| # | Institution | Articles | Citations | R ARank | R CRank |
|---|
| 1 | Emory University | 45 | 4,006 | 271 | 158 |
| 2 | Eötvös Loránd University | 18 | 2,070 | 176 | 33 |
| 3 | University of California | 50 | 2,000 | 2.9K | 2.6K |
| 4 | Georgia State University | 42 | 1,767 | 57 | 82 |
| 5 | University of Georgia | 24 | 1,343 | 726 | 474 |
| 6 | Yale University | 13 | 1,318 | 1.4K | 732 |
| 7 | Kyoto University | 45 | 1,225 | 824 | 738 |
| 8 | Ohio State University | 10 | 1,205 | 3.0K | 995 |
| 9 | Hungarian Academy of Sciences | 10 | 1,173 | 730 | 192 |
| 10 | University of Hawaii | 12 | 914 | 381 | 193 |
| 11 | Consiglio Nazionale delle Ricerche | 8 | 906 | 394 | 115 |
| 12 | University of Lethbridge | 21 | 869 | 36 | 40 |
| 13 | Rutgers University | 16 | 860 | 1.2K | 868 |
| 14 | Indiana University | 32 | 837 | 417 | 686 |
| 15 | University of Arizona | 13 | 814 | 1.6K | 1.0K |
| 16 | University of Maryland | 14 | 809 | 1.9K | 1.3K |
| 17 | University of St Andrews | 18 | 784 | 213 | 174 |
| 18 | York University | 8 | 740 | 948 | 296 |
| 19 | Northwestern University | 3 | 707 | 4.5K | 1.3K |
| 20 | State University of New York | 20 | 679 | 694 | 831 |
| 21 | University of Texas MD Anderson Cancer Center | 6 | 672 | 1.0K | 439 |
| 22 | University of Rennes 1 | 16 | 639 | 151 | 146 |
| 23 | University of Pennsylvania | 7 | 636 | 3.7K | 1.8K |
| 24 | University of Minnesota | 8 | 615 | 4.0K | 2.1K |
| 25 | McMaster University | 23 | 611 | 823 | 1.0K |
| 26 | University of Vienna | 17 | 588 | 832 | 822 |
| 27 | Brock University | 4 | 578 | 575 | 95 |
| 28 | University of Tennessee | 24 | 575 | 509 | 799 |
| 29 | University of Kentucky | 11 | 557 | 1.4K | 891 |
| 30 | Cornell University | 14 | 545 | 1.8K | 1.8K |
| 31 | University of Alberta | 13 | 543 | 2.1K | 1.6K |
| 32 | Harvard University | 13 | 534 | 1.6K | 1.7K |
| 33 | University of Queensland | 10 | 530 | 2.6K | 1.8K |
| 34 | Brandeis University | 6 | 521 | 456 | 260 |
| 35 | University of Padua | 12 | 520 | 1.6K | 1.1K |
| 36 | Kwansei Gakuin University | 2 | 518 | 426 | 43 |
| 37 | University of Oxford | 4 | 504 | 5.4K | 2.2K |
| 38 | University of Memphis | 8 | 501 | 371 | 189 |
| 39 | University of Louisiana at Lafayette | 2 | 480 | 636 | 49 |
| 40 | University of Zurich | 12 | 477 | 1.2K | 1.2K |
| 41 | University of Stirling | 16 | 461 | 170 | 256 |
| 42 | University of Texas at Austin | 15 | 448 | 2.3K | 2.4K |
| 43 | National Institute on Alcohol Abuse and Alcoholism | 2 | 443 | 280 | 128 |
| 44 | University of Nebraska–Lincoln | 6 | 430 | 986 | 493 |
| 45 | The Rockefeller University | 7 | 418 | 440 | 433 |
| 46 | Columbia University | 10 | 408 | 2.7K | 2.3K |
| 47 | Glendon College | 2 | 405 | 33 | 5 |
| 48 | Concordia University | 1 | 381 | 3.5K | 457 |
| 49 | Northern Arizona University | 2 | 352 | 927 | 192 |
| 50 | University of Pisa | 9 | 308 | 1.4K | 1.2K |
| 51 | Duke University | 9 | 301 | 1.7K | 1.7K |
| 52 | University of California, Los Angeles | 2 | 298 | 4.1K | 1.6K |
| 53 | University of Florida | 13 | 295 | 2.5K | 2.8K |
| 54 | University at Buffalo | 11 | 293 | 1.2K | 1.4K |
| 55 | Washington State University | 4 | 284 | 2.3K | 1.3K |
| 56 | University of Trento | 10 | 274 | 361 | 439 |
