| 1 | Brandeis University | 2 | 712 | 1182 | 208 |
| 2 | National Institutes of Health | 14 | 474 | 1681 | 1609 |
| 3 | Massachusetts Institute of Technology | 5 | 363 | 3071 | 2014 |
| 4 | Cornell University | 12 | 340 | 2561 | 2672 |
| 5 | Australian National University | 1 | 327 | 6662 | 1501 |
| 5 | John Curtin School of Medical Research | 1 | 327 | 649 | 160 |
| 7 | Baylor College of Medicine | 1 | 282 | 4407 | 1478 |
| 8 | Duke University | 5 | 247 | 4519 | 3022 |
| 9 | University of Wisconsin–Madison | 5 | 223 | 5734 | 3801 |
| 10 | University of California, Berkeley | 3 | 196 | 5568 | 3446 |
| 11 | Ohio State University | 4 | 173 | 6336 | 4154 |
| 12 | Stockholm University | 2 | 170 | 3813 | 1743 |
| 13 | Purdue University | 4 | 164 | 5079 | 3299 |
| 14 | Medical College of Wisconsin | 2 | 163 | 2344 | 1115 |
| 15 | University of Edinburgh | 1 | 131 | 8628 | 3376 |
| 15 | University of Bristol | 1 | 131 | 7050 | 2931 |
| 17 | Michigan State University | 2 | 130 | 6760 | 3572 |
| 18 | Harvard University | 5 | 120 | 7015 | 6163 |
| 19 | University of Oulu | 1 | 116 | 3933 | 1565 |
| 20 | University of Zurich | 1 | 106 | 6574 | 3075 |
| 21 | University of Texas at Austin | 3 | 103 | 8453 | 5966 |
| 22 | National Institute of Dental and Craniofacial Research | 3 | 102 | 331 | 437 |
| 23 | Texas A&M University | 3 | 100 | 6432 | 4495 |
| 24 | Columbia University | 3 | 91 | 6998 | 5331 |
| 25 | National Institute of Mental Health | 2 | 80 | 1041 | 857 |
| 26 | Case Western Reserve University | 2 | 73 | 4840 | 3595 |
| 27 | National Heart, Lung, and Blood Institute | 3 | 72 | 692 | 823 |
| 28 | University of Florida | 3 | 70 | 7169 | 5864 |
| 28 | University of Essex | 1 | 70 | 2735 | 1305 |
| 30 | Osaka University | 1 | 64 | 6419 | 3480 |
| 31 | Pennsylvania State University | 3 | 55 | 6716 | 6139 |
| 32 | Reichman University | 1 | 54 | 2253 | 1394 |
| 33 | University of California, San Diego | 1 | 53 | 8059 | 5427 |
| 34 | Yale University | 5 | 51 | 4271 | 5433 |
| 34 | Humboldt University of Berlin | 2 | 51 | 4988 | 3809 |
| 36 | Université catholique de Louvain | 1 | 49 | 5005 | 3369 |
| 37 | Albert Einstein College of Medicine | 1 | 48 | 3524 | 2494 |
| 38 | Academia Sinica | 1 | 47 | 3789 | 2446 |
| 39 | Iowa State University | 2 | 46 | 4209 | 3681 |
| 40 | North Carolina State University | 1 | 40 | 6314 | 4388 |
| 40 | National Eye Institute | 1 | 40 | 463 | 447 |
| 40 | Fox Chase Cancer Center | 1 | 40 | 986 | 827 |
| 43 | University of Michigan | 2 | 39 | 10011 | 8422 |
| 43 | St. Jude Children's Research Hospital | 1 | 39 | 1348 | 1097 |
| 45 | Emory University | 1 | 38 | 6684 | 4739 |
| 45 | Virginia Commonwealth University | 1 | 38 | 5193 | 3644 |
| 45 | VCU Medical Center | 1 | 38 | 1310 | 1122 |
| 48 | University College Dublin | 1 | 36 | 5154 | 3568 |
| 48 | NOVA University Lisbon | 1 | 36 | 3508 | 2146 |
| 48 | Aix-Marseille University | 1 | 36 | 5436 | 3739 |
| 48 | Centre for Food Safety | 1 | 36 | 82 | 97 |
| 52 | Karolinska Institute | 2 | 32 | 4248 | 4218 |
| 53 | Moscow State University | 2 | 31 | 3782 | 2982 |
| 54 | University of Düsseldorf | 1 | 30 | 3533 | 2814 |
| 54 | University of Miami | 2 | 30 | 4976 | 4993 |
| 54 | Scripps Research | 2 | 30 | 1197 | 1511 |
| 54 | Lawrence Berkeley National Laboratory | 2 | 30 | 1976 | 2339 |
| 58 | Johns Hopkins University | 4 | 28 | 5224 | 6870 |
| 59 | University of Bonn | 1 | 27 | 5197 | 4152 |
| 60 | University of Southern California | 2 | 26 | 6693 | 6762 |
| 61 | University of Pennsylvania | 2 | 25 | 7721 | 7613 |
| 61 | University of London | 1 | 25 | 7307 | 5766 |
| 61 | University of New England | 1 | 25 | 2730 | 2176 |
| 61 | New York Medical College | 1 | 25 | 1843 | 1576 |
| 61 | National University of Mar del Plata | 1 | 25 | 1112 | 675 |
| 61 | Royal Holloway, University of London | 1 | 25 | 2027 | 1705 |
| 61 | University of California, San Francisco | 1 | 25 | 5657 | 4845 |
