| # | Institution | Articles | Citations | R ARank | R CRank |
|---|
| 1 | University of California | 83 | 4,112 | 1.8K | 1.6K |
| 2 | Lund University | 49 | 3,403 | 376 | 244 |
| 3 | University of Northampton | 3 | 2,470 | 138 | 1 |
| 4 | Swedish University of Agricultural Sciences | 53 | 2,428 | 105 | 104 |
| 5 | National Center for Scientific Research | 102 | 2,382 | 593 | 782 |
| 6 | University of British Columbia | 38 | 2,303 | 909 | 675 |
| 7 | Umeå University | 39 | 2,265 | 170 | 141 |
| 8 | University of California, Berkeley | 13 | 2,215 | 771 | 294 |
| 9 | Imperial College London | 28 | 2,157 | 1.1K | 642 |
| 10 | Wageningen University & Research | 38 | 2,042 | 278 | 280 |
| 11 | Purdue University | 12 | 1,894 | 1.9K | 476 |
| 12 | University of Alberta | 26 | 1,771 | 1.1K | 587 |
| 13 | Stockholm University | 49 | 1,703 | 151 | 242 |
| 14 | Uppsala University | 36 | 1,702 | 491 | 454 |
| 15 | Aarhus University | 28 | 1,634 | 521 | 364 |
| 16 | University of Florida | 24 | 1,510 | 1.4K | 806 |
| 17 | University of Oslo | 32 | 1,502 | 425 | 392 |
| 18 | University of Arizona | 28 | 1,454 | 708 | 604 |
| 19 | Michigan State University | 14 | 1,443 | 1.6K | 624 |
| 20 | Cornell University | 24 | 1,345 | 1.1K | 926 |
| 21 | University of Zurich | 38 | 1,291 | 348 | 531 |
| 22 | University of Minnesota | 29 | 1,271 | 1.4K | 1.2K |
| 23 | Hendrix College | 1 | 1,244 | 99 | 2 |
| 24 | University of Washington | 19 | 1,242 | 2.2K | 1.5K |
| 25 | University of Helsinki | 34 | 1,227 | 541 | 637 |
| 26 | University of Connecticut | 16 | 1,185 | 740 | 388 |
| 27 | McGill University | 26 | 1,164 | 1.1K | 974 |
| 28 | University of Vermont | 12 | 1,093 | 453 | 214 |
| 29 | University of Tennessee | 17 | 1,088 | 784 | 435 |
| 30 | University of Michigan | 24 | 1,048 | 1.9K | 1.7K |
| 31 | University of Turku | 24 | 1,027 | 293 | 263 |
| 32 | University of Georgia | 15 | 1,012 | 1.2K | 627 |
| 33 | Spanish National Research Council | 21 | 992 | 82 | 130 |
| 34 | University of Oxford | 22 | 968 | 1.6K | 1.4K |
| 35 | Université de Sherbrooke | 6 | 950 | 808 | 151 |
| 36 | Ben-Gurion University of the Negev | 19 | 934 | 373 | 222 |
| 37 | Florida International University | 11 | 920 | 425 | 161 |
| 38 | University of Texas at Austin | 11 | 902 | 3.0K | 1.4K |
| 39 | University of Southampton | 12 | 881 | 1.3K | 667 |
| 40 | Ghent University | 20 | 869 | 884 | 805 |
| 41 | Oregon State University | 13 | 842 | 850 | 518 |
| 42 | Miami University | 12 | 836 | 218 | 98 |
| 43 | Princeton University | 8 | 831 | 1.4K | 754 |
| 44 | University of Sydney | 25 | 827 | 1.3K | 1.3K |
| 44 | Katholieke Universiteit Leuven | 21 | 827 | 904 | 937 |
| 46 | Washington State University | 9 | 805 | 1.2K | 545 |
| 47 | University of Basel | 6 | 804 | 1.2K | 461 |
| 48 | Norwegian University of Life Sciences | 18 | 795 | 101 | 116 |
| 49 | Australian National University | 14 | 784 | 987 | 655 |
| 50 | University of Leeds | 11 | 781 | 1.7K | 795 |
| 51 | University of Göttingen | 10 | 771 | 1.0K | 546 |
| 52 | Rutgers University | 15 | 766 | 1.2K | 947 |
| 53 | University of Melbourne | 14 | 748 | 2.1K | 1.4K |
| 54 | Commonwealth Scientific and Industrial Research Organisation | 7 | 729 | 571 | 239 |
| 55 | University of Bern | 17 | 715 | 776 | 723 |
| 56 | Oklahoma State University | 8 | 714 | 789 | 243 |
| 57 | University of Gothenburg | 8 | 712 | 1.7K | 800 |
| 57 | Chinese Academy of Sciences | 29 | 712 | 2.1K | 1.9K |
| 59 | ETH Zürich | 13 | 711 | 1.1K | 875 |
| 60 | University of Toronto | 25 | 697 | 2.0K | 2.3K |
| 61 | Autonomous University of Barcelona | 5 | 693 | 2.0K | 524 |
| 62 | University of Edinburgh | 8 | 686 | 2.6K | 1.2K |
| 63 | University of Montana | 12 | 655 | 164 | 148 |
| 64 | University of Amsterdam | 18 | 645 | 1.1K | 1.3K |
| 65 | University of Aberdeen | 12 | 626 | 767 | 600 |
| 65 | University of New Mexico | 11 | 626 | 727 | 510 |
| 67 | (former) Institut National de Recherche pour l'Agriculture, l'Alimentation et l'Environnement | 29 | 621 | 293 | 554 |
| 68 | University of Jyväskylä | 15 | 618 | 219 | 194 |
| 69 | University of Potsdam | 15 | 617 | 195 | 171 |
| 70 | University of North Carolina | 4 | 614 | 3.3K | 1.1K |
| 71 | University of Exeter | 12 | 608 | 680 | 492 |
| 72 | University of California, Santa Barbara | 12 | 602 | 283 | 324 |
| 73 | University of California, San Diego | 10 | 600 | 1.5K | 1.2K |
| 74 | University College Cork | 5 | 598 | 816 | 274 |
| 75 | University of Idaho | 6 | 591 | 585 | 218 |
| 76 | Yale University | 15 | 590 | 1.2K | 1.3K |
| 77 | University of Würzburg | 13 | 588 | 700 | 606 |
| 78 | University of California, Davis | 21 | 581 | 372 | 721 |
| 79 | University of Tartu | 13 | 580 | 265 | 184 |
| 80 | University of York | 8 | 579 | 1.1K | 565 |
| 81 | University of Freiburg | 12 | 578 | 696 | 601 |
| 82 | University of Sheffield | 13 | 572 | 1.4K | 1.1K |
| 83 | Rice University | 18 | 569 | 326 | 503 |
| 84 | Macquarie University | 13 | 568 | 532 | 427 |
| 85 | Utrecht University | 11 | 561 | 1.6K | 1.3K |
| 85 | University of Canterbury | 11 | 561 | 378 | 234 |
| 85 | Virginia Commonwealth University | 1 | 561 | 4.7K | 633 |
| 88 | University of Wisconsin–Madison | 20 | 558 | 2.0K | 2.3K |
| 89 | Norwegian University of Science and Technology | 24 | 556 | 314 | 544 |
| 90 | Nicolaus Copernicus University in Toruń | 9 | 549 | 348 | 78 |
| 91 | Kansas State University | 7 | 546 | 981 | 444 |
| 92 | University of Maryland | 13 | 545 | 2.1K | 1.7K |
| 93 | École Normale Supérieure | 8 | 542 | 336 | 234 |
| 94 | University of New Hampshire | 5 | 534 | 697 | 269 |
| 94 | University of Bergen | 11 | 534 | 806 | 655 |
| 96 | University of Groningen | 17 | 527 | 1.0K | 1.2K |
| 97 | University of Utah | 7 | 523 | 2.4K | 1.2K |
| 97 | University of Rennes 1 | 2 | 523 | 1.3K | 189 |
| 99 | Eötvös Loránd University | 7 | 522 | 497 | 157 |