| 57 | Konrad Lorenz Institute for Evolution and Cognition Research | 4 | 269 | 4 | 8 |
| 57 | San Francisco State University | 2 | 269 | 930 | 234 |
| 57 | Berry College | 6 | 269 | 20 | 11 |
| 60 | University of Edinburgh | 7 | 268 | 2.8K | 2.2K |
| 61 | University of Wisconsin–Madison | 9 | 262 | 3.6K | 3.4K |
| 62 | Virginia Tech | 10 | 252 | 1.4K | 1.6K |
| 63 | University of Western Australia | 3 | 249 | 3.3K | 1.7K |
| 64 | Arizona State University | 4 | 248 | 3.4K | 1.9K |
| 64 | University of Guelph | 1 | 248 | 3.9K | 1.1K |
| 66 | Laval University | 3 | 247 | 3.0K | 1.5K |
| 67 | University of Liverpool | 4 | 241 | 3.1K | 1.7K |
| 68 | University of Michigan | 6 | 240 | 5.2K | 4.0K |
| 69 | University of Toronto | 6 | 236 | 5.8K | 4.3K |
| 70 | University of Tromsø – The Arctic University of Norway | 5 | 234 | 714 | 558 |
| 71 | Primate Research Institute | 10 | 230 | 9 | 22 |
| 72 | City University of New York | 6 | 229 | 1.4K | 1.0K |
| 72 | National Eye Institute | 1 | 229 | 413 | 183 |
| 72 | National Institute of Mental Health | 1 | 229 | 1.4K | 558 |
| 75 | University of Washington | 5 | 224 | 5.3K | 4.0K |
| 76 | University of the Witwatersrand | 2 | 222 | 2.9K | 779 |
| 76 | Hunter College | 5 | 222 | 317 | 208 |
| 78 | Eunice Kennedy Shriver National Institute of Child Health and Human Development | 8 | 220 | 358 | 564 |
| 79 | Nagoya University | 3 | 219 | 2.9K | 1.3K |
| 80 | Central Washington University | 2 | 214 | 191 | 57 |
| 81 | University of California, Santa Cruz | 1 | 209 | 1.1K | 366 |
| 82 | Georgia Institute of Technology | 5 | 206 | 2.1K | 1.7K |
| 83 | Williams College | 2 | 187 | 357 | 149 |
| 84 | Polish Academy of Sciences | 3 | 186 | 2.9K | 1.5K |
| 85 | University of New England | 4 | 182 | 677 | 465 |
| 86 | University of Chicago | 4 | 179 | 3.6K | 2.7K |
| 87 | Zoo Atlanta | 4 | 176 | 3 | 5 |
| 88 | University of Texas Medical Branch | 3 | 175 | 1.7K | 1.1K |
| 89 | University of Cambridge | 8 | 174 | 3.3K | 3.6K |
| 90 | Sapienza University of Rome | 2 | 172 | 4.8K | 2.1K |
| 91 | University of Missouri | 6 | 169 | 2.5K | 2.3K |
| 92 | Istituto Superiore di Sanità | 3 | 166 | 937 | 673 |
| 93 | Indiana University Bloomington | 7 | 164 | 246 | 501 |
| 94 | Johns Hopkins University | 2 | 163 | 5.7K | 3.1K |
| 95 | Colgate University | 5 | 161 | 126 | 124 |
| 96 | Purdue University | 3 | 159 | 4.7K | 3.0K |
| 97 | University of Tasmania | 2 | 158 | 2.4K | 1.1K |
| 98 | Carleton College | 5 | 153 | 56 | 52 |
| 99 | Natural History Museum | 5 | 151 | 483 | 474 |
| 100 | Bucknell University | 8 | 149 | 77 | 142 |
| 101 | New York State Psychiatric Institute | 2 | 148 | 524 | 392 |
| 102 | California State University | 3 | 147 | 3.0K | 1.5K |
| 103 | Stony Brook University | 1 | 145 | 4.3K | 1.8K |
| 104 | Charles University | 5 | 144 | 1.9K | 1.5K |
| 104 | Central European University | 3 | 144 | 148 | 79 |
| 106 | Boston University | 3 | 142 | 3.2K | 2.3K |
| 107 | University of Toledo | 5 | 141 | 666 | 693 |
| 108 | Hiram College | 3 | 135 | 17 | 12 |
| 109 | DePaul University | 1 | 134 | 1.4K | 329 |
| 110 | Durham University | 3 | 131 | 2.2K | 1.5K |
| 111 | University College London | 5 | 129 | 4.1K | 3.8K |
| 112 | University of Seville | 2 | 126 | 3.0K | 1.3K |
