| 1 | Dartmouth College | 12 | 945 | 409 | 279 |
| 2 | California Institute of Technology | 14 | 477 | 839 | 989 |
| 3 | University of California | 12 | 462 | 7706 | 5979 |
| 4 | Case Western Reserve University | 4 | 403 | 3405 | 1593 |
| 5 | Albert Einstein College of Medicine | 10 | 366 | 1141 | 1089 |
| 6 | Marine Biological Laboratory | 7 | 355 | 210 | 216 |
| 7 | Baylor College of Medicine | 5 | 328 | 2359 | 1497 |
| 8 | University of Hawaii | 5 | 278 | 1174 | 843 |
| 9 | University of Pennsylvania | 4 | 261 | 5996 | 3650 |
| 10 | University of North Carolina at Chapel Hill | 5 | 256 | 4500 | 3235 |
| 10 | University of North Carolina | 5 | 256 | 4782 | 3364 |
| 10 | Worcester Foundation for Biomedical Research | 4 | 256 | 84 | 68 |
| 13 | Colorado State University | 1 | 239 | 5079 | 1589 |
| 14 | Princeton University | 2 | 225 | 5031 | 2390 |
| 15 | Johns Hopkins School of Medicine | 3 | 221 | 1949 | 1263 |
| 16 | Harvard University | 8 | 213 | 5778 | 5483 |
| 16 | University of Washington | 4 | 213 | 6378 | 4420 |
| 18 | The Rockefeller University | 4 | 183 | 746 | 750 |
| 19 | Stanford University | 2 | 180 | 8467 | 4847 |
| 19 | Carnegie Mellon University | 3 | 180 | 2390 | 1654 |
| 21 | University of Auckland | 1 | 175 | 6912 | 2498 |
| 22 | National Institute for Basic Biology, Japan | 2 | 153 | 258 | 193 |
| 23 | Purdue University | 2 | 145 | 6873 | 3712 |
| 23 | University at Buffalo | 9 | 145 | 1619 | 2476 |
| 23 | Koninklijke Philips NV | 1 | 145 | 1831 | 705 |
| 26 | State University of New York | 11 | 133 | 2098 | 3620 |
| 27 | Nagoya University | 3 | 132 | 3795 | 2460 |
| 28 | Scripps Research | 1 | 129 | 1549 | 962 |
| 29 | University of Siena | 2 | 125 | 2129 | 1274 |
| 30 | University of Alabama | 1 | 120 | 5487 | 2253 |
| 31 | University of Bern | 4 | 109 | 3102 | 2903 |
| 32 | University of Virginia | 4 | 105 | 3635 | 3415 |
| 33 | Duke University | 3 | 99 | 4895 | 4087 |
| 34 | Florida State University | 3 | 97 | 3124 | 2549 |
| 35 | University of California, San Diego | 2 | 94 | 4161 | 3129 |
| 36 | Cancer Research Center | 2 | 88 | 2680 | 2089 |
| 37 | Tokyo Institute of Technology | 3 | 87 | 2457 | 1868 |
| 37 | Leonard M. Miller School of Medicine | 3 | 87 | 2094 | 1914 |
| 39 | Rice University | 1 | 82 | 3437 | 1970 |
| 40 | Ochanomizu University | 2 | 81 | 479 | 293 |
| 41 | Yale University | 6 | 78 | 3816 | 4965 |
| 42 | Smith College | 1 | 77 | 892 | 424 |
| 42 | University of Maryland | 4 | 77 | 6018 | 6028 |
| 42 | University at Albany | 1 | 77 | 2263 | 1272 |
| 45 | Fordham University | 2 | 76 | 1140 | 789 |
| 46 | Cold Spring Harbor Laboratory | 2 | 75 | 300 | 389 |
| 47 | Katholieke Universiteit Leuven | 1 | 74 | 8624 | 4969 |
| 47 | Janssen Pharmaceuticals | 1 | 74 | 844 | 559 |
| 47 | interdisciplinarity | 1 | 74 | 10684 | 5548 |
| 47 | Medical Research Council | 2 | 74 | 2069 | 1938 |
| 51 | University of Tokyo | 3 | 73 | 6733 | 5401 |
| 52 | University of Kansas | 2 | 71 | 4053 | 3074 |
| 53 | Rensselaer Polytechnic Institute | 1 | 67 | 2485 | 1589 |
| 53 | Syracuse University | 4 | 67 | 1572 | 2011 |
| 55 | Osaka University | 2 | 62 | 5116 | 3733 |
| 55 | University of New Mexico | 2 | 62 | 4016 | 3438 |
| 57 | Otsuka Pharmaceutical | 1 | 61 | 1018 | 635 |
| 58 | University of Toronto | 1 | 57 | 12429 | 8009 |
| 58 | Alexandria University | 1 | 57 | 5792 | 3057 |
| 60 | University of Vermont | 1 | 55 | 3713 | 2374 |
| 61 | George Washington University | 1 | 53 | 5656 | 3569 |
| 61 | Montana State University | 1 | 53 | 2349 | 1609 |
| 61 | University Medical Center | 1 | 53 | 7963 | 5538 |
| 64 | Salk Institute for Biological Studies | 1 | 50 | 772 | 735 |
| 65 | Kansas State University | 1 | 49 | 3762 | 2494 |
| 65 | Goethe University Frankfurt | 1 | 49 | 4411 | 2926 |
| 67 | Wayne State University | 1 | 48 | 5966 | 4066 |
| 68 | Haverford College | 2 | 47 | 199 | 269 |
| 69 | Pennsylvania State University | 2 | 45 | 7958 | 6964 |
