alcalá | 1 | 25 | 2456 | 2000 |
| 859 | Biotechnology Institute | 2 | 25 | 1142 | 1639 |
| 859 | State Museum of Natural History Karlsruhe | 1 | 25 | 40 | 47 |
| 859 | Department of Archaeology | 2 | 25 | 532 | 659 |
| 865 | Leipzig University | 1 | 24 | 4534 | 3971 |
| 865 | Complutense University of Madrid | 1 | 24 | 6958 | 5110 |
| 865 | Norwegian University of Science and Technology | 2 | 24 | 1493 | 2234 |
| 865 | Cardiff University | 1 | 24 | 5848 | 5250 |
| 865 | Forest Research Institute | 1 | 24 | 1102 | 1068 |
| 865 | Stephens College | 2 | 24 | 227 | 554 |
| 865 | ComUE Paris-Saclay University | 2 | 24 | 2982 | 3154 |
| 865 | Department of Primary Industries | 2 | 24 | 621 | 899 |
| 865 | Museum of New Zealand Te Papa Tongarewa | 1 | 24 | 116 | 148 |
| 874 | University of Iceland | 3 | 23 | 1090 | 2199 |
| 874 | Seattle University | 1 | 23 | 578 | 623 |
| 874 | Mount Allison University | 2 | 23 | 258 | 475 |
| 874 | University of Florence | 1 | 23 | 5158 | 4550 |
| 874 | Consiglio Nazionale delle Ricerche | 1 | 23 | 2803 | 2774 |
| 874 | Sichuan University | 1 | 23 | 5750 | 4878 |
| 874 | University of Lille | 3 | 23 | 1812 | 3173 |
| 874 | Dominican University | 1 | 23 | 122 | 115 |
| 874 | Millersville University of Pennsylvania | 1 | 23 | 211 | 189 |
| 874 | Natural History Museum of Los Angeles County | 1 | 23 | 176 | 173 |
| 874 | onet.pl | 1 | 23 | 57 | 55 |
| 874 | Antiquities, Monuments and Museum Corporation | 1 | 23 | 7 | 10 |
| 874 | Poznań University of Life Sciences | 1 | 23 | 471 | 448 |
| 887 | University of Toyama | 1 | 22 | 2363 | 2021 |
| 887 | St. Jude Children's Research Hospital | 1 | 22 | 1341 | 1429 |
| 887 | Lanzhou University | 3 | 22 | 2160 | 3046 |
| 887 | Adelphi University | 1 | 22 | 698 | 648 |
| 887 | Center for Philosophy of Science | 1 | 22 | 40 | 52 |
| 887 | Forest Research Institute Malaysia | 1 | 22 | 174 | 201 |
| 887 | Luther University | 2 | 22 | 1076 | 1782 |
| 887 | National Institute for Agro-Environmental Sciences | 1 | 22 | 245 | 292 |
| 887 | Arizona Game and Fish Department | 1 | 22 | 57 | 56 |
| 887 | Australian National Wildlife Collection | 1 | 22 | 47 | 56 |
| 897 | University of Colorado Museum of Natural History | 1 | 21 | 24 | 36 |
| 897 | Duke University West Campus | 2 | 21 | 287 | 781 |
| 899 | Colby College | 2 | 20 | 220 | 489 |
| 899 | Central China Normal University | 2 | 20 | 1107 | 1499 |
| 899 | University of Cumbria | 1 | 20 | 166 | 194 |
| 899 | Chinese Academy of Tropical Agricultural Sciences | 2 | 20 | 308 | 478 |
| 899 | Stockton University | 1 | 20 | 313 | 363 |
| 904 | University of Alaska Southeast | 1 | 19 | 99 | 144 |
| 904 | Worcester Polytechnic Institute | 1 | 19 | 1375 | 1479 |
| 904 | Tennessee State University | 2 | 19 | 1705 | 2576 |
| 904 | Washington Department of Ecology | 1 | 19 | 56 | 87 |