| 112 | Woods Hole Oceanographic Institution | 1 | 126 | 640 | 387 |
| 114 | University of Konstanz | 2 | 123 | 1.5K | 956 |
| 114 | Bowling Green State University | 4 | 123 | 618 | 526 |
| 116 | Brown University | 3 | 120 | 2.8K | 2.2K |
| 116 | University of Hawaii at Manoa | 2 | 120 | 1.5K | 891 |
| 118 | University of Sussex | 5 | 118 | 1.4K | 1.5K |
| 119 | University of Barcelona | 7 | 117 | 2.1K | 2.6K |
| 120 | University of Exeter | 3 | 116 | 2.4K | 1.8K |
| 121 | Northern Michigan University | 1 | 114 | 227 | 39 |
| 121 | Wright State University | 6 | 114 | 401 | 650 |
| 123 | American University | 3 | 112 | 615 | 457 |
| 124 | Université de Moncton | 3 | 109 | 215 | 150 |
| 125 | Ruhr University Bochum | 1 | 108 | 4.1K | 1.7K |
| 125 | Queen's University Belfast | 4 | 108 | 1.7K | 1.6K |
| 127 | University of Parma | 4 | 105 | 1.2K | 1.2K |
| 128 | Oakland University | 6 | 103 | 269 | 495 |
| 129 | University of Massachusetts Amherst | 1 | 102 | 2.6K | 1.1K |
| 130 | University of Strasbourg | 3 | 101 | 1.2K | 964 |
| 130 | Liverpool John Moores University | 2 | 101 | 1.3K | 776 |
| 132 | University of Sydney | 1 | 100 | 9.3K | 4.7K |
| 132 | University of South Bohemia in České Budějovice | 3 | 100 | 398 | 352 |
| 134 | Florida Atlantic University | 3 | 99 | 1.0K | 827 |
| 135 | Massachusetts Institute of Technology | 1 | 95 | 5.6K | 3.4K |
| 135 | Southern Illinois University Carbondale | 2 | 95 | 2.1K | 1.2K |
| 137 | Tufts University | 4 | 92 | 1.5K | 1.9K |
| 138 | Bielefeld University | 2 | 90 | 1.7K | 1.1K |
| 138 | University of Pittsburgh | 1 | 90 | 8.0K | 4.3K |
| 138 | National Zoological Park | 2 | 90 | 141 | 140 |
| 141 | University of Milan | 2 | 89 | 4.6K | 3.1K |
| 142 | Auburn University | 3 | 88 | 2.2K | 1.7K |
| 142 | Louis Herman | 1 | 88 | 1 | 2 |
| 144 | Université du Québec à Montréal | 5 | 87 | 693 | 1.0K |
| 145 | State University of New York at Fredonia | 2 | 86 | 69 | 40 |
| 145 | University of Texas Health Science Center at Houston | 3 | 86 | 1.3K | 1.3K |
| 147 | Leiden University | 3 | 85 | 2.4K | 2.3K |
| 147 | Franklin & Marshall College | 3 | 85 | 113 | 135 |
| 149 | Lund University | 1 | 84 | 6.9K | 3.8K |
| 150 | University of Tübingen | 1 | 83 | 4.6K | 2.5K |
| 151 | University of British Columbia | 4 | 82 | 5.7K | 5.6K |
| 152 | University of Birmingham | 2 | 81 | 5.5K | 3.8K |
| 152 | Carleton University | 2 | 81 | 2.4K | 1.6K |
| 152 | Eastern Kentucky University | 2 | 81 | 199 | 131 |
| 155 | Pennsylvania State University | 2 | 80 | 7.3K | 5.1K |
| 155 | Santa Fe Institute | 1 | 80 | 153 | 136 |
| 157 | University of Leicester | 4 | 77 | 2.0K | 2.3K |
| 158 | University of Connecticut | 2 | 76 | 4.4K | 3.2K |
| 159 | University of Waterloo | 3 | 75 | 3.3K | 3.0K |
| 159 | Bar-Ilan University | 3 | 75 | 1.6K | 1.5K |
| 161 | Whitman College | 4 | 74 | 38 | 90 |
| 162 | McGill University | 3 | 73 | 5.7K | 5.2K |
| 163 | University of Nottingham | 4 | 72 | 3.5K | 3.9K |
| 163 | Osaka University | 3 | 72 | 3.4K | 2.8K |
| 165 | University of Valencia | 5 | 71 | 1.9K | 2.3K |
| 166 | Niigata University | 1 | 70 | 2.4K | 1.1K |
| 166 | University of North Carolina at Greensboro | 2 | 70 | 1.1K | 846 |
| 166 | MIT Computer Science and Artificial Intelligence Laboratory | 1 | 70 | 89 | 56 |