| 100 | Kyoto University | 13 | 513 | 2.1K | 1.5K |
| 101 | Liverpool John Moores University | 5 | 501 | 553 | 159 |
| 102 | University of California, Los Angeles | 7 | 480 | 1.8K | 1.2K |
| 103 | Bucknell University | 4 | 479 | 204 | 31 |
| 104 | University of Massachusetts Boston | 3 | 467 | 412 | 65 |
| 105 | University of Plymouth | 8 | 466 | 384 | 266 |
| 106 | Hebrew University of Jerusalem | 12 | 465 | 946 | 878 |
| 107 | Simon Fraser University | 7 | 463 | 1.1K | 592 |
| 108 | Indiana University | 13 | 460 | 1.1K | 1.1K |
| 109 | Colorado State University | 10 | 453 | 1.1K | 878 |
| 110 | Eawag | 5 | 450 | 96 | 76 |
| 111 | University of Notre Dame | 5 | 424 | 1.4K | 600 |
| 112 | University of New South Wales | 14 | 423 | 1.7K | 1.7K |
| 113 | University of Queensland | 11 | 422 | 2.4K | 2.1K |
| 114 | Monash University | 8 | 419 | 2.7K | 1.7K |
| 115 | University of Pretoria | 7 | 414 | 971 | 361 |
| 115 | North Dakota State University | 3 | 414 | 994 | 240 |
| 117 | Czech Academy of Sciences | 21 | 412 | 458 | 619 |
| 118 | University of Alabama | 3 | 411 | 2.8K | 729 |
| 119 | Idaho State University | 3 | 406 | 347 | 51 |
| 120 | University of St Andrews | 9 | 401 | 444 | 333 |
| 121 | University of Copenhagen | 12 | 397 | 1.9K | 1.8K |
| 122 | University of Jena | 7 | 395 | 830 | 513 |
| 123 | King Juan Carlos University | 13 | 393 | 111 | 115 |
| 124 | Vrije Universiteit Amsterdam | 10 | 391 | 1.0K | 1.1K |
| 125 | Northern Arizona University | 7 | 386 | 276 | 183 |
| 126 | Manaaki Whenua – Landcare Research | 10 | 385 | 72 | 105 |
| 127 | Leiden University | 7 | 377 | 1.3K | 909 |
| 128 | Montana State University | 6 | 371 | 576 | 349 |
| 129 | Arizona State University | 8 | 361 | 2.0K | 1.5K |
| 129 | Iowa State University | 10 | 361 | 1.4K | 1.2K |
| 131 | Hungarian Academy of Sciences | 6 | 358 | 1.1K | 594 |
| 132 | University of Cape Town | 6 | 357 | 1.4K | 778 |
| 133 | Carleton University | 9 | 349 | 619 | 495 |
| 134 | Stanford University | 15 | 346 | 2.0K | 2.8K |
| 134 | California State University | 5 | 346 | 2.0K | 672 |
| 136 | University of Oulu | 13 | 342 | 533 | 672 |
| 137 | Crown Research Institute | 2 | 341 | 13 | 3 |
| 138 | Pennsylvania State University | 10 | 338 | 2.8K | 2.4K |
| 139 | San Francisco State University | 7 | 337 | 215 | 176 |
| 139 | Swansea University | 3 | 337 | 1.6K | 468 |
| 141 | University of North Carolina Wilmington | 3 | 332 | 337 | 76 |
| 142 | University of Cambridge | 12 | 331 | 2.5K | 2.7K |
| 143 | St. Olaf College | 1 | 330 | 238 | 12 |
| 144 | Stellenbosch University | 8 | 324 | 852 | 547 |
| 145 | Deakin University | 10 | 320 | 766 | 732 |
| 146 | University of Guelph | 9 | 319 | 949 | 907 |
| 147 | United States Geological Survey | 10 | 314 | 239 | 296 |
| 148 | University of Maine | 5 | 301 | 630 | 381 |
| 149 | University of Konstanz | 11 | 299 | 360 | 529 |
| 150 | University of Bayreuth | 4 | 296 | 854 | 448 |
| 151 | University of Ottawa | 6 | 295 | 2.3K | 1.5K |
| 151 | Utah State University | 8 | 295 | 585 | 584 |
| 153 | University of Illinois at Urbana–Champaign | 9 | 291 | 1.7K | 1.7K |
| 154 | University of East Anglia | 8 | 282 | 819 | 835 |
| 155 | University of the Witwatersrand | 10 | 279 | 753 | 632 |
| 156 | Technische Universität Darmstadt | 4 | 278 | 1.1K | 525 |
| 157 | University of Calgary | 10 | 275 | 1.9K | 1.9K |
| 158 | University of Massachusetts | 4 | 274 | 2.5K | 1.4K |
| 159 | University of Barcelona | 6 | 270 | 2.4K | 1.5K |
| 160 | University of Concepción | 5 | 265 | 596 | 248 |
| 161 | James Cook University | 11 | 263 | 322 | 480 |
| 162 | University of Pisa | 3 | 261 | 3.2K | 1.3K |
| 162 | Centre d'Écologie Fonctionnelle et Évolutive | 14 | 261 | 14 | 40 |
| 164 | Radboud University Nijmegen | 7 | 253 | 1.3K | 1.3K |
| 164 | University of Colorado | 6 | 253 | 2.2K | 1.8K |
| 166 | Andrews University | 2 | 251 | 71 | 19 |
| 166 | California State University, Los Angeles | 2 | 251 | 149 | 34 |
| 168 | Brown University | 5 | 250 | 2.0K | 1.5K |
| 168 | State University of New York | 2 | 250 | 4.9K | 1.8K |
| 170 | University of Neuchâtel | 11 | 247 | 133 | 253 |
| 171 | McMaster University | 2 | 246 | 5.1K | 2.0K |
| 172 | University of Liverpool | 14 | 243 | 1.1K | 1.7K |
| 172 | University of Oklahoma | 6 | 243 | 1.3K | 959 |
| 174 | Joseph Fourier University | 3 | 241 | 521 | 369 |
| 175 | University of Wyoming | 6 | 240 | 693 | 570 |
| 176 | University of Pennsylvania | 1 | 235 | 8.8K | 3.2K |
| 177 | University of Haifa | 5 | 234 | 967 | 569 |
| 178 | American Museum of Natural History | 2 | 232 | 379 | 158 |
| 178 | Linköping University | 5 | 232 | 1.4K | 1.0K |
| 180 | Charles University | 8 | 226 | 1.3K | 1.1K |
| 181 | Helmholtz Centre for Environmental Research | 5 | 223 | 30 | 50 |
| 182 | University of the Fraser Valley | 1 | 219 | 112 | 6 |
| 183 | Ohio Wesleyan University | 1 | 218 | 154 | 12 |
| 184 | San Diego State University | 4 | 215 | 1.3K | 797 |
| 185 | Louisiana State University | 6 | 211 | 1.8K | 1.5K |
| 186 | Queen's University Belfast | 6 | 210 | 1.2K | 1.0K |
| 187 | Adam Mickiewicz University in Poznań | 5 | 209 | 654 | 287 |
| 187 | University of Tasmania | 8 | 209 | 769 | 851 |
| 187 | Université du Québec à Montréal | 5 | 209 | 693 | 520 |
| 190 | University of Marburg | 8 | 206 | 767 | 977 |
| 190 | University of Lille | 3 | 206 | 507 | 183 |
| 192 | University of Tübingen | 9 | 204 | 1.3K | 1.6K |
| 192 | Penryn Campus | 5 | 204 | 57 | 78 |
| 192 | Swiss Federal Institute for Forest, Snow and Landscape Research | 1 | 204 | 78 | 40 |
| 195 | Wake Forest University | 2 | 202 | 1.4K | 586 |
| 196 | Hokkaido University | 12 | 201 | 1.4K | 1.6K |
| 196 | Hopkins Marine Station | 2 | 201 | 93 | 66 |
| 198 | University of Antwerp | 10 | 197 | 678 | 1.1K |
| 198 | University of Alaska System | 1 | 197 | 7 | 1 |