| 904 | Universidade do Vale do Itajaí | 1 | 19 | 495 | 252 |
| 904 | University of Puerto Rico at Mayaguez | 1 | 19 | 1868 | 1272 |
| 904 | Sitka Sound Science Center | 1 | 19 | 4 | 6 |
| 904 | Dauphin Island Sea Lab | 1 | 19 | 113 | 134 |
| 912 | Meiji University | 1 | 18 | 1137 | 912 |
| 912 | Smith College | 1 | 18 | 892 | 988 |
| 912 | University of Paris | 1 | 18 | 4164 | 4134 |
| 912 | University of Caen Normandy | 1 | 18 | 1742 | 1893 |
| 912 | Irkutsk State University | 1 | 18 | 450 | 308 |
| 912 | Saint Louis University | 1 | 18 | 3157 | 3219 |
| 912 | University of South Dakota | 1 | 18 | 1334 | 1462 |
| 912 | Nanjing Normal University | 2 | 18 | 1670 | 2161 |
| 912 | Santos | 1 | 18 | 2233 | 1957 |
| 912 | Sorbonne Universités | 1 | 18 | 2057 | 2384 |
| 912 | Norwich Research Park | 1 | 18 | 1053 | 1395 |
| 912 | Theodore Roosevelt High School | 1 | 18 | 3185 | 3300 |
| 912 | Universidad Nacional Mayor de San Marcos | 1 | 18 | 1048 | 588 |
| 912 | California State University Channel Islands | 1 | 18 | 94 | 139 |
| 926 | University of Pisa | 1 | 17 | 5425 | 5182 |
| 926 | Temple University | 2 | 17 | 3861 | 5109 |
| 926 | University of L'Aquila | 1 | 17 | 2218 | 2393 |
| 926 | Unity College | 1 | 17 | 21 | 34 |
| 926 | Université de Perpignan | 1 | 17 | 513 | 617 |
| 926 | Universidade Federal do Sul da Bahia | 1 | 17 | 230 | 87 |
| 932 | University of Marburg | 1 | 16 | 2603 | 2808 |
| 932 | AgResearch | 1 | 16 | 769 | 965 |
| 932 | Chalmers University of Technology | 1 | 16 | 3491 | 3727 |
| 932 | Grenoble University - Grenoble Alpes University | 1 | 16 | 1817 | 1848 |
| 932 | Old Dominion University | 1 | 16 | 2557 | 2731 |
| 932 | Swedish Meteorological and Hydrological Institute | 1 | 16 | 199 | 248 |
| 932 | Denver Museum of Nature and Science | 1 | 16 | 308 | 295 |
| 932 | University of Wisconsin–Eau Claire | 1 | 16 | 703 | 824 |
| 932 | European University of Brittany | 1 | 16 | 468 | 701 |
| 932 | Department of Public | 1 | 16 | 3167 | 2812 |
| 932 | Agrocampus Ouest | 1 | 16 | 373 | 537 |
| 932 | École Nationale Supérieure des Sciences Agronomiques de Bordeaux-Aquitaine | 1 | 16 | 174 | 254 |
| 932 | Institute for Condensed Matter Physics | 2 | 16 | 90 | 150 |
| 945 | Linköping University | 1 | 15 | 4358 | 4665 |
| 945 | Max Planck Institute for Developmental Biology | 1 | 15 | 164 | 312 |
| 945 | Capital University | 1 | 15 | 597 | 765 |
| 945 | Lock Haven University of Pennsylvania | 1 | 15 | 64 | 89 |
| 945 | California State University, Los Angeles | 1 | 15 | 285 | 407 |
| 945 | Institut national d'études supérieures agronomiques de Montpellier | 1 | 15 | 474 | 663 |
| 945 | Science Museum | 1 | 15 | 292 | 346 |
| 945 | City College | 1 | 15 | 1712 | 1811 |
| 953 | London School of Economics and Political Science | 1 | 14 | 2728 | 2769 |
| 953 | Aalto University | 1 | 14 | 2703 | 2983 |