| 169 | University of Veterinary Medicine Hanover, Foundation | 3 | 67 | 268 | 306 |
| 170 | University of New Mexico | 2 | 65 | 3.3K | 2.7K |
| 170 | National Institute of Amazonian Research | 1 | 65 | 402 | 216 |
| 170 | Federal University of Rio Grande do Norte | 1 | 65 | 2.5K | 743 |
| 170 | University of Wisconsin School of Medicine and Public Health | 1 | 65 | 998 | 728 |
| 170 | Zoological Society of San Diego | 2 | 65 | 66 | 84 |
| 175 | Radboud University Nijmegen | 1 | 64 | 4.3K | 2.8K |
| 175 | Vassar College | 3 | 64 | 170 | 193 |
| 177 | Dalhousie University | 2 | 63 | 3.6K | 3.0K |
| 178 | University of Veterinary Medicine Vienna | 5 | 61 | 161 | 327 |
| 178 | College of the Holy Cross | 3 | 61 | 122 | 161 |
| 178 | Lincoln Park Zoo | 1 | 61 | 19 | 17 |
| 181 | University of Central Oklahoma | 2 | 60 | 94 | 89 |
| 182 | Benemérita Universidad Autónoma de Puebla | 1 | 58 | 851 | 275 |
| 183 | San Diego State University | 1 | 57 | 3.2K | 2.0K |
| 184 | Appalachian State University | 1 | 56 | 1.2K | 534 |
| 184 | Agnes Scott College | 2 | 56 | 49 | 41 |
| 186 | Ludwig Maximilian University of Munich | 2 | 55 | 4.2K | 3.7K |
| 186 | University of Westminster | 1 | 55 | 1.1K | 610 |
| 188 | California State University San Marcos | 1 | 54 | 195 | 135 |
| 188 | Yeshiva University | 1 | 54 | 1.3K | 977 |
| 188 | Heriot-Watt University | 1 | 54 | 2.1K | 1.3K |
| 188 | Albert Einstein College of Medicine | 1 | 54 | 2.8K | 2.0K |
| 192 | University of Rochester | 2 | 53 | 3.5K | 3.2K |
| 193 | Clemson University | 1 | 52 | 3.7K | 2.2K |
| 193 | Chiba University | 6 | 52 | 1.3K | 1.9K |
| 195 | University of Bristol | 3 | 51 | 3.9K | 4.2K |
| 195 | Monterey Bay Aquarium Research Institute | 1 | 51 | 160 | 149 |
| 197 | University of Massachusetts | 1 | 50 | 5.6K | 3.8K |
| 198 | National Center for Scientific Research | 6 | 48 | 4.7K | 6.0K |
| 198 | Eindhoven University of Technology | 1 | 48 | 2.7K | 1.9K |
| 200 | California State University, San Bernardino | 1 | 47 | 203 | 127 |
| 201 | Université de Montréal | 1 | 46 | 6.2K | 4.2K |
| 201 | University of Chester | 1 | 46 | 532 | 282 |
| 201 | Binghamton University | 2 | 46 | 1.1K | 1.1K |
| 201 | University of North Carolina | 3 | 46 | 3.9K | 4.7K |
| 201 | Nova Southeastern University | 1 | 46 | 1.1K | 626 |
| 201 | Louis Pasteur University | 1 | 46 | 1.2K | 1.1K |
| 207 | Kansas State University | 2 | 45 | 2.5K | 2.4K |
| 207 | Medical University of Vienna | 3 | 45 | 1.8K | 2.1K |
| 209 | George Washington University | 2 | 44 | 3.1K | 3.0K |
| 209 | University of Miami | 1 | 44 | 5.4K | 3.7K |
| 209 | Keio University | 1 | 44 | 3.8K | 2.3K |
| 209 | University of Florence | 2 | 44 | 3.8K | 3.5K |
| 213 | Macquarie University | 5 | 43 | 1.5K | 2.9K |
| 214 | University of Southern California | 2 | 42 | 6.1K | 5.7K |
| 214 | University of Antwerp | 1 | 42 | 3.9K | 2.8K |
| 214 | Miami University | 2 | 42 | 1.5K | 1.6K |
| 214 | Nanyang Technological University | 1 | 42 | 4.8K | 3.3K |
| 214 | University of York | 1 | 42 | 4.1K | 2.9K |
| 219 | University of Tokyo | 1 | 41 | 8.5K | 5.9K |
| 219 | Walter Reed Army Institute of Research | 3 | 41 | 464 | 810 |
| 221 | University of New Hampshire | 2 | 40 | 1.6K | 1.7K |