| 68 | Merck & Co. | 1 | 22 | 1188 | 1326 |
| 69 | University of Washington | 2 | 21 | 8736 | 8967 |
| 69 | Aston University | 1 | 21 | 2405 | 2231 |
| 69 | University of Ferrara | 1 | 21 | 3200 | 2897 |
| 72 | University of Oklahoma | 1 | 20 | 6218 | 5202 |
| 73 | Tor Vergata University of Rome | 1 | 19 | 3224 | 2947 |
| 74 | University of Maryland | 1 | 18 | 10201 | 8660 |
| 74 | Joseph Fourier University | 1 | 18 | 1859 | 2214 |
| 74 | Swedish University of Agricultural Sciences | 1 | 18 | 2138 | 2205 |
| 77 | National Cancer Center | 1 | 17 | 2280 | 1996 |
| 78 | Russian Academy of Sciences | 2 | 16 | 6077 | 5533 |
| 78 | University of Akron | 1 | 16 | 2102 | 2130 |
| 80 | The Rockefeller University | 2 | 15 | 1121 | 1772 |
| 80 | University of California, Davis | 2 | 15 | 6710 | 7745 |
| 82 | University of Barcelona | 1 | 14 | 7120 | 6056 |
| 82 | University of Aberdeen | 1 | 14 | 4787 | 4791 |
| 82 | Texas Tech University | 1 | 14 | 5849 | 5342 |
| 85 | Université de Montréal | 1 | 13 | 7130 | 6683 |
| 86 | State University of New York Upstate Medical University | 1 | 12 | 1875 | 1970 |
| 87 | Fordham University | 3 | 11 | 814 | 1942 |
| 87 | University of Illinois at Chicago | 1 | 11 | 7001 | 7131 |
| 87 | United States Department of Veterans Affairs | 2 | 11 | 2261 | 3107 |
| 90 | Laboratory of Molecular Biology | 1 | 10 | 1849 | 2187 |
| 90 | Hungarian Academy of Sciences | 2 | 10 | 2353 | 3633 |
| 90 | Delft University of Technology | 1 | 10 | 4571 | 4730 |
| 90 | University of Debrecen | 2 | 10 | 1712 | 2569 |
| 90 | Howard University | 1 | 10 | 2173 | 2464 |
| 95 | University of Pittsburgh | 2 | 8 | 7072 | 8987 |
| 96 | Monsanto | 1 | 7 | 672 | 983 |
| 96 | University of Illinois at Urbana–Champaign | 1 | 7 | 7992 | 8467 |
| 96 | Washington University in St. Louis | 1 | 7 | 7519 | 7929 |
| 96 | Memorial Sloan Kettering Cancer Center | 1 | 7 | 2599 | 2989 |
| 96 | Uniformed Services University of the Health Sciences | 1 | 7 | 1949 | 2416 |
| 101 | University of Oslo | 1 | 6 | 7211 | 8010 |
| 101 | University of Leeds | 1 | 6 | 7632 | 8309 |
| 101 | Macquarie University | 1 | 6 | 5046 | 5955 |
| 101 | Hamedan University of Medical Sciences | 1 | 6 | 1054 | 1317 |
| 105 | Tufts University | 2 | 5 | 3803 | 6127 |
| 105 | University of Münster | 1 | 5 | 4776 | 5615 |
| 105 | Paris Descartes University | 1 | 5 | 3245 | 4149 |
| 108 | National Institute of Arthritis and Musculoskeletal and Skin Diseases | 1 | 4 | 294 | 536 |
| 109 | Stanford University | 2 | 3 | 8168 | 11591 |
| 109 | Northwestern University | 1 | 3 | 7017 | 8466 |
| 109 | Illinois College | 1 | 3 | 1073 | 1688 |
| 109 | Oak Ridge National Laboratory | 1 | 3 | 2646 | 3448 |
| 109 | University of Georgia | 1 | 3 | 6729 | 8090 |
| 109 | University of North Dakota | 1 | 3 | 2038 | 3131 |
| 109 | University of Virginia | 1 | 3 | 7348 | 8739 |
| 116 | University of Kansas | 1 | 2 | 6641 | 8441 |
| 116 | California Institute of Technology | 1 | 2 | 3239 | 4451 |
| 116 | Howard Hughes Medical Institute | 1 | 2 | 1503 | 2194 |
| 116 | Space Sciences Laboratory | 1 | 2 | 232 | 368 |
| 120 | Vanderbilt University | 1 | 1 | 6966 | 8949 |
| 120 | University of California | 1 | 1 | 9205 | 11520 |
| 120 | National Institute of Neurological Disorders and Stroke | 1 | 1 | 841 | 1416 |
| 120 | University of Campania "Luigi Vanvitelli" | 1 | 1 | 2310 | 3631 |
| 120 | DuPont Central Research | 1 | 1 | 3446 | 4695 |
| 120 | Indiana University | 1 | 1 | 9782 | 11953 |
| 120 | Beckman Center for Molecular and Genetic Medicine | 1 | 1 | 30 | 70 |
| 120 | University of California, Santa Barbara | 1 | 1 | 4268 | 6048 |
| 120 | University of Maryland, Baltimore | 1 | 1 | 3274 | 5116 |
| 120 | Microbial Ecology | 1 | 1 | 1273 | 1976 |
| 130 | University of Cambridge | 1 | 0 | 10005 | 13148 |
| 130 | Collège de France | 1 | 0 | 1269 | 2167 |
| 130 | Tel Aviv University | 1 | 0 | 7547 | 10121 |
| 130 | State University of New York | 1 | 0 | 8063 | 11153 |