| 1 | University of California, Davis | 14 | 311 | 884 | 1488 |
| 2 | University of Cambridge | 1 | 267 | 9676 | 3302 |
| 3 | Yale University | 3 | 252 | 5468 | 3081 |
| 4 | University of Münster | 3 | 247 | 2828 | 1426 |
| 5 | University of Würzburg | 8 | 245 | 1267 | 1322 |
| 6 | Schneider Children's Medical Center of Israel | 6 | 204 | 388 | 462 |
| 7 | John Radcliffe Hospital | 2 | 190 | 1414 | 856 |
| 8 | University of Pennsylvania | 5 | 189 | 5359 | 4196 |
| 9 | University of California | 10 | 179 | 8420 | 8416 |
| 10 | Oswaldo Cruz Foundation | 7 | 177 | 628 | 591 |
| 11 | Erasmus University Rotterdam | 3 | 174 | 2621 | 1778 |
| 12 | Vanderbilt University | 4 | 167 | 3862 | 2965 |
| 13 | United Arab Emirates University | 2 | 162 | 1680 | 629 |
| 14 | South University | 1 | 153 | 5756 | 2468 |
| 15 | University of Belgrade | 2 | 152 | 3499 | 1389 |
| 16 | Monash University | 11 | 133 | 2719 | 4287 |
| 17 | Howard Hughes Medical Institute | 3 | 126 | 990 | 1085 |
| 18 | Vrije Universiteit Brussel | 1 | 121 | 4039 | 1697 |
| 18 | Vrije Universiteit Amsterdam | 1 | 121 | 7960 | 3969 |
| 18 | French National Institute of Health and Medical Research | 8 | 121 | 3099 | 3392 |
| 21 | Royal Postgraduate Medical School | 5 | 116 | 374 | 500 |
| 22 | Weizmann Institute of Science | 4 | 113 | 1384 | 1448 |
| 22 | University of Chicago | 4 | 113 | 4751 | 4120 |
| 24 | University of Miami | 1 | 105 | 6090 | 3020 |
| 25 | Kumamoto University | 3 | 97 | 1832 | 1296 |
| 26 | University of Liège | 5 | 92 | 1607 | 2110 |
| 27 | University of Kansas Medical Center | 2 | 91 | 1682 | 1228 |
| 28 | University of Mississippi Medical Center | 3 | 90 | 991 | 987 |
| 29 | University of Copenhagen | 3 | 79 | 5249 | 4661 |
| 29 | Paris Diderot University | 6 | 79 | 1729 | 2536 |
| 31 | Maastricht University | 2 | 76 | 3282 | 2672 |
| 32 | Complutense University of Madrid | 6 | 74 | 2767 | 3249 |
| 33 | University of Oxford | 4 | 73 | 6198 | 6347 |
| 34 | University of British Columbia | 1 | 72 | 10589 | 6382 |
| 34 | Polish Academy of Sciences | 1 | 72 | 5349 | 3029 |
| 36 | Max Planck Society | 3 | 68 | 3248 | 3347 |
| 36 | Universidade do Estado do Rio de Janeiro | 2 | 68 | 1998 | 857 |
| 38 | University of Alabama at Birmingham | 6 | 66 | 2144 | 3256 |
| 38 | University of Crete | 2 | 66 | 1786 | 1596 |
| 40 | Pasteur Institute | 4 | 65 | 1233 | 1545 |
| 40 | Virginia Commonwealth University | 1 | 65 | 5112 | 3110 |
| 40 | VCU Medical Center | 1 | 65 | 1426 | 1039 |
| 43 | Cancer Research UK | 2 | 63 | 1028 | 1086 |
| 44 | Imperial College London | 2 | 60 | 6918 | 5851 |
| 44 | University of Massachusetts | 4 | 60 | 3680 | 4752 |
| 46 | Panum Building | 2 | 57 | 412 | 598 |
| 47 | Necker–Enfants Malades Hospital | 2 | 54 | 940 | 959 |
| 47 | Medical Research Council | 2 | 54 | 2069 | 2172 |
| 49 | University of Alabama | 4 | 53 | 2537 | 3474 |
| 50 | Paris Descartes University | 1 | 52 | 3149 | 2362 |
| 51 | University of Connecticut Health Center | 4 | 49 | 752 | 1438 |
| 52 | Bar-Ilan University | 1 | 47 | 3752 | 2342 |
| 53 | University of Graz | 2 | 46 | 3141 | 3006 |
| 53 | WEHI | 5 | 46 | 311 | 644 |
| 55 | California Institute of Technology | 2 | 45 | 2530 | 2693 |
| 55 | Jackson Laboratory | 4 | 45 | 322 | 605 |