| 221 | University of Portsmouth | 3 | 40 | 918 | 1.5K |
| 223 | Utrecht University | 2 | 39 | 4.7K | 4.8K |
| 224 | University of Caen Normandy | 2 | 38 | 570 | 810 |
| 224 | University of Roehampton | 2 | 38 | 401 | 466 |
| 226 | Pace University | 2 | 37 | 261 | 408 |
| 226 | University of Bern | 3 | 37 | 3.0K | 3.8K |
| 228 | University of Haifa | 2 | 36 | 2.1K | 2.1K |
| 228 | University of Southern Mississippi | 2 | 36 | 960 | 1.1K |
| 228 | University of Victoria | 2 | 36 | 2.5K | 2.7K |
| 231 | Northern Illinois University | 3 | 35 | 1.1K | 1.6K |
| 231 | University of Rennes | 3 | 35 | 58 | 289 |
| 233 | University of Denver | 1 | 34 | 1.9K | 1.3K |
| 234 | University of Auckland | 2 | 33 | 4.6K | 4.8K |
| 234 | Bangor University | 1 | 33 | 1.7K | 1.5K |
| 234 | University of Wales | 1 | 33 | 2.4K | 2.1K |
| 237 | University of Regensburg | 3 | 32 | 1.5K | 2.4K |
| 237 | Florida International University | 3 | 32 | 1.7K | 2.7K |
| 237 | E. R. Johnstone Training and Research Center | 1 | 32 | 3 | 2 |
| 240 | University of Minho | 3 | 31 | 1.4K | 2.2K |
| 240 | Villanova University | 1 | 31 | 1.3K | 963 |
| 240 | Aix-Marseille University | 4 | 31 | 1.2K | 2.3K |
| 240 | Christian Brothers University | 1 | 31 | 27 | 25 |
| 240 | University of South Alabama | 2 | 31 | 948 | 1.3K |
| 245 | University of South Florida | 1 | 30 | 5.8K | 4.5K |
| 245 | Mount Holyoke College | 1 | 30 | 427 | 352 |
| 245 | Budapest University of Technology and Economics | 2 | 30 | 1.2K | 1.2K |
| 245 | Southwestern University | 2 | 30 | 58 | 102 |
| 245 | University of Massachusetts Dartmouth | 1 | 30 | 674 | 588 |
| 245 | Goldsmiths, University of London | 2 | 30 | 284 | 456 |
| 251 | Pompeu Fabra University | 2 | 29 | 1.0K | 1.4K |
| 251 | Aarhus University | 2 | 29 | 4.6K | 5.0K |
| 251 | University of Saskatchewan | 1 | 29 | 4.8K | 3.9K |
| 251 | Kresge Hearing Research Institute | 2 | 29 | 65 | 145 |
| 251 | Museo Nacional de Ciencias Naturales | 2 | 29 | 199 | 292 |
| 256 | Newcastle University | 2 | 28 | 4.0K | 4.4K |
| 256 | Beltsville Agricultural Research Center | 2 | 28 | 301 | 440 |
| 256 | Michigan Medicine | 1 | 28 | 3.0K | 2.7K |
| 259 | University of Helsinki | 1 | 27 | 6.6K | 5.4K |
| 259 | Freie Universität Berlin | 1 | 27 | 4.4K | 3.7K |
| 259 | University of South Carolina | 1 | 27 | 5.4K | 4.5K |
| 259 | Ghent University | 2 | 27 | 5.0K | 5.6K |
| 259 | Paris Descartes University | 1 | 27 | 2.4K | 2.2K |
| 259 | Indian Institutes of Science Education and Research | 1 | 27 | 483 | 407 |
| 259 | Wofford College | 1 | 27 | 44 | 29 |
| 266 | Leipzig University | 3 | 26 | 2.2K | 3.3K |
| 266 | University of Wrocław | 1 | 26 | 1.9K | 1.3K |
| 266 | University of Oregon | 1 | 26 | 3.2K | 2.8K |
| 266 | Linköping University | 2 | 26 | 2.8K | 3.6K |
| 266 | Gettysburg College | 1 | 26 | 304 | 279 |
| 266 | University of Pittsburgh at Johnstown | 1 | 26 | 53 | 39 |
| 266 | National Institute of Neurosciences & Hospital | 2 | 26 | 287 | 464 |
| 273 | University of Utah | 3 | 25 | 4.1K | 5.8K |
| 273 | North Carolina State University | 1 | 25 | 5.8K | 4.9K |
| 273 | Maynooth University | 1 | 25 | 838 | 789 |
| 276 | Complutense University of Madrid | 2 | 24 | 4.3K | 4.3K |
| 276 | University of Ottawa | 2 | 24 | 4.8K | 5.4K |
