| 1 | Max Planck Society | 191 | 15.0K | 169 | 125 |
| 2 | French National Institute of Health and Medical Research | 142 | 7.6K | 475 | 345 |
| 3 | Howard Hughes Medical Institute | 103 | 7.6K | 95 | 130 |
| 4 | University of California | 119 | 7.2K | 1477 | 1204 |
| 5 | Harvard University | 95 | 7.2K | 1034 | 778 |
| 6 | University of Tokyo | 125 | 5.7K | 546 | 346 |
| 7 | Medway School of Science | 107 | 4.8K | 487 | 265 |
| 8 | Osaka University | 79 | 4.7K | 452 | 206 |
| 9 | Kyoto University | 86 | 4.7K | 512 | 282 |
| 10 | University of Helsinki | 65 | 3.8K | 333 | 277 |
| 11 | University of Cambridge | 95 | 3.8K | 360 | 549 |
| 12 | Collège de France | 39 | 3.7K | 51 | 38 |
| 13 | University of Pennsylvania | 47 | 3.2K | 1044 | 739 |
| 14 | Columbia University | 46 | 3.1K | 928 | 714 |
| 15 | University of Washington | 35 | 3.0K | 1502 | 898 |
| 16 | University of Southern California | 31 | 2.9K | 916 | 467 |
| 17 | Cancer Research UK | 42 | 2.9K | 124 | 137 |
| 18 | National Institutes of Health | 44 | 2.8K | 726 | 617 |
| 19 | Guy's Hospital | 21 | 2.7K | 187 | 71 |
| 20 | University of Toronto | 43 | 2.7K | 1454 | 989 |
| 20 | Japan Science and Technology Agency | 49 | 2.7K | 131 | 135 |
| 22 | Vanderbilt University | 39 | 2.6K | 651 | 461 |
| 23 | University of Texas at Austin | 43 | 2.2K | 1155 | 990 |
| 24 | University Medical Center | 31 | 2.2K | 2184 | 1184 |
| 25 | Bio-Medical Science (South Korean) | 56 | 2.2K | 1089 | 908 |
| 26 | National Institute of Genetics | 41 | 2.1K | 30 | 43 |
| 27 | Western General Hospital | 23 | 2.1K | 129 | 71 |
| 28 | Lunenfeld-Tanenbaum Research Institute | 21 | 2.1K | 61 | 58 |
| 28 | Sinai Hospital | 22 | 2.1K | 311 | 154 |
| 30 | Wellcome Trust | 26 | 2.0K | 138 | 148 |
| 31 | Ulm University | 29 | 2.0K | 151 | 67 |
| 32 | MRC Human Genetics Unit | 34 | 1.9K | 10 | 24 |
| 33 | King's College London | 66 | 1.8K | 282 | 519 |
| 34 | European Molecular Biology Laboratory | 43 | 1.7K | 56 | 111 |
| 35 | Pasteur Institute | 47 | 1.7K | 196 | 246 |
| 36 | Baylor College of Medicine | 34 | 1.6K | 595 | 502 |
| 37 | Nagoya University | 30 | 1.6K | 875 | 410 |
| 38 | Princeton University | 28 | 1.6K | 811 | 727 |
| 39 | University of Würzburg | 21 | 1.6K | 548 | 321 |
| 40 | Icahn School of Medicine at Mount Sinai | 36 | 1.6K | 499 | 475 |
| 41 | University of Cologne | 27 | 1.6K | 479 | 311 |
| 42 | University of Göttingen | 31 | 1.5K | 411 | 361 |
| 43 | Laboratory of Molecular Biology | 24 | 1.5K | 183 | 169 |
| 44 | Babraham Institute | 7 | 1.5K | 118 | 58 |
| 45 | University of Virginia | 24 | 1.5K | 806 | 587 |
| 46 | University College London | 59 | 1.4K | 631 | 1138 |
| 47 | Children's Hospital Los Angeles | 9 | 1.4K | 305 | 78 |
| 48 | University of Freiburg | 18 | 1.4K | 671 | 384 |
| 49 | California Institute of Technology | 27 | 1.4K | 503 | 546 |
| 50 | University of Texas MD Anderson Cancer Center | 28 | 1.4K | 527 | 440 |
| 51 | Weizmann Institute of Science | 14 | 1.4K | 592 | 332 |
| 52 | University of Chicago | 22 | 1.3K | 1393 | 1013 |
| 53 | University of Bergen | 17 | 1.3K | 681 | 361 |
| 54 | Mitsubishi Electric | 13 | 1.3K | 190 | 29 |
| 55 | Saarland University | 6 | 1.3K | 1276 | 194 |
| 56 | University of Edinburgh | 71 | 1.3K | 313 | 879 |