| 57 | Bio-Medical Science (South Korean) | 2 | 43 | 6712 | 5747 |
| 58 | Sheba Medical Center | 1 | 42 | 1984 | 1578 |
| 58 | HaEmek Medical Center | 1 | 42 | 266 | 199 |
| 60 | Wageningen University & Research | 1 | 41 | 3542 | 3020 |
| 60 | Institut de chimie | 3 | 41 | 788 | 1042 |
| 60 | Halifax | 1 | 41 | 5834 | 4315 |
| 63 | University of Tokyo | 1 | 40 | 9472 | 6591 |
| 63 | Dokkyo University | 1 | 40 | 599 | 462 |
| 63 | Icahn School of Medicine at Mount Sinai | 2 | 40 | 3250 | 3201 |
| 63 | Tokyo Tokyo | 1 | 40 | 685 | 466 |
| 63 | Ludwig Cancer Research | 3 | 40 | 430 | 789 |
| 68 | Tel Aviv University | 1 | 39 | 7475 | 5382 |
| 68 | University of New South Wales | 1 | 39 | 9129 | 6833 |
| 68 | Tel Aviv Sourasky Medical Center | 1 | 39 | 1491 | 1209 |
| 68 | Kaplan Medical Center | 1 | 39 | 491 | 407 |
| 68 | Babraham Institute | 2 | 39 | 275 | 444 |
| 73 | Bernardo O'Higgins University | 3 | 38 | 12234 | 9083 |
| 73 | Leonard M. Miller School of Medicine | 1 | 38 | 3270 | 2668 |
| 73 | Westminster Hospital | 1 | 38 | 788 | 621 |
| 76 | Biomedical Primate Research Centre | 1 | 37 | 141 | 165 |
| 76 | Roswell Park Comprehensive Cancer Center | 1 | 37 | 1368 | 1176 |
| 78 | Harvard University | 3 | 36 | 8935 | 9516 |
| 78 | University of Bristol | 2 | 36 | 5281 | 5339 |
| 78 | Mayo Clinic | 2 | 36 | 3950 | 3774 |
| 81 | University of Rochester Medical Center | 2 | 35 | 1688 | 1930 |
| 81 | National Institute of Public Health | 2 | 35 | 1329 | 1566 |
| 81 | University of California, San Francisco | 5 | 35 | 1911 | 3232 |
| 84 | Johannes Gutenberg University Mainz | 1 | 33 | 3395 | 2810 |
| 84 | University of Toronto | 6 | 33 | 6435 | 9302 |
| 84 | Max Planck Institute for Biophysical Chemistry | 1 | 33 | 339 | 415 |
| 84 | University of Western Australia | 1 | 33 | 6626 | 5335 |
| 88 | Tor Vergata University of Rome | 1 | 32 | 2393 | 2076 |
| 89 | Fukuoka University | 1 | 31 | 1797 | 1289 |
| 89 | University of Maryland, Baltimore | 1 | 31 | 1205 | 1164 |
| 91 | Emory University | 4 | 30 | 3803 | 5365 |
| 92 | National Institutes of Health | 1 | 29 | 5077 | 4526 |
| 92 | Johns Hopkins School of Medicine | 1 | 29 | 3073 | 2787 |
| 92 | University of South Carolina | 1 | 29 | 7508 | 6050 |
| 92 | Center for Research and Advanced Studies of the National Polytechnic Institute | 1 | 29 | 1963 | 1632 |
| 92 | Geisel School of Medicine | 1 | 29 | 2038 | 1967 |
| 97 | University of Washington | 1 | 28 | 10505 | 8682 |
| 97 | University of Groningen | 3 | 28 | 4101 | 5567 |
| 97 | Centenary Institute | 3 | 28 | 153 | 456 |
| 100 | Australian National University | 2 | 27 | 4933 | 5469 |
| 100 | University of Göttingen | 1 | 27 | 4839 | 4168 |
| 100 | Nippon Zoki Pharmaceutical | 3 | 27 | 9 | 32 |
| 100 | Society for Endocrinology | 1 | 27 | 296 | 370 |
| 104 | University College London | 2 | 26 | 7711 | 7927 |
| 104 | University of Birmingham | 2 | 26 | 6060 | 6517 |
| 104 | University of Salamanca | 1 | 26 | 3343 | 2462 |
| 104 | Kyoto University | 1 | 26 | 7583 | 5991 |
| 104 | MRC London Institute of Medical Sciences | 1 | 26 | 388 | 506 |
| 109 | Stanford University | 2 | 25 | 8467 | 8879 |
| 109 | Royal Melbourne Hospital | 3 | 25 | 938 | 1612 |
