| # | Title | Journal | Year | Citations |
|---|
| 1 | Terrestrial Ecoregions of the World: A New Map of Life on Earth | BioScience | 2001 | 6,067 |
| 2 | The Natural Flow Regime | BioScience | 1997 | 5,166 |
| 3 | Marine Ecoregions of the World: A Bioregionalization of Coastal and Shelf Areas | BioScience | 2007 | 2,773 |
| 4 | Urbanization, Biodiversity, and Conservation | BioScience | 2002 | 2,618 |
| 5 | Quantifying Threats to Imperiled Species in the United States | BioScience | 1998 | 2,421 |
| 6 | A Global Crisis for Seagrass Ecosystems | BioScience | 2006 | 2,318 |
| 7 | The Nitrogen Cascade | BioScience | 2003 | 2,278 |
| 8 | Cascading Trophic Interactions and Lake Productivity | BioScience | 1985 | 2,183 |
| 9 | Proximate Causes and Underlying Driving Forces of Tropical Deforestation | BioScience | 2002 | 2,155 |
| 10 | Environmental and Economic Costs of Nonindigenous Species in the United States | BioScience | 2000 | 2,144 |
| 11 | Nitrogen Saturation in Northern Forest Ecosystems | BioScience | 1989 | 2,074 |
| 12 | Carbon Isotopes in Photosynthesis | BioScience | 1988 | 1,951 |
| 13 | Log-normal Distributions across the Sciences: Keys and Clues | BioScience | 2001 | 1,939 |
| 14 | The Identification, Conservation, and Management of Estuarine and Marine Nurseries for Fish and Invertebrates | BioScience | 2001 | 1,934 |
| 15 | An Ecosystem Perspective of Riparian Zones | BioScience | 1991 | 1,898 |
| 16 | The Human Footprint and the Last of the Wild | BioScience | 2002 | 1,852 |
| 17 | A Continuous Satellite-Derived Measure of Global Terrestrial Primary Production | BioScience | 2004 | 1,778 |
| 18 | Climate Change and Forest Disturbances | BioScience | 2001 | 1,682 |
| 19 | Citizen Science: A Developing Tool for Expanding Science Knowledge and Scientific Literacy | BioScience | 2009 | 1,681 |
| 20 | Nitrogen Saturation in Temperate Forest Ecosystems | BioScience | 1998 | 1,630 |
| 21 | Minimum Population Sizes for Species Conservation | BioScience | 1981 | 1,607 |
| 22 | Challenges in the Quest for Keystones | BioScience | 1996 | 1,557 |
| 23 | Freshwater Ecoregions of the World: A New Map of Biogeographic Units for Freshwater Biodiversity Conservation | BioScience | 2008 | 1,483 |
| 24 | Nonlethal Effects in the Ecology of Predator-Prey Interactions | BioScience | 1998 | 1,477 |
| 25 | Cross-scale Drivers of Natural Disturbances Prone to Anthropogenic Amplification: The Dynamics of Bark Beetle Eruptions | BioScience | 2008 | 1,410 |
| 26 | Vulnerability of Permafrost Carbon to Climate Change: Implications for the Global Carbon Cycle | BioScience | 2008 | 1,379 |
| 27 | Lidar Remote Sensing for Ecosystem Studies | BioScience | 2002 | 1,330 |
| 28 | Mangrove Forests: One of the World's Threatened Major Tropical Environments | BioScience | 2001 | 1,320 |
| 29 | The Economic Value of Ecological Services Provided by Insects | BioScience | 2006 | 1,316 |
| 30 | Human Appropriation of the Products of Photosynthesis | BioScience | 1986 | 1,301 |
| 31 | The Global Decline of Reptiles, Déjà Vu Amphibians | BioScience | 2000 | 1,212 |
| 32 | Landscapes to Riverscapes: Bridging the Gap between Research and Conservation of Stream Fishes | BioScience | 2002 | 1,193 |
| 33 | Effects of Invasive Alien Plants on Fire Regimes | BioScience | 2004 | 1,193 |
| 34 | An Ecoregion-Based Approach to Protecting Half the Terrestrial Realm | BioScience | 2017 | 1,178 |
| 35 | What Is Conservation Biology? | BioScience | 1985 | 1,146 |
| 36 | Plant Responses to Multiple Environmental Factors | BioScience | 1987 | 1,109 |
| 37 | The Ocean's Food Web, A Changing Paradigm | BioScience | 1974 | 1,104 |
| 38 | Structure and Function of Stream Ecosystems | BioScience | 1974 | 1,093 |
| 39 | Progressive Nitrogen Limitation of Ecosystem Responses to Rising Atmospheric Carbon Dioxide | BioScience | 2004 | 1,092 |
| 40 | The Evolutionary Significance of Phenotypic Plasticity | BioScience | 1989 | 1,068 |
| 41 | Prairie Conservation in North America | BioScience | 1994 | 1,064 |
| 42 | Resilience in Agriculture through Crop Diversification: Adaptive Management for Environmental Change | BioScience | 2011 | 1,040 |
| 43 | Climate Change and Bark Beetles of the Western United States and Canada: Direct and Indirect Effects | BioScience | 2010 | 1,009 |
| 44 | Viruses and Nutrient Cycles in the Sea | BioScience | 1999 | 996 |
| 45 | Oyster Reefs at Risk and Recommendations for Conservation, Restoration, and Management | BioScience | 2011 | 978 |
| 46 | Hydraulic Limits to Tree Height and Tree Growth | BioScience | 1997 | 974 |
| 47 | Reef Corals: Mutualistic Symbioses Adapted to Nutrient-Poor Environments | BioScience | 1977 | 968 |
| 48 | Consequences of More Extreme Precipitation Regimes for Terrestrial Ecosystems | BioScience | 2008 | 959 |
| 49 | The Demise of Fire and “Mesophication” of Forests in the Eastern United States | BioScience | 2008 | 959 |
| 50 | The Keystone-Species Concept in Ecology and Conservation | BioScience | 1993 | 955 |