| 57 | Massachusetts General Hospital | 20 | 1.3K | 1309 | 928 |
| 58 | National Institute for Basic Biology, Japan | 29 | 1.3K | 28 | 53 |
| 59 | University of Queensland | 29 | 1.2K | 1300 | 1211 |
| 60 | McGill University | 27 | 1.2K | 1300 | 1139 |
| 60 | University of Tokushima | 32 | 1.2K | 176 | 104 |
| 62 | Kumamoto University | 33 | 1.2K | 229 | 183 |
| 63 | Hadassah Medical Center | 16 | 1.2K | 256 | 144 |
| 64 | Eunice Kennedy Shriver National Institute of Child Health and Human Development | 23 | 1.1K | 148 | 179 |
| 65 | Tohoku University | 23 | 1.1K | 1305 | 734 |
| 66 | The Honourable Society of Lincoln's Inn | 13 | 1.1K | 80 | 76 |
| 67 | University of Michigan | 23 | 1.1K | 2537 | 2011 |
| 68 | Centre de Biologie du Développement | 20 | 1.1K | 3 | 6 |
| 69 | Karolinska Institute | 20 | 1.1K | 1388 | 1040 |
| 70 | Paris Diderot University | 23 | 1.1K | 526 | 462 |
| 70 | University of Bath | 27 | 1.1K | 247 | 309 |
| 72 | Tokyo Medical and Dental University | 12 | 1.0K | 676 | 237 |
| 73 | Technical University of Braunschweig | 12 | 1.0K | 152 | 62 |
| 74 | Johns Hopkins University | 17 | 1.0K | 2022 | 1453 |
| 75 | Boston University | 5 | 1.0K | 3354 | 996 |
| 76 | University of Massachusetts | 20 | 1.0K | 910 | 851 |
| 77 | National University of Singapore | 69 | 1.0K | 327 | 1086 |
| 78 | University of Minnesota | 15 | 977 | 2828 | 1772 |
| 79 | Nara Institute of Science and Technology | 17 | 963 | 134 | 73 |
| 80 | University of California, San Diego | 10 | 953 | 1593 | 850 |
| 81 | German Cancer Research Center | 14 | 950 | 185 | 145 |
| 82 | Autonomous University of Madrid | 28 | 943 | 272 | 314 |
| 83 | Department of Embryology | 9 | 941 | 61 | 36 |
| 84 | University of California, San Francisco | 9 | 935 | 1343 | 640 |
| 85 | Newcastle University | 17 | 929 | 916 | 675 |
| 86 | Aichi University of Education | 1 | 906 | 292 | 4 |
| 87 | University of Zurich | 23 | 896 | 700 | 760 |
| 88 | Research Institute of Molecular Pathology | 8 | 879 | 44 | 47 |
| 89 | Carnegie Institution for Science | 5 | 878 | 338 | 162 |
| 90 | Case Western Reserve University | 14 | 871 | 1416 | 925 |
| 91 | Heidelberg University | 29 | 863 | 550 | 771 |
| 92 | Istituto di Genetica e Biofisica | 8 | 858 | 46 | 22 |
| 93 | The Graduate University for Advanced Studies | 16 | 853 | 74 | 49 |
| 94 | Cancer Research Center | 17 | 843 | 747 | 630 |
| 95 | National Institute of Neurological Disorders and Stroke | 5 | 840 | 329 | 170 |
| 96 | Hokkaido University | 15 | 825 | 1602 | 800 |
| 97 | Great Ormond Street Hospital | 22 | 824 | 639 | 617 |
| 98 | Children's Medical Research Institute | 10 | 823 | 32 | 35 |
| 99 | Wellcome | 14 | 814 | 297 | 260 |
| 100 | Jackson Laboratory | 13 | 811 | 103 | 113 |
| 101 | Duke University Hospital | 11 | 808 | 1139 | 771 |
| 102 | University of Basel | 20 | 806 | 582 | 635 |
| 103 | Washington University School of Medicine | 14 | 805 | 1164 | 861 |
| 104 | University of Texas Southwestern Medical Center | 17 | 804 | 874 | 730 |
| 105 | The Rockefeller University | 23 | 787 | 208 | 353 |
| 106 | Whitehead Institute | 11 | 783 | 69 | 113 |
| 107 | King's Buildings | 8 | 779 | 281 | 159 |
| 108 | University of Hamburg | 8 | 778 | 746 | 324 |
| 109 | University of Oxford | 44 | 773 | 1061 | 2017 |