| 200 | University of KwaZulu-Natal | 6 | 196 | 799 | 533 |
| 200 | National University of Singapore | 5 | 196 | 3.6K | 2.7K |
| 200 | University of New Brunswick | 2 | 196 | 1.3K | 432 |
| 200 | Trent University | 9 | 196 | 139 | 206 |
| 204 | College of William & Mary | 1 | 195 | 474 | 105 |
| 205 | Royal Holloway University of London | 4 | 193 | 249 | 241 |
| 206 | York University | 7 | 191 | 1.1K | 1.2K |
| 207 | Mount Allison University | 2 | 188 | 231 | 76 |
| 208 | Lakehead University | 8 | 186 | 161 | 261 |
| 209 | Miguel Hernández University of Elche | 5 | 183 | 367 | 287 |
| 210 | Skidmore College | 3 | 182 | 107 | 61 |
| 211 | Leipzig University | 4 | 179 | 1.8K | 1.3K |
| 211 | University of Pittsburgh | 5 | 179 | 4.0K | 3.1K |
| 213 | United States Forest Service | 3 | 178 | 505 | 372 |
| 214 | University of Glasgow | 8 | 176 | 1.9K | 2.2K |
| 214 | Trinity College Dublin | 2 | 176 | 2.2K | 998 |
| 214 | Dalhousie University | 3 | 176 | 2.8K | 1.7K |
| 217 | Wright State University | 1 | 174 | 1.9K | 452 |
| 217 | University of South Alabama | 3 | 174 | 628 | 373 |
| 219 | University of Buenos Aires | 5 | 173 | 1.7K | 1.0K |
| 220 | Clarion University of Pennsylvania | 2 | 172 | 35 | 5 |
| 221 | University of Otago | 9 | 170 | 1.1K | 1.5K |
| 221 | Federal University of Uberlândia | 1 | 170 | 1.7K | 200 |
| 223 | Vanderbilt University | 2 | 169 | 3.9K | 2.0K |
| 224 | University of Minnesota Duluth | 6 | 168 | 80 | 133 |
| 225 | Åbo Akademi University | 7 | 167 | 269 | 377 |
| 226 | Humboldt University of Berlin | 1 | 165 | 4.7K | 1.6K |
| 227 | Bournemouth University | 5 | 163 | 259 | 246 |
| 228 | University of Milan | 3 | 162 | 3.8K | 2.4K |
| 229 | University of Adelaide | 4 | 161 | 2.8K | 2.2K |
| 230 | Michigan Technological University | 6 | 160 | 441 | 498 |
| 230 | Université catholique de Louvain | 1 | 160 | 4.5K | 1.7K |
| 230 | University of Tromso | 5 | 160 | 69 | 155 |
| 233 | Zoological Society of London | 6 | 158 | 93 | 131 |
| 234 | Zhejiang University | 6 | 157 | 3.0K | 2.4K |
| 235 | Technical University of Denmark | 7 | 156 | 1.2K | 1.5K |
| 236 | University of Warsaw | 3 | 153 | 1.8K | 696 |
| 236 | Polish Academy of Sciences | 7 | 153 | 1.7K | 1.7K |
| 236 | University of Liège | 3 | 153 | 2.2K | 1.5K |
| 239 | University of Miami | 4 | 150 | 2.5K | 2.1K |
| 239 | James Hutton Institute | 2 | 150 | 264 | 169 |
| 239 | North American Butterfly Association | 1 | 150 | 1 | 1 |
| 242 | University of Extremadura | 5 | 149 | 617 | 504 |
| 242 | University of Minnesota Twin Cities | 5 | 149 | 75 | 176 |
| 244 | Harvard University | 4 | 148 | 3.7K | 3.3K |
| 244 | Museo Nacional de Ciencias Naturales | 6 | 148 | 76 | 110 |
| 246 | North Carolina State University | 7 | 147 | 2.0K | 2.4K |
| 246 | Truman State University | 2 | 147 | 102 | 40 |
| 248 | University of Bristol | 5 | 144 | 2.9K | 2.7K |
| 248 | Indiana State University | 1 | 144 | 737 | 143 |
| 248 | University of Oregon | 3 | 144 | 1.8K | 1.3K |
| 251 | Ben Gurion University of the Negev | 3 | 143 | 648 | 416 |
| 252 | University of Nebraska–Lincoln | 4 | 142 | 1.4K | 1.2K |
| 253 | University of Bordeaux | 5 | 141 | 818 | 803 |
| 253 | Teton Science Schools | 1 | 141 | 2 | 2 |
| 255 | Natural History Museum | 5 | 140 | 483 | 493 |
| 256 | University of Tokyo | 7 | 139 | 4.1K | 3.7K |
| 256 | Leuphana University of Lüneburg | 4 | 139 | 97 | 124 |
| 256 | Pontifical Catholic University of Chile | 1 | 139 | 3.1K | 825 |
| 259 | Moscow State University | 5 | 138 | 1.8K | 1.2K |
| 259 | Concordia University | 3 | 138 | 1.8K | 1.1K |
| 259 | Victoria University of Wellington | 6 | 138 | 628 | 708 |
| 262 | University of Nevada, Las Vegas | 1 | 137 | 1.2K | 282 |
| 263 | National Marine Fisheries Service | 3 | 136 | 286 | 266 |
| 263 | University of Stirling | 3 | 136 | 1.0K | 777 |
| 265 | Stony Brook University | 4 | 134 | 2.0K | 1.8K |
| 265 | University of Wollongong | 7 | 134 | 951 | 1.3K |
| 267 | Acadia University | 2 | 132 | 291 | 127 |
| 267 | Södertörn University | 4 | 132 | 53 | 62 |
| 269 | National Autonomous University of Mexico | 5 | 130 | 2.3K | 1.6K |
| 269 | University of Regensburg | 1 | 130 | 2.8K | 1.2K |
| 269 | Syracuse University | 4 | 130 | 1.4K | 1.3K |
| 269 | University of Kassel | 1 | 130 | 1.4K | 397 |
| 269 | Federal Government of Belgium | 1 | 130 | 73 | 42 |
| 274 | University of Tromsø – The Arctic University of Norway | 6 | 128 | 610 | 889 |
| 274 | University of Nottingham | 3 | 128 | 4.1K | 2.9K |
| 274 | University of South Bohemia in České Budějovice | 8 | 128 | 162 | 284 |
| 274 | Palacký University Olomouc | 5 | 128 | 433 | 440 |
| 274 | AgroParisTech, Paris-Saclay University | 3 | 128 | 109 | 121 |
| 279 | University of Lausanne | 4 | 127 | 1.4K | 1.4K |
| 280 | University of New England | 4 | 126 | 671 | 658 |
| 280 | Fudan University | 5 | 126 | 2.8K | 2.3K |
| 282 | Milwaukee Public Museum | 1 | 125 | 44 | 13 |
| 283 | Pierre and Marie Curie University | 1 | 124 | 2.7K | 1.4K |
| 283 | Tulane University | 3 | 124 | 1.6K | 1.2K |
| 283 | University of Alcalá | 1 | 124 | 2.3K | 685 |
| 286 | Auburn University | 2 | 123 | 2.8K | 1.4K |
| 286 | University of Nevada, Reno | 5 | 123 | 1.4K | 1.4K |
| 288 | Russian Academy of Sciences | 2 | 122 | 4.6K | 2.2K |
| 288 | University of Kentucky | 5 | 122 | 2.7K | 2.7K |
| 288 | Chinese Academy of Agricultural Sciences | 4 | 122 | 818 | 691 |
| 291 | Kyushu University | 5 | 121 | 2.9K | 2.4K |
| 292 | Smithsonian Institution | 2 | 120 | 603 | 387 |
| 292 | Université de Montréal | 4 | 120 | 3.1K | 2.7K |
| 292 | University of Jaén | 2 | 120 | 949 | 415 |
| 292 | University of Reading | 3 | 120 | 2.0K | 1.6K |
| 296 | University of Chicago | 3 | 119 | 4.2K | 3.2K |
| 296 | University of Colorado Boulder | 4 | 119 | 1.3K | 1.4K |
| 298 | University of Fribourg | 2 | 118 | 992 | 623 |
| 298 | Cardiff University | 3 | 118 | 3.0K | 2.2K |