| 111 | University of Bern | 1 | 24 | 5717 | 5053 |
| 111 | California State University | 1 | 24 | 5645 | 4504 |
| 111 | University of Bari | 1 | 24 | 3405 | 3266 |
| 111 | Salk Institute for Biological Studies | 1 | 24 | 772 | 927 |
| 111 | Moss Landing Marine Laboratories | 1 | 24 | 147 | 182 |
| 111 | Oklahoma Medical Research Foundation | 1 | 24 | 526 | 650 |
| 117 | Keele University | 1 | 23 | 2239 | 2217 |
| 118 | University of Alberta | 1 | 22 | 9586 | 8138 |
| 118 | Autonomous University of Madrid | 1 | 22 | 4587 | 3784 |
| 118 | University of Innsbruck | 1 | 22 | 4125 | 3734 |
| 118 | University of Verona | 1 | 22 | 3138 | 2884 |
| 118 | Grinnell College | 1 | 22 | 1377 | 1472 |
| 118 | Cross Cancer Institute | 1 | 22 | 432 | 536 |
| 118 | Hospital Universitario de La Princesa | 1 | 22 | 609 | 619 |
| 118 | interdisciplinarity | 1 | 22 | 10684 | 8390 |
| 118 | University of Iowa | 1 | 22 | 6880 | 6069 |
| 127 | Collège de France | 3 | 21 | 618 | 1265 |
| 127 | Kanazawa University | 2 | 21 | 2531 | 2869 |
| 127 | Military Medical Academy | 4 | 21 | 615 | 1195 |
| 130 | University Hospital Zurich | 1 | 20 | 1947 | 1996 |
| 130 | University of Arizona | 2 | 20 | 6655 | 7646 |
| 130 | Robert Koch Institute | 1 | 20 | 645 | 788 |
| 130 | Scripps Research | 2 | 20 | 1183 | 1746 |
| 130 | University of Saskatchewan | 2 | 20 | 3794 | 4908 |
| 135 | German Cancer Research Center | 1 | 19 | 1073 | 1194 |
| 135 | Fukushima Medical University | 1 | 19 | 1364 | 1226 |
| 135 | Hammersmith Hospital | 4 | 19 | 696 | 1550 |
| 138 | University of Szeged | 1 | 18 | 2504 | 2429 |
| 138 | Centre for Blood Research | 1 | 18 | 248 | 375 |
| 138 | Ontario Cancer Institute | 2 | 18 | 362 | 708 |
| 141 | University of León | 1 | 17 | 1325 | 1341 |
| 141 | St. Jude Children's Research Hospital | 1 | 17 | 1341 | 1545 |
| 141 | University of Brescia | 1 | 17 | 2799 | 2901 |
| 141 | Erasmus Hospital | 1 | 17 | 984 | 1096 |
| 141 | Mario Negri Institute for Pharmacological Research | 1 | 17 | 849 | 1069 |
| 141 | Marie Curie | 2 | 17 | 3545 | 4687 |
| 147 | Max Planck Institute for Informatics | 1 | 16 | 249 | 337 |
| 147 | University of Queensland | 1 | 16 | 9836 | 9253 |
| 149 | Imperial College School of Medicine | 4 | 15 | 391 | 1160 |
| 150 | University of Minnesota | 1 | 14 | 10588 | 9909 |
| 150 | Ohio State University | 2 | 14 | 9547 | 10576 |
| 150 | Loma Linda University | 3 | 14 | 1039 | 2213 |
| 150 | Carlos III Health Institute | 2 | 14 | 1524 | 2257 |
| 154 | University of Texas at Austin | 2 | 13 | 9263 | 10663 |
| 154 | University of Turku | 1 | 13 | 4672 | 4924 |
| 154 | University of Texas MD Anderson Cancer Center | 2 | 13 | 2485 | 3360 |
| 157 | University of Freiburg | 1 | 12 | 4721 | 5086 |
| 157 | Wellesley Hospital | 3 | 12 | 94 | 351 |
| 159 | TU Dresden | 1 | 11 | 5012 | 5322 |
| 159 | Michigan State University | 1 | 11 | 8627 | 8794 |
| 159 | Osaka University | 1 | 11 | 6351 | 6186 |
| 162 | University of Tübingen | 1 | 10 | 4561 | 5102 |
| 162 | University of Edinburgh | 1 | 10 | 8499 | 8676 |
| 162 | University of Auckland | 2 | 10 | 5188 | 7479 |
| 162 | University of Geneva | 1 | 10 | 5143 | 5684 |
| 162 | Royal Infirmary of Edinburgh | 1 | 10 | 835 | 1086 |