| 276 | University of Bayreuth | 1 | 24 | 1.9K | 1.7K |
| 276 | University of North Carolina Wilmington | 1 | 24 | 897 | 827 |
| 276 | University of Iowa | 3 | 24 | 4.0K | 5.5K |
| 281 | University of Richmond | 2 | 23 | 362 | 621 |
| 281 | Temple University | 1 | 23 | 4.4K | 3.7K |
| 281 | Instituto de Biología y Medicina Experimental | 1 | 23 | 125 | 173 |
| 284 | University of Illinois at Urbana–Champaign | 3 | 22 | 3.8K | 5.6K |
| 284 | Bowdoin College | 1 | 22 | 470 | 446 |
| 284 | Western University | 3 | 22 | 1.2K | 2.3K |
| 284 | Wildlife Conservation Society | 1 | 22 | 180 | 248 |
| 288 | Mount Allison University | 1 | 21 | 415 | 424 |
| 288 | University of Málaga | 1 | 21 | 2.7K | 2.3K |
| 288 | Reed College | 1 | 21 | 375 | 378 |
| 288 | Memorial University of Newfoundland | 2 | 21 | 2.2K | 3.0K |
| 288 | Istituto Italiano di Tecnologia | 1 | 21 | 845 | 994 |
| 288 | Hyogo College of Medicine | 1 | 21 | 1.2K | 1.0K |
| 288 | Muğla University | 1 | 21 | 762 | 577 |
| 295 | Humboldt University of Berlin | 2 | 20 | 3.4K | 4.3K |
| 295 | Hebrew University of Jerusalem | 1 | 20 | 5.1K | 4.6K |
| 295 | University of Leeds | 1 | 20 | 6.9K | 6.1K |
| 295 | University of New South Wales | 1 | 20 | 8.2K | 7.3K |
| 295 | Massey University | 1 | 20 | 3.0K | 2.9K |
| 295 | (former) Institut National de Recherche pour l'Agriculture, l'Alimentation et l'Environnement | 1 | 20 | 2.4K | 2.6K |
| 295 | Georgia Southern University-Armstrong Campus | 1 | 20 | 70 | 84 |
| 295 | University of California, Berkeley | 2 | 20 | 3.5K | 4.9K |
| 295 | Augustana College | 1 | 20 | 179 | 197 |
| 304 | Brooklyn College | 1 | 19 | 775 | 736 |
| 304 | University of Neuchâtel | 3 | 19 | 509 | 1.2K |
| 304 | University of Warwick | 1 | 19 | 5.2K | 4.8K |
| 304 | University of Colorado Denver | 1 | 19 | 2.7K | 2.9K |
| 304 | Max Planck Institute for the Science of Human History | 1 | 19 | 84 | 114 |
| 304 | University of Toronto Mississauga | 1 | 19 | 675 | 830 |
| 310 | National Autonomous University of Mexico | 2 | 18 | 4.1K | 4.2K |
| 310 | University of Burgundy | 1 | 18 | 1.5K | 1.7K |
| 310 | Avila University | 1 | 18 | 11 | 15 |
| 310 | Randolph College | 1 | 18 | 27 | 33 |
| 314 | University of Aveiro | 1 | 17 | 2.7K | 2.6K |
| 314 | Monash University | 1 | 17 | 8.0K | 7.4K |
| 314 | Pontifícia Universidade Católica do Rio Grande do Sul | 1 | 17 | 1.2K | 992 |
| 314 | Oklahoma State University–Stillwater | 1 | 17 | 218 | 278 |
| 314 | Victoria University of Wellington | 2 | 17 | 1.7K | 2.6K |
| 314 | University of Mysore | 1 | 17 | 878 | 811 |
| 314 | Kent State University | 1 | 17 | 2.7K | 2.7K |
| 314 | New College of Florida | 1 | 17 | 102 | 143 |
| 322 | Oregon Health & Science University | 1 | 16 | 2.8K | 3.0K |
| 322 | Nottingham Trent University | 2 | 16 | 1.2K | 2.1K |
| 322 | University of Wisconsin–Oshkosh | 1 | 16 | 480 | 530 |
| 325 | Collège de France | 1 | 15 | 609 | 845 |
| 325 | University of Liège | 1 | 15 | 3.9K | 4.1K |
| 325 | University of Alabama in Huntsville | 1 | 15 | 917 | 1.0K |
| 328 | Michigan State University | 2 | 14 | 6.1K | 7.7K |
| 328 | Griffith University | 1 | 14 | 4.7K | 4.9K |
| 328 | University of Porto | 1 | 14 | 3.5K | 3.8K |
| 328 | Thomas Jefferson University | 1 | 14 | 3.1K | 3.2K |