| 300 | University College London | 4 | 117 | 4.7K | 4.0K |
| 301 | ComUE Paris-Saclay University | 1 | 116 | 2.8K | 932 |
| 302 | Tufts University | 2 | 115 | 2.5K | 1.7K |
| 302 | University of Illinois at Chicago | 2 | 115 | 4.3K | 2.6K |
| 304 | Sun Yat-sen University | 3 | 113 | 3.4K | 2.3K |
| 304 | Chicago Botanic Garden | 2 | 113 | 33 | 26 |
| 306 | University of North Carolina at Chapel Hill | 4 | 112 | 3.0K | 2.9K |
| 306 | Western Washington University | 1 | 112 | 959 | 303 |
| 306 | Muséum national d'histoire naturelle | 8 | 112 | 226 | 476 |
| 306 | University of California, San Francisco | 1 | 112 | 3.7K | 2.1K |
| 310 | University of Coimbra | 6 | 111 | 946 | 1.3K |
| 311 | Universidad Nacional del Comahue | 2 | 109 | 222 | 105 |
| 312 | University of Western Australia | 5 | 107 | 2.4K | 2.7K |
| 312 | Bavarian Forest National Park | 2 | 107 | 30 | 28 |
| 314 | University of Vigo | 3 | 106 | 790 | 609 |
| 314 | Florida State University | 4 | 106 | 2.2K | 2.1K |
| 314 | Grinnell College | 1 | 106 | 279 | 49 |
| 317 | University of Cologne | 3 | 105 | 2.7K | 2.1K |
| 317 | University of Saskatchewan | 5 | 105 | 1.8K | 2.1K |
| 317 | University of the Free State | 3 | 105 | 700 | 393 |
| 317 | Sorbonne University | 8 | 105 | 485 | 657 |
| 321 | Virginia Tech | 4 | 103 | 2.9K | 2.8K |
| 321 | University of Granada | 1 | 103 | 4.7K | 1.8K |
| 321 | Loughborough University | 1 | 103 | 3.3K | 1.4K |
| 321 | University of Worcester | 1 | 103 | 329 | 89 |
| 325 | University of the Aegean | 3 | 102 | 391 | 308 |
| 325 | Chinese Academy of Tropical Agricultural Sciences | 2 | 102 | 219 | 105 |
| 327 | University of Debrecen | 3 | 101 | 1.1K | 793 |
| 327 | Université de Moncton | 1 | 101 | 617 | 162 |
| 329 | University of Houston | 3 | 99 | 2.5K | 1.9K |
| 329 | University of Mons | 2 | 99 | 280 | 179 |
| 331 | Peking University | 3 | 98 | 3.6K | 2.7K |
| 331 | University of Lethbridge | 2 | 98 | 626 | 422 |
| 333 | Central China Normal University | 2 | 97 | 830 | 420 |
| 334 | University of Hamburg | 2 | 96 | 3.0K | 1.9K |
| 334 | Duke University | 5 | 96 | 2.7K | 3.2K |
| 334 | Tokyo University of Agriculture and Technology | 1 | 96 | 1.7K | 684 |
| 337 | Natural Resources Canada | 3 | 95 | 404 | 409 |
| 338 | Nelson Mandela University | 3 | 94 | 318 | 188 |
| 338 | University of Toledo | 2 | 94 | 1.6K | 981 |
| 340 | Queen Mary University of London | 2 | 93 | 1.7K | 1.2K |
| 341 | Tohoku University | 5 | 92 | 2.6K | 2.5K |
| 341 | University of the Balearic Islands | 1 | 92 | 1.2K | 481 |
| 343 | University of North Dakota | 2 | 91 | 888 | 593 |
| 343 | Clark University | 1 | 91 | 960 | 463 |
| 345 | Nanjing University | 2 | 90 | 2.8K | 1.8K |
| 345 | Charles Darwin University | 1 | 90 | 915 | 397 |
| 347 | Towson University | 1 | 87 | 1.0K | 278 |
| 347 | University of California, Riverside | 5 | 87 | 458 | 935 |
| 349 | University of Kiel | 1 | 85 | 2.1K | 1.2K |
| 350 | University of Victoria | 2 | 84 | 2.5K | 1.8K |
| 350 | French Agricultural Research Centre for International Development | 7 | 84 | 200 | 382 |
| 352 | Technical University of Munich | 2 | 83 | 4.4K | 3.2K |
| 352 | Rhodes University | 2 | 83 | 786 | 478 |
| 354 | Memorial University of Newfoundland | 5 | 81 | 1.1K | 1.5K |
| 354 | South Dakota State University | 1 | 81 | 1.1K | 539 |
| 354 | Lanzhou University | 4 | 81 | 1.3K | 1.2K |
| 354 | UMR BIOdiversity, GEnes & Communities | 3 | 81 | 34 | 51 |
| 358 | European Telecommunications Standards Institute | 1 | 80 | 22 | 9 |
| 358 | Bridgewater State University | 1 | 80 | 170 | 49 |
| 358 | Wollo University | 1 | 80 | 187 | 44 |
| 361 | Senckenberg Biodiversity and Climate Research Centre | 3 | 79 | 19 | 40 |
| 362 | University of Padua | 2 | 77 | 5.2K | 3.8K |
| 362 | Ohio State University | 4 | 77 | 5.5K | 5.4K |
| 362 | Appalachian State University | 4 | 77 | 306 | 385 |
| 362 | Macaulay Institute | 3 | 77 | 143 | 217 |
| 366 | Columbia University | 3 | 75 | 5.7K | 5.1K |
| 367 | Wellesley College | 2 | 74 | 394 | 367 |
| 367 | Murdoch University | 3 | 74 | 946 | 1.1K |
| 369 | Ryerson University | 2 | 73 | 1.3K | 891 |
| 369 | University of Salento | 2 | 73 | 1.1K | 811 |
| 369 | Furman University | 1 | 73 | 303 | 120 |
| 369 | Penn State Cancer Institute | 2 | 73 | 171 | 202 |
| 373 | University of Waterloo | 2 | 71 | 4.1K | 3.1K |
| 374 | University of Giessen | 3 | 70 | 1.8K | 1.9K |
| 375 | National University of La Pampa | 1 | 69 | 149 | 45 |
| 376 | Masaryk University | 3 | 68 | 1.2K | 1.1K |
| 376 | Nottingham Trent University | 2 | 68 | 1.2K | 912 |
| 378 | Bangor University | 2 | 67 | 1.2K | 1.0K |
| 378 | Case Western Reserve University | 5 | 67 | 2.4K | 3.3K |
| 378 | University of Eastern Finland | 2 | 67 | 1.4K | 1.1K |
| 381 | Dartmouth College | 3 | 66 | 1.3K | 1.6K |
| 381 | University of Notre Dame Australia | 1 | 66 | 1.4K | 807 |
| 381 | Henan University of Science and Technology | 1 | 66 | 1.0K | 389 |
| 381 | California State University Channel Islands | 1 | 66 | 74 | 27 |
| 385 | Freie Universität Berlin | 2 | 65 | 3.1K | 2.5K |
| 385 | Indian Institute of Science | 2 | 65 | 2.2K | 1.6K |
| 385 | Colorado Mesa University | 1 | 65 | 77 | 49 |
| 385 | University of North Carolina at Charlotte | 2 | 65 | 1.4K | 1.1K |
| 385 | Federal University of Viçosa | 2 | 65 | 1.3K | 746 |
| 385 | Mediterranean Center for Environmental Studies | 1 | 65 | 14 | 16 |
| 391 | United States Environmental Protection Agency | 3 | 64 | 463 | 674 |
| 392 | National Tsing Hua University | 1 | 63 | 2.4K | 1.3K |
| 392 | University of Kansas | 3 | 63 | 3.3K | 3.3K |
| 392 | TU Dresden | 3 | 63 | 2.2K | 2.4K |
| 392 | University of Porto | 3 | 63 | 1.8K | 1.9K |
| 392 | Institute for Biodiversity and Ecosystem Dynamics | 2 | 63 | 18 | 39 |
| 392 | National Institute for Environmental Studies | 3 | 63 | 498 | 575 |