| 110 | Massachusetts Institute of Technology | 11 | 772 | 1829 | 1408 |
| 111 | Karlsruhe Institute of Technology | 13 | 771 | 956 | 569 |
| 112 | University of Geneva | 21 | 770 | 598 | 691 |
| 113 | Max Planck Institute for Brain Research | 9 | 762 | 45 | 39 |
| 114 | University of Murcia | 13 | 759 | 443 | 202 |
| 115 | Medical Research Council | 10 | 754 | 782 | 565 |
| 116 | Max Planck Institute of Molecular Cell Biology and Genetics | 20 | 753 | 24 | 56 |
| 117 | Katholieke Universiteit Leuven | 17 | 752 | 1593 | 1405 |
| 118 | Yale University | 20 | 735 | 1585 | 1737 |
| 119 | Hebrew University of Jerusalem | 16 | 716 | 889 | 780 |
| 120 | Salk Institute for Biological Studies | 17 | 700 | 124 | 217 |
| 121 | Humboldt University of Berlin | 10 | 699 | 2950 | 1432 |
| 122 | University of Sheffield | 46 | 692 | 378 | 1112 |
| 123 | Norwegian Nobel Institute | 5 | 688 | 66 | 42 |
| 124 | Weill Cornell Medicine | 12 | 684 | 1347 | 878 |
| 125 | Institut de chimie | 7 | 682 | 499 | 225 |
| 126 | Ludwig Cancer Research | 17 | 679 | 119 | 195 |
| 127 | Kobe University | 14 | 674 | 881 | 411 |
| 127 | Graduate University for Advanced Studies | 15 | 674 | 122 | 97 |
| 129 | Baylor University | 15 | 672 | 490 | 422 |
| 130 | Albert Einstein College of Medicine | 15 | 660 | 803 | 740 |
| 131 | University of Wisconsin School of Medicine and Public Health | 5 | 653 | 765 | 284 |
| 132 | ETH Zürich | 10 | 652 | 1435 | 1087 |
| 132 | Johns Hopkins School of Medicine | 7 | 652 | 1145 | 618 |
| 134 | Cincinnati Children's Hospital Medical Center | 15 | 650 | 514 | 485 |
| 135 | Pierre and Marie Curie University | 14 | 646 | 530 | 507 |
| 136 | Stanford University | 26 | 645 | 2014 | 2664 |
| 137 | Ohio State University | 20 | 643 | 2723 | 2541 |
| 138 | University of Nottingham | 29 | 638 | 753 | 1262 |
| 139 | Hunt's | 12 | 628 | 12 | 26 |
| 140 | University of Utah | 18 | 625 | 1362 | 1393 |
| 141 | St. Jude Children's Research Hospital | 7 | 623 | 446 | 273 |
| 142 | University of South Carolina | 11 | 600 | 1968 | 1289 |
| 143 | Bernardo O'Higgins University | 21 | 599 | 4081 | 2203 |
| 144 | Pitié-Salpêtrière Hospital | 6 | 590 | 863 | 423 |
| 145 | University of Oregon | 15 | 588 | 452 | 544 |
| 145 | University of Kentucky | 8 | 588 | 2489 | 1178 |
| 147 | Brigham and Women's Hospital | 6 | 586 | 2263 | 1235 |
| 147 | Royal Melbourne Hospital | 14 | 586 | 289 | 301 |
| 149 | University of Adelaide | 16 | 582 | 1037 | 1079 |
| 149 | Pennsylvania State University | 10 | 582 | 3197 | 2001 |
| 151 | Harvard School of Dental Medicine | 5 | 573 | 154 | 95 |
| 152 | Institut Jacques Monod | 19 | 566 | 38 | 80 |
| 153 | University of Warwick | 16 | 542 | 816 | 860 |
| 154 | Karger Publishers | 2 | 539 | 1492 | 278 |
| 155 | University of Tsukuba | 13 | 538 | 1233 | 708 |
| 156 | Chinese Academy of Sciences | 24 | 532 | 2764 | 2669 |
| 157 | Indiana University | 13 | 523 | 2272 | 1908 |
| 158 | Hammersmith Hospital | 1 | 521 | 1429 | 345 |
| 159 | Niigata University | 8 | 520 | 975 | 352 |
| 160 | University of Illinois at Chicago | 14 | 518 | 1138 | 1111 |
| 160 | National Institute of Mental Health | 4 | 518 | 695 | 368 |
| 162 | NYU Langone Medical Center | 3 | 516 | 356 | 93 |
| 163 | Alexandria University | 5 | 514 | 2575 | 666 |