| 332 | National Tsing Hua University | 1 | 13 | 2.4K | 2.4K |
| 332 | Oregon State University | 2 | 13 | 3.2K | 4.8K |
| 332 | Colorado State University | 2 | 13 | 3.5K | 5.1K |
| 332 | Universidad Nacional del Sur | 1 | 13 | 770 | 765 |
| 332 | Academia Sinica | 1 | 13 | 3.5K | 3.6K |
| 332 | National Scientific and Technical Research Council | 1 | 13 | 2.5K | 2.4K |
| 338 | Amherst College | 1 | 12 | 478 | 610 |
| 338 | University of Göttingen | 2 | 12 | 3.1K | 4.8K |
| 338 | James Madison University | 1 | 12 | 1.1K | 1.3K |
| 338 | Jean Monnet University | 1 | 12 | 447 | 675 |
| 338 | Mohammed V University | 1 | 12 | 907 | 907 |
| 338 | State University of New York at Oneonta | 1 | 12 | 87 | 137 |
| 338 | École Normale Supérieure | 1 | 12 | 1.5K | 1.9K |
| 345 | Southern Connecticut State University | 1 | 11 | 331 | 443 |
| 345 | University of Trieste | 2 | 11 | 1.9K | 3.3K |
| 345 | Portland State University | 1 | 11 | 2.0K | 2.5K |
| 345 | Kenyon College | 1 | 11 | 251 | 347 |
| 345 | Drake University | 1 | 11 | 354 | 471 |
| 345 | Graduate Center, CUNY | 2 | 11 | 139 | 468 |
| 345 | Luther College | 1 | 11 | 128 | 202 |
| 345 | University of Virginia | 2 | 11 | 4.7K | 6.6K |
| 353 | University of Kansas | 1 | 10 | 5.7K | 6.1K |
| 353 | University of Texas at Tyler | 1 | 10 | 345 | 524 |
| 353 | University of Waikato | 1 | 10 | 1.8K | 2.4K |
| 353 | University of São Paulo | 1 | 10 | 9.4K | 8.3K |
| 353 | University of Otago | 2 | 10 | 3.3K | 5.2K |
| 353 | George Mason University | 1 | 10 | 3.6K | 4.0K |
| 353 | Loyola University Chicago | 1 | 10 | 2.2K | 2.6K |
| 353 | Hendrix College | 1 | 10 | 99 | 171 |
| 353 | University of Tulsa | 2 | 10 | 542 | 1.2K |
| 353 | Università degli Studi Roma Tre | 1 | 10 | 776 | 1.0K |
| 353 | Oklahoma State University | 2 | 10 | 2.5K | 4.1K |
| 364 | Dartmouth College | 1 | 9 | 2.6K | 3.3K |
| 364 | University of Debrecen | 1 | 9 | 2.2K | 2.5K |
| 364 | Bloomfield College | 1 | 9 | 19 | 30 |
| 364 | University of Newcastle | 1 | 9 | 5.0K | 5.9K |
| 368 | Wichita State University | 1 | 8 | 1.2K | 1.6K |
| 368 | Syracuse University | 1 | 8 | 3.4K | 4.1K |
| 368 | SeaWorld Parks & Entertainment | 1 | 8 | 10 | 14 |
| 368 | National University of Colombia | 2 | 8 | 1.5K | 2.5K |
| 368 | University of New Orleans | 1 | 8 | 1.2K | 1.7K |
| 373 | Saint Petersburg State University | 1 | 7 | 1.5K | 1.5K |
| 373 | West Virginia University | 1 | 7 | 4.3K | 5.3K |
| 373 | Tilburg University | 1 | 7 | 1.9K | 2.5K |
| 373 | Koç University | 1 | 7 | 1.3K | 1.8K |
| 373 | Université Catholique de Lille | 1 | 7 | 117 | 261 |
| 373 | Scientific and Technological Research Council of Turkey | 1 | 7 | 161 | 321 |
| 373 | Hubert Curien Multi-disciplinary Institute | 1 | 7 | 164 | 282 |
| 373 | California University of Pennsylvania | 1 | 7 | 307 | 376 |
| 373 | Universidad CEU Cardenal Herrera | 1 | 7 | 93 | 187 |
| 382 | University of Canterbury | 1 | 6 | 2.9K | 3.9K |
| 382 | Universidad de Guanajuato | 1 | 6 | 946 | 1.2K |
| 382 | National Chengchi University | 1 | 6 | 673 | 997 |
| 382 | Cardiff University | 2 | 6 | 3.8K | 6.4K |
| 382 | Shiga University | 1 | 6 | 183 | 282 |
| 382 | Eastern Michigan University | 1 | 6 | 881 | 1.4K |