| 398 | University of London | 3 | 62 | 4.6K | 4.5K |
| 399 | University of South Florida | 2 | 61 | 4.2K | 3.3K |
| 399 | Laval University | 3 | 61 | 3.0K | 3.2K |
| 399 | Indiana University Bloomington | 2 | 61 | 980 | 989 |
| 399 | Claude Bernard University Lyon 1 | 4 | 61 | 1.2K | 1.6K |
| 403 | Boise State University | 4 | 60 | 315 | 604 |
| 403 | University of Puerto Rico, Río Piedras Campus | 1 | 60 | 73 | 62 |
| 405 | The Nature Conservancy | 2 | 59 | 218 | 241 |
| 405 | Hope College | 1 | 59 | 289 | 155 |
| 405 | Northeastern University | 2 | 59 | 3.6K | 2.9K |
| 408 | University of Auckland | 4 | 58 | 3.0K | 3.8K |
| 408 | University of Missouri | 4 | 58 | 3.3K | 4.0K |
| 410 | Aristotle University of Thessaloniki | 1 | 57 | 5.3K | 2.9K |
| 410 | University of Florence | 2 | 57 | 3.8K | 3.1K |
| 410 | Slovak Academy of Sciences | 2 | 57 | 1.6K | 1.3K |
| 410 | Catholic University of the Most Holy Conception | 1 | 57 | 304 | 68 |
| 410 | University of Abertay Dundee | 2 | 57 | 173 | 204 |
| 410 | University of Hawaii at Manoa | 3 | 57 | 1.0K | 1.5K |
| 416 | China Agricultural University | 3 | 56 | 1.3K | 1.4K |
| 416 | University of Burgundy | 1 | 56 | 1.5K | 1.1K |
| 416 | Nanyang Technological University | 3 | 56 | 2.8K | 2.9K |
| 416 | Balaton Limnological Research Institute | 1 | 56 | 28 | 21 |
| 416 | University of Toronto Scarborough | 1 | 56 | 609 | 425 |
| 421 | Wesleyan University | 2 | 55 | 567 | 555 |
| 421 | University of Greifswald | 3 | 55 | 783 | 1.0K |
| 421 | Brigham Young University | 4 | 55 | 1.3K | 2.0K |
| 421 | Alfred Wegener Institute for Polar and Marine Research | 1 | 55 | 409 | 359 |
| 421 | Newcastle University | 2 | 55 | 4.0K | 3.4K |
| 421 | University of Essex | 3 | 55 | 1.2K | 1.4K |
| 421 | King Abdullah University of Science and Technology | 1 | 55 | 1.4K | 935 |
| 421 | Wilfrid Laurier University | 2 | 55 | 932 | 836 |
| 421 | Aix-Marseille University | 4 | 55 | 1.2K | 1.7K |
| 421 | Australian Museum | 1 | 55 | 202 | 144 |
| 421 | Kristianstad University | 2 | 55 | 120 | 145 |
| 432 | University of Regina | 2 | 54 | 888 | 762 |
| 433 | Technical University of Braunschweig | 1 | 53 | 2.0K | 1.3K |
| 433 | Binghamton University | 1 | 53 | 1.7K | 972 |
| 433 | University of Innsbruck | 3 | 53 | 1.6K | 2.0K |
| 433 | University of South Carolina | 1 | 53 | 5.4K | 3.5K |
| 433 | Yokohama National University | 2 | 53 | 1.0K | 707 |
| 433 | University of Louisville | 2 | 53 | 2.8K | 2.4K |
| 433 | University of Canberra | 2 | 53 | 733 | 706 |
| 433 | Virginia Institute of Marine Science | 2 | 53 | 218 | 242 |
| 433 | Sorbonne Universités | 3 | 53 | 601 | 957 |
| 442 | Rensselaer Polytechnic Institute | 4 | 52 | 1.1K | 1.7K |
| 442 | University of Pannonia | 2 | 52 | 385 | 361 |
| 442 | Smithsonian Conservation Biology Institute | 1 | 52 | 186 | 133 |
| 442 | University of Mississippi | 1 | 52 | 2.1K | 1.2K |
| 442 | East Carolina University | 4 | 52 | 953 | 1.6K |
| 442 | National Zoological Park | 1 | 52 | 209 | 183 |
| 442 | University of Virginia | 5 | 52 | 2.7K | 4.1K |
| 449 | Oxford Brookes University | 1 | 51 | 1.6K | 908 |
| 449 | Moredun Research Institute | 1 | 51 | 180 | 145 |
| 451 | Jagiellonian University | 3 | 50 | 1.9K | 1.8K |
| 451 | University of Chile | 1 | 50 | 4.2K | 2.5K |
| 451 | University of California, Irvine | 2 | 50 | 2.0K | 2.2K |
| 454 | Goethe University Frankfurt | 4 | 48 | 1.6K | 2.5K |
| 454 | University of Trento | 1 | 48 | 2.5K | 1.6K |
| 454 | University of Arkansas | 3 | 48 | 1.7K | 2.0K |
| 454 | PSL University | 2 | 48 | 885 | 806 |
| 454 | Texas Tech University | 5 | 48 | 1.2K | 2.2K |
| 454 | Roskilde University | 2 | 48 | 533 | 588 |
| 454 | Jiangxi Normal University | 2 | 48 | 526 | 405 |
| 461 | Algoma University | 2 | 47 | 21 | 56 |
| 461 | Czech University of Life Sciences Prague | 3 | 47 | 266 | 325 |
| 461 | Shanghai University of Medicine and Health Sciences | 1 | 47 | 136 | 67 |
| 464 | École polytechnique fédérale de Lausanne | 2 | 45 | 1.8K | 1.9K |
| 464 | University of Seville | 1 | 45 | 4.3K | 2.3K |
| 464 | New Mexico State University | 3 | 45 | 1.0K | 1.4K |
| 464 | Southern Illinois University Carbondale | 2 | 45 | 2.1K | 1.9K |
| 464 | University of Puerto Rico | 2 | 45 | 1.2K | 1.2K |
| 464 | National Scientific and Technical Research Council | 6 | 45 | 646 | 1.4K |
| 470 | Tel Aviv University | 3 | 44 | 4.4K | 4.8K |
| 470 | National Institute of Water and Atmospheric Research | 1 | 44 | 314 | 292 |
| 470 | Loyola University Chicago | 1 | 44 | 2.2K | 1.4K |
| 470 | Halmstad University | 1 | 44 | 313 | 239 |
| 474 | University of Ljubljana | 1 | 43 | 4.1K | 2.6K |
| 474 | Ehime University | 2 | 43 | 1.2K | 1.1K |
| 474 | Peking Union Medical College | 1 | 43 | 2.7K | 1.9K |
| 474 | Fram Centre | 1 | 43 | 69 | 71 |
| 474 | École Normale Supérieure de Lyon | 1 | 43 | 1.0K | 784 |
| 479 | California Institute of Technology | 1 | 42 | 3.0K | 2.5K |
| 479 | Academia Sinica | 2 | 42 | 2.6K | 2.4K |
| 481 | Drexel University | 1 | 41 | 3.1K | 2.2K |
| 481 | University of Southern Mississippi | 4 | 41 | 515 | 1.0K |
| 481 | Senckenberg Nature Research Society | 1 | 41 | 54 | 69 |
| 481 | University of Manitoba | 2 | 41 | 4.0K | 3.8K |
| 481 | Rhodes College | 1 | 41 | 228 | 137 |
| 481 | Academy of Natural Sciences of Drexel University | 1 | 41 | 146 | 141 |
| 481 | Nicholas School of the Environment | 3 | 41 | 206 | 420 |
| 488 | National Science Foundation | 1 | 40 | 432 | 364 |
| 488 | Technion – Israel Institute of Technology | 2 | 40 | 2.8K | 2.8K |
| 488 | University of Hohenheim | 3 | 40 | 786 | 1.2K |
| 488 | City University of New York | 2 | 40 | 3.5K | 3.2K |
| 488 | Washington University in St. Louis | 2 | 40 | 165 | 338 |
| 488 | University of Nebraska system | 1 | 40 | 3 | 7 |
| 488 | University of Namur | 2 | 40 | 469 | 662 |
| 488 | University of Bath | 2 | 40 | 2.5K | 2.7K |
