| # | Title | Journal | Year | Citations |
|---|
| 1 | Community structure in social and biological networks | Proceedings of the National Academy of Sciences of the United States of America | 2002 | 11,888 |
| 2 | Finding and evaluating community structure in networks | Physical Review E | 2004 | 9,503 |
| 3 | Swarm Intelligence | | 1999 | 4,417 |
| 4 | The structure of scientific collaboration networks | Proceedings of the National Academy of Sciences of the United States of America | 2001 | 3,277 |
| 5 | Random graphs with arbitrary degree distributions and their applications | Physical Review E | 2001 | 2,651 |
| 6 | Network Robustness and Fragility: Percolation on Random Graphs | Physical Review Letters | 2000 | 1,970 |
| 7 | Embedology | Journal of Statistical Physics | 1991 | 1,895 |
| 8 | Fast folding and comparison of RNA secondary structures | Monatshefte Für Chemie | 1994 | 1,862 |
| 9 | Scientific collaboration networks. II. Shortest paths, weighted networks, and centrality | Physical Review E | 2001 | 1,763 |
| 10 | Hierarchical structure and the prediction of missing links in networks | Nature | 2008 | 1,674 |
| 11 | Inferring friendship network structure by using mobile phone data | Proceedings of the National Academy of Sciences of the United States of America | 2009 | 1,516 |
| 12 | Scientific collaboration networks. I. Network construction and fundamental results | Physical Review E | 2001 | 1,474 |
| 13 | Clustering and preferential attachment in growing networks | Physical Review E | 2001 | 1,365 |
| 14 | Renormalization group analysis of the small-world network model | Physics Letters, Section A: General, Atomic and Solid State Physics | 1999 | 1,198 |
| 15 | The structure of scientific collaboration networks | Proceedings of the National Academy of Sciences of the United States of America | 2001 | 1,186 |
| 16 | A framework for the unification of the behavioral sciences | Behavioral and Brain Sciences | 2007 | 1,089 |
| 17 | Ecological networks and their fragility | Nature | 2006 | 1,064 |
| 18 | The Origins of Scaling in Cities | Science | 2013 | 1,001 |
| 19 | Scaling and percolation in the small-world network model | Physical Review E | 1999 | 950 |
| 20 | Random graph models of social networks | Proceedings of the National Academy of Sciences of the United States of America | 2002 | 946 |
| 21 | The economy needs agent-based modelling | Nature | 2009 | 943 |
| 22 | Costly Signaling and Cooperation | Journal of Theoretical Biology | 2001 | 867 |
| 23 | Stochasticity in Transcriptional Regulation: Origins, Consequences, and Mathematical Representations | Biophysical Journal | 2001 | 840 |
| 24 | Inspiration for optimization from social insect behaviour | Nature | 2000 | 811 |
| 25 | Identity and Search in Social Networks | Science | 2002 | 804 |
| 26 | Effects of size and temperature on developmental time | Nature | 2002 | 798 |
| 27 | Dominant influence of HLA-B in mediating the potential co-evolution of HIV and HLA | Nature | 2004 | 784 |
| 28 | Effects of Body Size and Temperature on Population Growth | American Naturalist | 2004 | 767 |
| 29 | HIV-1 Nomenclature Proposal | Science | 2000 | 756 |
| 30 | Epidemics and percolation in small-world networks | Physical Review E | 2000 | 741 |
| 31 | A unified theory of urban living | Nature | 2010 | 728 |
| 32 | Parasites in food webs: the ultimate missing links | Ecology Letters | 2008 | 716 |
| 33 | The Structure and Dynamics of Networks | | 2011 | 707 |
| 34 | Performance of modularity maximization in practical contexts | Physical Review E | 2010 | 698 |
| 35 | From sequences to shapes and back: a case study in RNA secondary structures | Proceedings of the Royal Society B: Biological Sciences | 1994 | 685 |
| 36 | Complexity and the Economy | Science | 1999 | 685 |
| 37 | Language Is a Complex Adaptive System: Position Paper | Language Learning | 2009 | 678 |
| 38 | Increased temperature variation poses a greater risk to species than climate warming | Proceedings of the Royal Society B: Biological Sciences | 2014 | 674 |
| 39 | Architecture for an Artificial Immune System | Evolutionary Computation | 2000 | 645 |
| 40 | Did Warfare Among Ancestral Hunter-Gatherers Affect the Evolution of Human Social Behaviors? | Science | 2009 | 638 |
| 41 | The evolution of strong reciprocity: cooperation in heterogeneous populations | Theoretical Population Biology | 2004 | 636 |
| 42 | Policies Designed for Self-Interested Citizens May Undermine "The Moral Sentiments": Evidence from Economic Experiments | Science | 2008 | 610 |
| 43 | Time series properties of an artificial stock market | Journal of Economic Dynamics and Control | 1999 | 574 |
| 44 | The origin and evolution of cell types | Nature Reviews Genetics | 2016 | 572 |
| 45 | Email networks and the spread of computer viruses | Physical Review E | 2002 | 559 |
| 46 | Network Diversity and Economic Development | Science | 2010 | 559 |
| 47 | Policing stabilizes construction of social niches in primates | Nature | 2006 | 545 |
| 48 | Models of the Small World | Journal of Statistical Physics | 2000 | 544 |
| 49 | Global ecosystem thresholds driven by aridity | Science | 2020 | 526 |
| 50 | Food webs: reconciling the structure and function of biodiversity | Trends in Ecology and Evolution | 2012 | 521 |