| 70 | Laboratory of Molecular Biology | 4 | 44 | 919 | 1525 |
| 70 | Hadassah Medical Center | 1 | 44 | 1949 | 1539 |
| 72 | Centre for Blood Research | 1 | 42 | 248 | 272 |
| 73 | University of Texas System | 2 | 41 | 1994 | 2213 |
| 74 | Weizmann Institute of Science | 2 | 34 | 1968 | 2313 |
| 74 | Icahn School of Medicine at Mount Sinai | 1 | 34 | 4110 | 3357 |
| 74 | SUNY Downstate Medical Center | 2 | 34 | 1276 | 1399 |
| 74 | The Mount | 1 | 34 | 943 | 918 |
| 78 | Johns Hopkins University | 3 | 33 | 5759 | 6737 |
| 79 | Oregon State University | 3 | 32 | 2804 | 3994 |
| 79 | Bio-Medical Science (South Korean) | 2 | 32 | 6712 | 6155 |
| 81 | University of Rochester Medical Center | 1 | 29 | 2286 | 2059 |
| 82 | University of Bristol | 1 | 28 | 6944 | 5782 |
| 82 | Washington University School of Medicine | 1 | 28 | 3942 | 3537 |
| 84 | Penn State Milton S. Hershey Medical Center | 1 | 26 | 1617 | 1476 |
| 85 | University of Toledo Medical Center | 1 | 25 | 757 | 746 |
| 86 | University of Notre Dame Australia | 1 | 23 | 4988 | 4297 |
| 87 | University of Southampton | 1 | 22 | 6555 | 5944 |
| 87 | Wake Forest University | 1 | 22 | 2609 | 2592 |
| 87 | University of Colorado Boulder | 2 | 22 | 2319 | 3277 |
| 90 | Thomas Jefferson University | 1 | 21 | 4608 | 4214 |
| 90 | Robert Wood Johnson Medical School | 1 | 21 | 2137 | 2107 |
| 92 | Technion – Israel Institute of Technology | 1 | 20 | 5041 | 4746 |
| 92 | University of Wisconsin–Madison | 2 | 20 | 9072 | 9688 |
| 92 | University of Medicine and Dentistry of New Jersey | 1 | 20 | 1502 | 1822 |
| 92 | John Curtin School of Medical Research | 1 | 20 | 650 | 808 |
| 96 | University of Southern California | 2 | 19 | 6611 | 7631 |
| 96 | Lone Star College System | 1 | 19 | 7525 | 7143 |
| 96 | Madras Christian College | 1 | 19 | 3837 | 3459 |
| 99 | Massachusetts Institute of Technology | 1 | 18 | 6245 | 6277 |
| 99 | University of South Carolina | 2 | 18 | 5793 | 6889 |
| 99 | University of Washington Friday Harbor Laboratories | 1 | 18 | 134 | 220 |
| 102 | Los Alamos National Laboratory | 1 | 17 | 2312 | 2434 |
| 102 | Millersville University of Pennsylvania | 1 | 17 | 211 | 241 |
| 102 | Michigan Medicine | 1 | 17 | 3180 | 3270 |
| 105 | University of Denver | 1 | 16 | 2221 | 2227 |
| 105 | Emory University | 2 | 16 | 5178 | 6315 |
| 105 | State University of New York at Albany | 1 | 16 | 1126 | 1326 |
| 105 | state health agency | 1 | 16 | 1170 | 1407 |
| 109 | Bielefeld University | 1 | 14 | 2247 | 2615 |
| 109 | Temple University | 1 | 14 | 5401 | 5390 |
| 111 | University of Minnesota | 1 | 13 | 10588 | 10049 |
| 111 | University of Colorado | 1 | 13 | 7552 | 7589 |
| 113 | University of Salzburg | 1 | 12 | 1828 | 2213 |
| 113 | Queens College | 1 | 12 | 1433 | 1724 |
| 115 | Ben-Gurion University of the Negev | 1 | 11 | 4912 | 5379 |
| 115 | Xavier University | 1 | 11 | 1380 | 1748 |
| 115 | Jacob Blaustein Institutes for Desert Research | 1 | 11 | 420 | 591 |
| 118 | University of Texas at Austin | 1 | 10 | 11438 | 11162 |
| 118 | Ruhr University Bochum | 1 | 10 | 4905 | 5278 |
| 118 | University of Connecticut Health Center | 1 | 10 | 1697 | 2247 |
| 121 | Cornell University | 1 | 9 | 9061 | 9480 |
| 121 | Oakland University | 1 | 9 | 2157 | 2768 |
| 123 | Moscow State University | 1 | 7 | 4925 | 4900 |
| 123 | University of Wollongong | 2 | 7 | 2972 | 5322 |
| 123 | Oglethorpe University | 1 | 7 | 28 | 95 |
| 126 | Mount Sinai Medical Center | 1 | 6 | 1201 | 1667 |
| 127 | Boston University | 1 | 5 | 7209 | 8370 |
| 128 | Russell Sage College | 1 | 4 | 80 | 191 |
| 128 | Duke University Hospital | 1 | 4 | 3602 | 4525 |
| 128 | University of Tennessee | 1 | 4 | 6891 | 8570 |
| 131 | National Institutes of Health | 1 | 3 | 5077 | 6318 |
| 132 | Great Ormond Street Hospital | 1 | 2 | 3385 | 4445 |
| 132 | Warner–Lambert | 1 | 2 | 273 | 456 |
| 134 | West Virginia University | 1 | 1 | 5033 | 7475 |