| 496 | University of São Paulo | 4 | 39 | 5.5K | 5.7K |
| 496 | University of Alaska Fairbanks | 4 | 39 | 354 | 763 |
| 498 | University of Oldenburg | 1 | 38 | 1.1K | 836 |
| 498 | Trinity University | 1 | 38 | 522 | 342 |
| 498 | University of Wisconsin–Milwaukee | 1 | 38 | 3.1K | 2.2K |
| 498 | University of Hawaiʻi at Hilo | 1 | 38 | 299 | 273 |
| 498 | Sokoine University of Agriculture | 2 | 38 | 257 | 282 |
| 498 | Nature Conservation Foundation | 2 | 38 | 26 | 39 |
| 504 | Boston University | 2 | 37 | 4.1K | 4.2K |
| 504 | University of California, Santa Cruz | 2 | 37 | 681 | 1.0K |
| 506 | University of Münster | 3 | 36 | 2.3K | 3.1K |
| 506 | Oak Ridge National Laboratory | 2 | 36 | 1.8K | 1.9K |
| 506 | Oklahoma State University–Stillwater | 1 | 36 | 216 | 185 |
| 506 | University of Oviedo | 1 | 36 | 2.8K | 2.0K |
| 506 | University of Massachusetts Dartmouth | 1 | 36 | 672 | 519 |
| 506 | Australian Wildlife Conservancy | 1 | 36 | 28 | 28 |
| 512 | Bar-Ilan University | 2 | 35 | 2.2K | 2.2K |
| 512 | University of Parma | 2 | 35 | 2.0K | 2.3K |
| 512 | Nanjing Agricultural University | 2 | 35 | 1.2K | 1.2K |
| 512 | Louisiana State University Agricultural Center | 2 | 35 | 325 | 455 |
| 516 | University of Rochester | 2 | 34 | 3.5K | 3.7K |
| 516 | Wayne State University | 2 | 34 | 3.8K | 3.9K |
| 516 | National and Kapodistrian University of Athens | 1 | 34 | 5.2K | 3.8K |
| 516 | University of Kent | 1 | 34 | 2.9K | 2.1K |
| 516 | University of Windsor | 2 | 34 | 1.5K | 1.6K |
| 516 | Sea Mammal Research Unit | 2 | 34 | 39 | 76 |
| 516 | Tyndall Centre | 1 | 34 | 79 | 122 |
| 516 | Natural England | 1 | 34 | 63 | 74 |
| 516 | Lancaster University | 4 | 34 | 1.2K | 2.4K |
| 525 | Cranfield University | 1 | 33 | 1.6K | 1.4K |
| 525 | University of Dundee | 1 | 33 | 2.9K | 2.5K |
| 525 | California State University, Long Beach | 1 | 33 | 671 | 528 |
| 525 | National Geographic Society | 1 | 33 | 42 | 56 |
| 529 | University of Montpellier | 2 | 32 | 1.6K | 1.9K |
| 529 | Microsoft Research | 1 | 32 | 589 | 576 |
| 529 | University of Lapland | 1 | 32 | 239 | 189 |
| 529 | Capital Normal University | 1 | 32 | 1.0K | 746 |
| 529 | Xinjiang Agricultural University | 1 | 32 | 242 | 159 |
| 534 | University of Valladolid | 1 | 31 | 2.2K | 1.5K |
| 534 | Charles Darwin Foundation | 1 | 31 | 52 | 69 |
| 534 | University College Dublin | 2 | 31 | 2.9K | 3.4K |
| 534 | Avignon University | 3 | 31 | 86 | 237 |
| 534 | Fujian Normal University | 1 | 31 | 1.0K | 750 |
| 539 | University of Hong Kong | 1 | 30 | 6.7K | 5.3K |
| 539 | University of Northern British Columbia | 1 | 30 | 607 | 564 |
| 539 | Shinshu University | 1 | 30 | 2.0K | 1.4K |
| 539 | Edith Cowan University | 1 | 30 | 1.6K | 1.4K |
| 539 | University of Colorado Museum of Natural History | 1 | 30 | 16 | 18 |
| 539 | Drake University | 1 | 30 | 352 | 262 |
| 539 | Penn State Schuylkill | 1 | 30 | 10 | 9 |
| 539 | CSIRO Oceans and Atmosphere | 2 | 30 | 195 | 311 |
| 539 | Duke University Marine Laboratory | 1 | 30 | 142 | 183 |
| 548 | Nagasaki University | 1 | 29 | 2.0K | 1.4K |
| 548 | Austrian Academy of Sciences | 1 | 29 | 1.1K | 1.0K |
| 548 | Bowdoin College | 1 | 29 | 465 | 363 |
| 548 | Musée de l'Homme | 1 | 29 | 41 | 47 |
| 548 | Cévennes National Park | 1 | 29 | 1 | 1 |
| 553 | Johannes Gutenberg University Mainz | 1 | 28 | 2.8K | 2.5K |
| 553 | University of Siena | 2 | 28 | 2.1K | 2.6K |
| 553 | Western Michigan University | 1 | 28 | 1.8K | 1.3K |
| 553 | Royal Botanic Garden Edinburgh | 3 | 28 | 70 | 136 |
| 553 | Mississippi State University | 2 | 28 | 1.9K | 2.2K |
| 553 | Smithsonian Environmental Research Center | 3 | 28 | 103 | 262 |
| 553 | Tanzania Wildlife Research Institute | 1 | 28 | 43 | 50 |
| 560 | Technical University of Madrid | 1 | 27 | 2.4K | 1.9K |
| 560 | University of the Western Cape | 1 | 27 | 1.4K | 1.1K |
| 560 | Ohio University | 1 | 27 | 2.5K | 2.1K |
| 560 | Wuhan University | 1 | 27 | 4.0K | 3.2K |
| 560 | Middlebury College | 3 | 27 | 178 | 420 |
| 560 | Thompson Rivers University | 1 | 27 | 317 | 243 |
| 560 | University of the Littoral Opal Coast | 1 | 27 | 258 | 257 |
| 567 | University of Alaska Anchorage | 1 | 26 | 430 | 397 |
| 567 | National University of Córdoba | 1 | 26 | 1.9K | 1.3K |
| 569 | University of Central Florida | 1 | 25 | 3.8K | 3.2K |
| 569 | National Centre for Biological Sciences | 2 | 25 | 160 | 281 |
| 569 | University of Southern Denmark | 2 | 25 | 2.2K | 2.9K |
| 569 | University of Wisconsin–Oshkosh | 1 | 25 | 475 | 410 |
| 569 | Royal Society for the Protection of Birds | 3 | 25 | 16 | 49 |
| 569 | Universidad Politécnica Estatal del Carchi | 4 | 25 | 47 | 260 |
| 575 | Université du Québec à Chicoutimi | 1 | 24 | 440 | 434 |
| 575 | Denison University | 1 | 24 | 318 | 225 |
| 575 | Black Hills State University | 1 | 24 | 56 | 62 |
| 575 | Georgetown University Medical Center | 1 | 24 | 1.4K | 1.5K |
| 575 | Natural History Museum of Utah | 1 | 24 | 42 | 58 |
| 580 | Complutense University of Madrid | 2 | 23 | 4.3K | 4.3K |
| 580 | King's College London | 2 | 23 | 4.3K | 5.0K |
| 580 | Nanzan University | 1 | 23 | 143 | 75 |
| 580 | Agricultural Research Service | 3 | 23 | 800 | 1.3K |
| 580 | Doane University | 1 | 23 | 31 | 29 |
| 580 | Universidade Federal de São Carlos | 1 | 23 | 2.5K | 1.7K |
| 580 | Academy of Scientific and Innovative Research | 1 | 23 | 558 | 513 |
| 580 | Yukon College | 1 | 23 | 3 | 18 |
| 588 | University of Vienna | 1 | 22 | 5.5K | 4.8K |
| 588 | University of Zagreb | 1 | 22 | 3.8K | 3.0K |
| 588 | University of Nice Sophia Antipolis | 1 | 22 | 1.2K | 1.3K |
| 588 | Chapman University | 1 | 22 | 670 | 665 |
| 588 | Gansu Agricultural University | 1 | 22 | 418 | 351 |
| 588 | Presbyterian College | 1 | 22 | 37 | 43 |
| 588 | Universidade de Santiago de Compostela | 1 | 22 | 1.4K | 1.2K |