| 953 | Rhodes University | 1 | 14 | 1331 | 1450 |
| 953 | University of Sassari | 1 | 14 | 1791 | 2218 |
| 953 | Yunnan University | 1 | 14 | 2163 | 2190 |
| 953 | Max Planck Institute for Human Development | 1 | 14 | 365 | 586 |
| 953 | College of Wooster | 1 | 14 | 320 | 398 |
| 953 | Murdoch University | 1 | 14 | 2601 | 3131 |
| 953 | University of Namur | 1 | 14 | 1093 | 1404 |
| 953 | University of Western Brittany | 1 | 14 | 1240 | 1510 |
| 953 | University of Ghana | 1 | 14 | 2298 | 2334 |
| 953 | Australian Institute of Marine Science | 1 | 14 | 313 | 445 |
| 953 | Helsinki Institute for Information Technology | 1 | 14 | 33 | 81 |
| 953 | Armed Forces Research Institute of Medical Sciences | 1 | 14 | 165 | 227 |
| 953 | Noguchi Memorial Institute for Medical Research | 1 | 14 | 235 | 327 |
| 953 | Menzies Institute for Medical Research | 1 | 14 | 617 | 848 |
| 953 | United States Army | 1 | 14 | 1971 | 2429 |
| 953 | Cleveland Museum of Natural History | 1 | 14 | 56 | 84 |
| 953 | Natural History Museum of Utah | 1 | 14 | 59 | 94 |
| 972 | Laboratoire Jacques-Louis Lions | 1 | 13 | 235 | 259 |
| 972 | Koninklijke Philips NV | 1 | 13 | 1831 | 2149 |
| 972 | University of California, Riverside | 1 | 13 | 1712 | 2228 |
| 972 | Sorbonne | 1 | 13 | 4213 | 4089 |
| 972 | Scottish Parliament | 1 | 13 | 73 | 123 |
| 977 | Pompeu Fabra University | 1 | 12 | 2522 | 2835 |
| 977 | Université catholique de Louvain | 2 | 12 | 3835 | 5542 |
| 977 | Autonomous University of Madrid | 1 | 12 | 4587 | 4577 |
| 977 | Clermont Auvergne University | 1 | 12 | 1544 | 1730 |
| 977 | Università degli Studi Roma Tre | 2 | 12 | 614 | 1241 |
| 977 | National Polytechnic Institute of Toulouse | 1 | 12 | 236 | 340 |
| 977 | Higher University of San Andrés | 1 | 12 | 241 | 335 |
| 977 | Hospital de Clínicas de Porto Alegre | 1 | 12 | 1062 | 1261 |
| 977 | University of French Guiana | 1 | 12 | 143 | 205 |
| 977 | Fairchild Tropical Botanic Garden | 1 | 12 | 79 | 118 |
| 977 | Tall Timbers Research Station and Land Conservancy | 1 | 12 | 59 | 94 |
| 977 | Ecology of Guianan Forests | 1 | 12 | 104 | 176 |
| 977 | Perot Museum of Nature and Science | 1 | 12 | 53 | 88 |
| 990 | Technical University of Madrid | 1 | 11 | 2955 | 3080 |
| 990 | University of Kiel | 2 | 11 | 1860 | 3307 |
| 990 | Loyola University | 2 | 11 | 2541 | 3916 |
| 990 | California State University San Marcos | 1 | 11 | 248 | 470 |
| 990 | Vilnius University | 1 | 11 | 2191 | 2376 |
| 990 | Assumption University | 1 | 11 | 125 | 155 |
| 990 | Xi'an Jiaotong University | 1 | 11 | 4929 | 5077 |
| 990 | Charles Sturt University | 1 | 11 | 1911 | 2444 |
| 990 | Long Island University | 1 | 11 | 854 | 1192 |
| 990 | Universidade Federal do Amazonas | 1 | 11 | 1463 | 1144 |
| 990 | Kyoto University of Advanced Science | 1 | 11 | 146 | 162 |