| 162 | Kansai Medical University | 1 | 10 | 1162 | 1278 |
| 162 | University of California, Los Angeles | 1 | 10 | 5041 | 5948 |
| 162 | Henry M. Jackson Foundation | 1 | 10 | 331 | 517 |
| 170 | Waseda University | 1 | 9 | 3554 | 3798 |
| 170 | University of Gothenburg | 1 | 9 | 6324 | 7034 |
| 170 | University of Wisconsin–Madison | 2 | 9 | 9072 | 11352 |
| 170 | Umeå University | 1 | 9 | 4165 | 5107 |
| 170 | Biology Institute of Shandong Academy of Sciences | 1 | 9 | 3437 | 3805 |
| 175 | Princeton University | 1 | 8 | 6623 | 7323 |
| 175 | University of Basel | 1 | 8 | 4555 | 5320 |
| 175 | Los Alamos National Laboratory | 1 | 8 | 2312 | 2802 |
| 175 | University of Cape Town | 1 | 8 | 5019 | 5898 |
| 175 | Stony Brook University | 1 | 8 | 4942 | 5874 |
| 175 | University of Basilicata | 1 | 8 | 995 | 1494 |
| 175 | University of Lille | 1 | 8 | 3367 | 4256 |
| 175 | Sysmex Corporation | 1 | 8 | 104 | 174 |
| 175 | UCL Medical School | 1 | 8 | 1563 | 2066 |
| 175 | The Honourable Society of Lincoln's Inn | 1 | 8 | 446 | 686 |
| 185 | University of Michigan | 1 | 7 | 12506 | 12759 |
| 185 | University of Melbourne | 1 | 7 | 9897 | 10661 |
| 185 | Czech Academy of Sciences | 1 | 7 | 4014 | 4586 |
| 185 | Amgen | 1 | 7 | 706 | 1091 |
| 185 | Pennsylvania State University | 1 | 7 | 10050 | 10752 |
| 185 | University of Calgary | 1 | 7 | 7875 | 8795 |
| 185 | Thomas Jefferson University | 2 | 7 | 3448 | 5362 |
| 185 | Centre d'Immunologie de Marseille-Luminy | 1 | 7 | 161 | 294 |
| 185 | John Curtin School of Medical Research | 1 | 7 | 650 | 1007 |
| 185 | Lone Star College System | 1 | 7 | 7525 | 8671 |
| 195 | University of Glasgow | 1 | 6 | 7093 | 8192 |
| 195 | University of Georgia | 1 | 6 | 6660 | 7669 |
| 195 | MRC Human Genetics Unit | 1 | 6 | 323 | 550 |
| 195 | Washington University School of Medicine | 1 | 6 | 3942 | 4639 |
| 195 | Western General Hospital | 1 | 6 | 1493 | 2245 |
| 200 | Korea Research Institute of Bioscience and Biotechnology | 1 | 5 | 840 | 1236 |
| 201 | Ludwig Maximilian University of Munich | 1 | 4 | 8526 | 9912 |
| 201 | University of Burgundy | 1 | 4 | 2645 | 3723 |
| 201 | Courtauld Institute of Art | 1 | 4 | 118 | 188 |
| 201 | Japan Science and Technology Agency | 1 | 4 | 1816 | 2520 |
| 201 | Nebraska Medical Center | 1 | 4 | 2865 | 3836 |
| 206 | University of Milan | 1 | 3 | 6969 | 8608 |
| 206 | Massachusetts Institute of Technology | 1 | 3 | 6245 | 8168 |
| 206 | Durham University | 1 | 3 | 4505 | 6050 |
| 206 | University of Nottingham | 1 | 3 | 7729 | 9743 |
| 206 | Queen's Medical Centre | 1 | 3 | 1397 | 2175 |
| 206 | University Medical Center | 1 | 3 | 7963 | 8835 |
| 212 | Columbia University | 1 | 2 | 10018 | 12106 |
| 212 | University of Marburg | 1 | 2 | 2603 | 3914 |
| 212 | Centers for Disease Control and Prevention | 1 | 2 | 3634 | 5027 |
| 212 | University of Virginia | 1 | 2 | 7028 | 9136 |
| 216 | University of Cologne | 1 | 1 | 5026 | 7056 |
| 216 | Sanofi | 1 | 1 | 1589 | 2480 |
| 216 | University of Debrecen | 1 | 1 | 2505 | 3899 |
| 216 | Hokkaido University | 1 | 1 | 5870 | 7537 |
| 216 | University of Sussex | 1 | 1 | 4172 | 6395 |
| 216 | Institut Jacques Monod | 1 | 1 | 346 | 672 |
| 216 | Medway School of Science | 1 | 1 | 8584 | 10869 |