| 595 | University of Iceland | 2 | 21 | 1.2K | 1.9K |
| 595 | Saint Louis University | 1 | 21 | 2.4K | 2.2K |
| 595 | Nara Women's University | 1 | 21 | 521 | 480 |
| 595 | Grand Valley State University | 1 | 21 | 748 | 734 |
| 595 | Field Museum of Natural History | 1 | 21 | 360 | 386 |
| 600 | National Cheng Kung University | 3 | 20 | 2.2K | 3.4K |
| 600 | Mie University | 1 | 20 | 1.5K | 1.3K |
| 600 | Colgate University | 1 | 20 | 637 | 596 |
| 600 | East China Normal University | 1 | 20 | 2.5K | 2.2K |
| 600 | École nationale des travaux publics de l'État | 1 | 20 | 160 | 175 |
| 605 | Amherst College | 1 | 19 | 476 | 511 |
| 605 | Johns Hopkins University | 1 | 19 | 7.4K | 6.8K |
| 605 | University of Hawaii | 1 | 19 | 3.0K | 3.1K |
| 605 | Loyola University | 1 | 19 | 984 | 1.0K |
| 605 | United States Fish and Wildlife Service | 2 | 19 | 96 | 175 |
| 605 | Baylor University | 1 | 19 | 2.0K | 1.9K |
| 605 | Reed College | 2 | 19 | 199 | 404 |
| 605 | Curtin University | 2 | 19 | 3.0K | 4.2K |
| 605 | University of Central Arkansas | 1 | 19 | 398 | 378 |
| 605 | Prairie Research Institute | 1 | 19 | 7 | 19 |
| 605 | University of Maryland, Baltimore County | 1 | 19 | 1.1K | 1.3K |
| 605 | Brazilian Agricultural Research Corporation | 1 | 19 | 499 | 528 |
| 617 | Sapienza University of Rome | 1 | 18 | 6.2K | 5.7K |
| 617 | Chongqing University | 1 | 18 | 2.6K | 2.3K |
| 617 | California Polytechnic State University | 2 | 18 | 446 | 978 |
| 617 | Kenyon College | 1 | 18 | 247 | 285 |
| 617 | Westmont College | 1 | 18 | 65 | 55 |
| 622 | University of Bologna | 1 | 17 | 6.8K | 6.3K |
| 622 | University of Denver | 1 | 17 | 1.9K | 1.9K |
| 622 | University of Surrey | 1 | 17 | 3.7K | 3.8K |
| 622 | Murray State University | 1 | 17 | 421 | 462 |
| 622 | Osnabrück University | 1 | 17 | 1.3K | 1.4K |
| 622 | Ryukoku University | 1 | 17 | 487 | 395 |
| 622 | African Institute for Mathematical Sciences | 1 | 17 | 37 | 52 |
| 622 | Plantentuin Meise | 1 | 17 | 94 | 122 |
| 622 | Southwest Jiaotong University | 1 | 17 | 1.6K | 1.4K |
| 631 | Tata Institute of Fundamental Research | 1 | 16 | 1.0K | 998 |
| 631 | University of Geneva | 1 | 16 | 4.1K | 4.2K |
| 631 | Estonian University of Life Sciences | 1 | 16 | 416 | 473 |
| 631 | Doshisha University | 1 | 16 | 993 | 848 |
| 631 | California Department of Parks and Recreation | 1 | 16 | 6 | 7 |
| 636 | World Conservation Monitoring Centre | 1 | 15 | 26 | 54 |
| 636 | Bryn Mawr College | 1 | 15 | 574 | 652 |
| 636 | Paris Diderot University | 2 | 15 | 1.6K | 2.5K |
| 636 | University of North Carolina at Greensboro | 1 | 15 | 1.7K | 1.9K |
| 636 | University of Rhode Island | 1 | 15 | 2.4K | 2.7K |
| 636 | Beijing Forestry University | 1 | 15 | 1.0K | 1.1K |
| 636 | Medaille College | 1 | 15 | 12 | 29 |
| 636 | Otsuma Women's University | 1 | 15 | 101 | 117 |
| 636 | Patuxent Wildlife Research Center | 2 | 15 | 103 | 191 |
| 636 | California State University, Dominguez Hills | 2 | 15 | 35 | 132 |
| 646 | Aalborg University | 1 | 14 | 2.9K | 3.1K |
| 646 | Texas State University | 1 | 14 | 1.8K | 1.9K |
| 646 | University of Cádiz | 1 | 14 | 1.6K | 1.6K |
| 646 | Northwest University | 1 | 14 | 1.6K | 1.6K |
| 646 | University of Central Oklahoma | 1 | 14 | 221 | 288 |
| 646 | Gävle University College | 1 | 14 | 325 | 456 |
| 646 | National Museums Scotland | 1 | 14 | 90 | 106 |
| 653 | University of Valencia | 4 | 13 | 2.3K | 4.6K |
| 653 | Baruch College | 1 | 13 | 617 | 700 |
| 653 | Shizuoka University | 1 | 13 | 1.6K | 1.6K |
| 653 | Saint Petersburg State Agrarian University | 1 | 13 | 20 | 6 |
| 653 | Autonomous University of Yucatán | 1 | 13 | 510 | 542 |
| 653 | Federal University of the State of Rio de Janeiro | 1 | 13 | 407 | 357 |
| 653 | ICRA Limited | 1 | 13 | 31 | 50 |
| 660 | Wichita State University | 1 | 12 | 1.2K | 1.4K |
| 660 | University of Limerick | 1 | 12 | 1.9K | 2.3K |
| 660 | University of Waikato | 1 | 12 | 1.8K | 2.2K |
| 660 | Emory University | 1 | 12 | 5.2K | 5.5K |
| 660 | University of Delhi | 1 | 12 | 3.1K | 3.1K |
| 660 | Institut National de Recherche en Informatique et en Automatique | 1 | 12 | 796 | 952 |
| 660 | Santa Fe Institute | 2 | 12 | 84 | 289 |
| 660 | Villanova University | 1 | 12 | 1.3K | 1.5K |
| 660 | University of Santiago de Compostela | 1 | 12 | 2.4K | 2.7K |
| 660 | University of North Florida | 1 | 12 | 802 | 952 |
| 660 | University of Dayton | 1 | 12 | 1.2K | 1.4K |
| 660 | Mote Marine Laboratory | 1 | 12 | 70 | 126 |
| 660 | University of Côte d'Azur | 1 | 12 | 682 | 872 |
| 660 | European Commission | 1 | 12 | 1.4K | 1.8K |
| 674 | University of Dar es Salaam | 1 | 11 | 653 | 821 |
| 674 | University of Missouri–St. Louis | 1 | 11 | 153 | 331 |
| 674 | Budapest University of Technology and Economics | 1 | 11 | 1.7K | 1.8K |
| 674 | Central University of Kerala | 1 | 11 | 137 | 208 |
| 674 | Institut national des sciences appliquées Centre Val de Loire | 1 | 11 | 119 | 141 |
| 674 | Royal Holloway, University of London | 1 | 11 | 1.6K | 2.1K |
| 674 | Conservation International | 1 | 11 | 125 | 215 |
| 674 | New England Aquarium | 1 | 11 | 70 | 113 |
| 682 | Flinders University | 1 | 10 | 3.2K | 3.9K |
| 682 | University of San Diego | 1 | 10 | 634 | 957 |
| 682 | Kasetsart University | 1 | 10 | 1.5K | 1.7K |
| 682 | Mahasarakham University | 1 | 10 | 419 | 558 |
| 682 | Chulalongkorn University | 1 | 10 | 3.1K | 3.5K |
| 682 | Colorado College | 1 | 10 | 234 | 340 |
| 682 | University of the Ryukyus | 1 | 10 | 1.8K | 1.9K |
| 682 | University of South Dakota | 1 | 10 | 1.0K | 1.4K |
| 682 | University of Évora | 1 | 10 | 878 | 1.0K |
| 682 | Harper Adams University | 1 | 10 | 198 | 298 |
| 692 | National Taiwan University | 1 | 9 | 5.4K | 5.8K |
| 692 | Hiroshima University | 1 | 9 | 4.0K | 4.2K |
| 692 | Stazione Zoologica Anton Dohrn | 1 | 9 | 263 | 432 |