| 164 | Cellectis | 1 | 513 | 25 | 8 |
| 165 | Freie Universität Berlin | 2 | 508 | 177 | 10 |
| 165 | University of Melbourne | 35 | 508 | 1112 | 2302 |
| 167 | Tokyo Institute of Technology | 12 | 495 | 1148 | 714 |
| 168 | Lund University | 10 | 494 | 2295 | 1675 |
| 169 | Uppsala University | 11 | 492 | 1887 | 1503 |
| 170 | Kyushu University | 13 | 487 | 1835 | 1189 |
| 171 | University of Hohenheim | 15 | 479 | 221 | 281 |
| 172 | Universidad Metropolitana | 10 | 478 | 1359 | 589 |
| 173 | Medizinische Hochschule Hannover | 6 | 473 | 341 | 157 |
| 174 | Umeå University | 8 | 472 | 1170 | 806 |
| 175 | Royal Victoria Hospital | 4 | 471 | 473 | 185 |
| 176 | Stony Brook University | 9 | 467 | 1225 | 941 |
| 176 | University of Dundee | 25 | 467 | 219 | 644 |
| 178 | University of Connecticut Health Center | 8 | 458 | 374 | 345 |
| 179 | University of Wisconsin–Madison | 16 | 457 | 2798 | 2921 |
| 180 | University of Bari | 4 | 454 | 1543 | 546 |
| 181 | Lone Star College System | 8 | 449 | 2740 | 1930 |
| 182 | Max Planck Institute for Developmental Biology | 6 | 441 | 55 | 61 |
| 183 | Michigan Medicine | 4 | 432 | 1758 | 842 |
| 184 | King's College | 9 | 429 | 1203 | 991 |
| 185 | Leonard M. Miller School of Medicine | 8 | 427 | 1110 | 784 |
| 186 | University of Oulu | 5 | 425 | 1573 | 705 |
| 187 | University of Marburg | 9 | 422 | 619 | 473 |
| 188 | McMaster University | 5 | 419 | 3384 | 1602 |
| 189 | University of Liège | 8 | 414 | 1067 | 727 |
| 190 | RIKEN | 6 | 412 | 402 | 231 |
| 191 | University of Lübeck | 3 | 409 | 997 | 332 |
| 191 | Sanford Burnham Prebys Medical Discovery Institute | 6 | 409 | 166 | 196 |
| 193 | Montpellier 2 University | 5 | 407 | 721 | 442 |
| 194 | University of Birmingham | 12 | 403 | 1946 | 1824 |
| 195 | Utrecht University | 8 | 397 | 2416 | 1920 |
| 195 | UCL Medical School | 5 | 397 | 673 | 468 |
| 197 | University of Hawaii | 8 | 396 | 757 | 677 |
| 198 | Gurdon Institute | 14 | 393 | 23 | 70 |
| 199 | WEHI | 13 | 390 | 159 | 229 |
| 200 | Tufts University | 11 | 389 | 1083 | 1209 |
| 201 | University of Valladolid | 4 | 388 | 1108 | 272 |
| 202 | Marie Curie | 17 | 384 | 866 | 1272 |
| 203 | University of Münster | 10 | 383 | 1216 | 1073 |
| 203 | Memorial Sloan Kettering Cancer Center | 13 | 383 | 710 | 796 |
| 205 | University of Düsseldorf | 7 | 381 | 777 | 555 |
| 206 | National Eye Institute | 6 | 379 | 147 | 150 |
| 207 | National Institute of Neurosciences & Hospital | 8 | 374 | 137 | 128 |
| 208 | University of Pisa | 11 | 368 | 1306 | 1191 |
| 209 | National Scientific and Technical Research Council | 10 | 366 | 2330 | 1316 |
| 210 | University of Montpellier | 6 | 365 | 1615 | 915 |
| 210 | Developmental Biology Institute of Marseille | 6 | 365 | 6 | 16 |
| 212 | University of Padua | 8 | 356 | 2524 | 1816 |
| 212 | University of New Mexico | 8 | 356 | 1484 | 1148 |
| 212 | The Roslin Institute | 27 | 356 | 48 | 163 |
| 215 | Tata Institute of Fundamental Research | 9 | 353 | 279 | 184 |
| 216 | Medical College of Wisconsin | 5 | 351 | 1431 | 770 |
| 217 | Center for Biologics Evaluation and Research | 3 | 349 | 256 | 134 |
| 218 | Technion – Israel Institute of Technology | 10 | 347 | 1451 | 1249 |
| 219 | University of Calgary | 18 | 338 | 1255 | 1955 |