| 382 | University of Alaska Anchorage | 1 | 6 | 431 | 795 |
| 382 | Louisiana State University | 1 | 6 | 5.3K | 6.4K |
| 382 | Universidad Veracruzana | 1 | 6 | 965 | 1.1K |
| 382 | Mississippi State University | 1 | 6 | 2.8K | 3.7K |
| 382 | University of Udine | 1 | 6 | 2.0K | 2.9K |
| 382 | Tamagawa University | 1 | 6 | 235 | 308 |
| 394 | University of Turin | 1 | 5 | 5.5K | 6.7K |
| 394 | University of Illinois at Springfield | 1 | 5 | 168 | 372 |
| 394 | Grenoble University - Grenoble Alpes University | 1 | 5 | 1.3K | 2.0K |
| 394 | University of Wyoming | 1 | 5 | 2.6K | 3.6K |
| 394 | New Mexico State University | 1 | 5 | 2.2K | 3.1K |
| 394 | University of North Florida | 1 | 5 | 805 | 1.3K |
| 394 | University of Abertay Dundee | 1 | 5 | 338 | 693 |
| 394 | Udayana University | 1 | 5 | 626 | 506 |
| 394 | Idaho State University | 1 | 5 | 962 | 1.6K |
| 394 | University of Texas at Dallas | 1 | 5 | 2.5K | 3.5K |
| 394 | University of Girona | 1 | 5 | 1.4K | 2.1K |
| 394 | Hope College | 1 | 5 | 292 | 562 |
| 394 | United States Department of Homeland Security | 1 | 5 | 4 | 12 |
| 407 | University of Notre Dame | 1 | 4 | 3.9K | 5.2K |
| 407 | University of North Carolina at Chapel Hill | 1 | 4 | 5.8K | 7.4K |
| 407 | University of Amsterdam | 1 | 4 | 6.8K | 8.3K |
| 407 | Central Queensland University | 1 | 4 | 1.0K | 1.8K |
| 407 | Weber State University | 1 | 4 | 319 | 646 |
| 407 | University of Notre Dame Australia | 1 | 4 | 1.4K | 2.4K |
| 407 | Rochester Institute of Technology | 1 | 4 | 1.2K | 2.0K |
| 407 | Texas Christian University | 1 | 4 | 1.2K | 1.8K |
| 407 | Autonomous University of Tlaxcala | 1 | 4 | 166 | 290 |
| 407 | Gordon–Conwell Theological Seminary | 1 | 4 | 5 | 8 |
| 407 | Universidad de la República | 1 | 4 | 1.8K | 2.5K |
| 418 | Australian National University | 1 | 3 | 5.6K | 7.4K |
| 418 | National Institutes of Health | 1 | 3 | 4.9K | 6.1K |
| 418 | Tours University | 1 | 3 | 826 | 1.5K |
| 418 | Kindai University | 1 | 3 | 2.1K | 2.8K |
| 418 | Doshisha University | 1 | 3 | 997 | 1.5K |
| 418 | University of Manitoba | 1 | 3 | 5.6K | 7.4K |
| 418 | University of Hull | 1 | 3 | 2.7K | 4.0K |
| 418 | Novosibirsk State University | 1 | 3 | 1.1K | 1.6K |
| 418 | Wilfrid Laurier University | 1 | 3 | 1.5K | 2.5K |
| 418 | University of Brighton | 1 | 3 | 1.3K | 2.3K |
| 428 | University of Cincinnati | 1 | 2 | 5.3K | 7.2K |
| 428 | University of Melbourne | 1 | 2 | 9.1K | 11.4K |
| 428 | University of Zaragoza | 1 | 2 | 3.4K | 4.7K |
| 428 | University of Lausanne | 1 | 2 | 3.3K | 5.0K |
| 428 | Katholieke Universiteit Leuven | 1 | 2 | 7.2K | 9.5K |
| 428 | University of Colorado | 1 | 2 | 6.2K | 8.4K |
| 428 | University of Louisville | 1 | 2 | 4.1K | 6.1K |
| 428 | Konrad Lorenz Institute of Ethology | 1 | 2 | 36 | 78 |
| 428 | Northeastern Illinois University | 1 | 2 | 288 | 713 |
| 428 | Birmingham City University | 1 | 2 | 595 | 1.2K |
| 428 | Arcadia University | 1 | 2 | 162 | 416 |
| 428 | Piedmont University | 1 | 2 | 20 | 51 |
| 440 | Princeton University | 1 | 1 | 4.5K | 6.3K |
| 440 | Boston College | 1 | 1 | 2.3K | 3.6K |
| 440 | Smith College | 2 | 1 | 439 | 1.4K |
| 440 | Senshu University | 1 | 1 | 161 | 344 |
| 440 | Universidad Nacional de Cuyo | 1 | 1 | 465 | 947 |