| 692 | Yunnan University | 1 | 9 | 1.4K | 1.7K |
| 692 | Ștefan cel Mare University of Suceava | 1 | 9 | 135 | 186 |
| 692 | University of New Haven | 1 | 9 | 253 | 408 |
| 692 | Griffith University | 1 | 9 | 4.7K | 5.5K |
| 692 | University of Campinas | 1 | 9 | 4.6K | 4.4K |
| 692 | Huazhong Agricultural University | 1 | 9 | 1.5K | 1.8K |
| 692 | Agricultural University of Hebei | 1 | 9 | 568 | 736 |
| 692 | Brandon University | 1 | 9 | 299 | 456 |
| 692 | Ikerbasque | 1 | 9 | 268 | 509 |
| 692 | Natural Resources Research Institute | 1 | 9 | 94 | 184 |
| 692 | Canadian Museum of Nature | 1 | 9 | 171 | 279 |
| 692 | British Antarctic Survey | 1 | 9 | 439 | 589 |
| 692 | Natural Environment Research Council | 1 | 9 | 316 | 488 |
| 708 | Williams College | 1 | 8 | 610 | 872 |
| 708 | University of Tsukuba | 1 | 8 | 4.8K | 5.1K |
| 708 | Georgia Institute of Technology | 1 | 8 | 4.5K | 5.3K |
| 708 | Evergreen State College | 1 | 8 | 104 | 218 |
| 708 | College of Wooster | 1 | 8 | 185 | 312 |
| 708 | French National Institute of Health and Medical Research | 1 | 8 | 3.0K | 3.6K |
| 708 | Pima Community College | 1 | 8 | 17 | 34 |
| 708 | Tokyo City University | 1 | 8 | 493 | 493 |
| 708 | Inland Norway University of Applied Sciences | 1 | 8 | 152 | 259 |
| 708 | University of Agriculture | 1 | 8 | 1.6K | 1.9K |
| 718 | University of Southern California | 1 | 7 | 7.9K | 8.8K |
| 718 | University of Cincinnati | 1 | 7 | 5.3K | 6.2K |
| 718 | Osaka Prefecture University | 1 | 7 | 1.9K | 2.2K |
| 718 | Georgia Southern University | 1 | 7 | 1.2K | 1.8K |
| 718 | University of Lisbon | 2 | 7 | 3.3K | 5.4K |
| 718 | Mount Holyoke College | 1 | 7 | 424 | 705 |
| 718 | Queensland University of Technology | 1 | 7 | 4.1K | 5.4K |
| 718 | University of Murcia | 1 | 7 | 3.0K | 3.5K |
| 718 | Florida Gulf Coast University | 1 | 7 | 333 | 561 |
| 718 | Kent State University | 1 | 7 | 2.7K | 3.5K |
| 718 | University of Algarve | 1 | 7 | 1.1K | 1.7K |
| 718 | Netherlands Environmental Assessment Agency | 1 | 7 | 54 | 126 |
| 718 | University of Savoy | 1 | 7 | 14 | 65 |
| 718 | Northeast Normal University | 1 | 7 | 1.3K | 1.6K |
| 718 | Mianyang Normal University | 1 | 7 | 96 | 159 |
| 718 | Toulouse III University - Paul-Sabatier | 1 | 7 | 483 | 1.0K |
| 734 | Meiji University | 1 | 6 | 1.0K | 1.2K |
| 734 | NOAA | 1 | 6 | 807 | 1.1K |
| 734 | Pondicherry University | 1 | 6 | 855 | 1.2K |
| 734 | Université du Québec | 1 | 6 | 1.1K | 1.9K |
| 734 | Kagoshima University | 1 | 6 | 2.1K | 2.6K |
| 734 | University of Delaware | 2 | 6 | 3.1K | 5.3K |
| 734 | University of Mauritius | 1 | 6 | 447 | 718 |
| 734 | DePaul University | 1 | 6 | 1.4K | 1.9K |
| 734 | Botanical Survey of India | 1 | 6 | 78 | 92 |
| 734 | Universidade Federal de Sergipe | 1 | 6 | 1.4K | 1.4K |
| 734 | Wittenberg University | 1 | 6 | 136 | 199 |
| 734 | Agriculture and Agriculture-Food Canada | 1 | 6 | 583 | 883 |
| 746 | Southern Utah University | 1 | 5 | 84 | 214 |
| 746 | Monmouth University | 1 | 5 | 250 | 444 |
| 746 | Xiamen University | 1 | 5 | 3.2K | 4.0K |
| 746 | Tallinn University | 1 | 5 | 346 | 561 |
| 746 | Sam Houston State University | 1 | 5 | 808 | 1.3K |
| 746 | Hatfield Marine Science Center | 1 | 5 | 122 | 207 |
| 746 | Beijing Normal University | 1 | 5 | 2.9K | 3.7K |
| 746 | Audubon | 1 | 5 | 36 | 82 |
| 746 | Tuscia University | 1 | 5 | 849 | 1.4K |
| 746 | University of Texas at El Paso | 1 | 5 | 1.8K | 2.7K |
| 746 | Graduate University for Advanced Studies | 1 | 5 | 139 | 261 |
| 746 | Taizhou University | 1 | 5 | 467 | 733 |
| 746 | Animal and Plant Health Agency | 1 | 5 | 57 | 114 |
| 746 | Florida Museum of Natural History | 1 | 5 | 283 | 426 |
| 760 | Bielefeld University | 1 | 4 | 2.4K | 3.5K |
| 760 | University of Aveiro | 1 | 4 | 2.6K | 3.7K |
| 760 | University of Koblenz - Landau | 1 | 4 | 371 | 801 |
| 760 | University of Naples Federico II | 1 | 4 | 4.9K | 6.3K |
| 760 | University of Duisburg-Essen | 1 | 4 | 3.4K | 4.6K |
| 760 | University of Warwick | 1 | 4 | 5.2K | 6.7K |
| 760 | University of Rennes | 1 | 4 | 287 | 795 |
| 760 | University of Udine | 1 | 4 | 2.0K | 3.1K |
| 760 | Norwegian Environment Agency | 1 | 4 | 4 | 23 |
| 760 | Royal Zoological Society of Antwerp | 1 | 4 | 27 | 66 |
| 770 | Ludwig Maximilian University of Munich | 1 | 3 | 5.7K | 7.2K |
| 770 | University of Łódź | 1 | 3 | 1.8K | 2.3K |
| 770 | Ruhr University Bochum | 1 | 3 | 4.0K | 5.4K |
| 770 | University of Sussex | 1 | 3 | 3.6K | 5.0K |
| 770 | University of Ostrava | 1 | 3 | 427 | 806 |
| 770 | University of Gloucestershire | 1 | 3 | 357 | 720 |
| 770 | Wrocław University of Environmental and Life Sciences | 1 | 3 | 632 | 1.0K |
| 770 | University of Toronto Mississauga | 1 | 3 | 674 | 1.3K |
| 770 | Poznań University of Life Sciences | 1 | 3 | 630 | 1.1K |
| 779 | Massey University | 1 | 2 | 3.0K | 4.7K |
| 779 | Western Sydney University | 1 | 2 | 2.8K | 4.7K |
| 779 | Heriot-Watt University | 1 | 2 | 2.1K | 3.4K |
| 779 | National Ecological Observatory Network | 1 | 2 | 24 | 80 |
| 779 | Federal Rural University of Rio de Janeiro | 1 | 2 | 1.0K | 1.3K |
| 779 | University of Iowa | 1 | 2 | 6.5K | 8.5K |
| 785 | Georgia State University | 1 | 1 | 3.4K | 5.3K |
| 785 | Rocky Mountain Research Station | 1 | 1 | 291 | 489 |
| 785 | TERI University | 1 | 1 | 87 | 255 |
| 785 | Oswaldo Cruz Foundation | 1 | 1 | 1.6K | 2.6K |
| 785 | Butterfly Conservation | 1 | 1 | 26 | 46 |
| 785 | Swedish Institute of Space Physics | 1 | 1 | 86 | 152 |
| 785 | École Nationale Supérieure des Sciences Agronomiques de Bordeaux-Aquitaine | 1 | 1 | 84 | 191 |
| 785 | Bordeaux INP | 1 | 1 | 269 | 616 |
| 793 | British Trust for Ornithology | 1 | 0 | 9 | 25 |
| 793 | California State University, Fullerton | 1 | 0 | 364 | 1.0K |
| 793 | New Mexico Museum of Natural History and Science | 1 | 0 | 56 | 104 |