| 220 | Los Alamos National Laboratory | 2 | 335 | 1779 | 767 |
| 221 | Johannes Gutenberg University Mainz | 8 | 334 | 894 | 797 |
| 222 | Rutgers University | 8 | 333 | 3269 | 2401 |
| 223 | University of Konstanz | 6 | 322 | 781 | 582 |
| 224 | City of Hope National Medical Center | 1 | 321 | 1402 | 414 |
| 225 | National Research Centre | 8 | 319 | 1653 | 1089 |
| 226 | University of Genoa | 6 | 318 | 1730 | 1017 |
| 227 | Leiden University | 9 | 316 | 1982 | 1860 |
| 227 | Imperial College London | 11 | 316 | 2962 | 2976 |
| 229 | Fox Chase Cancer Center | 4 | 311 | 435 | 309 |
| 230 | University of Wales | 2 | 309 | 2241 | 832 |
| 231 | University of Maryland | 10 | 308 | 3465 | 3307 |
| 231 | University of Newcastle | 6 | 308 | 1877 | 1351 |
| 233 | Kansas State University | 7 | 306 | 1120 | 857 |
| 234 | National Defense Medical College | 3 | 304 | 572 | 158 |
| 235 | Emory University | 11 | 303 | 2089 | 2171 |
| 236 | Stowers Institute for Medical Research | 9 | 302 | 39 | 74 |
| 237 | RIKEN Brain Science Institute | 5 | 301 | 123 | 116 |
| 238 | National Institute of Health Sciences | 5 | 300 | 208 | 127 |
| 239 | University of Amsterdam | 5 | 299 | 3863 | 2506 |
| 240 | Kyowa Hakko Kirin | 1 | 298 | 359 | 73 |
| 241 | Universiti Sains Malaysia | 7 | 296 | 2372 | 1230 |
| 242 | Paris Descartes University | 6 | 293 | 1260 | 1003 |
| 242 | Royal Children's Hospital | 6 | 293 | 534 | 464 |
| 244 | Jichi Medical University | 6 | 289 | 674 | 351 |
| 245 | Helsinki University Central Hospital | 5 | 286 | 1100 | 788 |
| 246 | John Radcliffe Hospital | 6 | 283 | 763 | 679 |
| 247 | Cornell University | 9 | 281 | 3110 | 3134 |
| 248 | University of Chile | 21 | 280 | 485 | 990 |
| 248 | Tokai University | 4 | 280 | 1207 | 439 |
| 250 | Dana–Farber Cancer Institute | 8 | 279 | 640 | 713 |
| 250 | University of Agriculture | 2 | 279 | 3082 | 726 |
| 252 | Gunma University | 4 | 275 | 1251 | 492 |
| 253 | University of Naples Federico II | 8 | 274 | 2499 | 2114 |
| 254 | Hiroshima University | 9 | 273 | 1535 | 1052 |
| 255 | University of North Carolina at Chapel Hill | 8 | 271 | 3396 | 3154 |
| 256 | Swarthmore College | 2 | 269 | 340 | 97 |
| 257 | Thomas Jefferson University | 6 | 266 | 1781 | 1374 |
| 258 | Arizona State University | 5 | 264 | 3378 | 2159 |
| 259 | Oregon Health & Science University | 6 | 263 | 1726 | 1359 |
| 260 | United States Department of Agriculture | 2 | 261 | 1540 | 757 |
| 260 | Agricultural Research Service | 2 | 261 | 2095 | 1126 |
| 262 | Barnard College | 5 | 260 | 156 | 142 |
| 263 | Wesleyan University | 5 | 259 | 337 | 251 |
| 263 | Namiki | 2 | 259 | 704 | 291 |
| 263 | Academia Sinica | 21 | 259 | 566 | 1072 |
| 266 | University of Missouri | 8 | 258 | 3004 | 2583 |
| 266 | University of Ottawa | 6 | 258 | 2792 | 2036 |
| 266 | Yokohama City University | 6 | 258 | 792 | 486 |
| 269 | University of Alabama at Birmingham | 7 | 257 | 1955 | 1666 |
| 270 | Statens Serum Institut | 2 | 256 | 483 | 240 |
| 271 | Tulane University | 6 | 255 | 1488 | 1153 |
| 271 | University of Kent | 6 | 255 | 856 | 629 |
| 273 | Bryn Mawr College | 2 | 253 | 327 | 90 |
| 274 | Roswell Park Comprehensive Cancer Center | 2 | 252 | 980 | 452 |
| 275 | North Carolina State University | 4 | 251 | 3265 | 1940 |
| 276 | Academic Medical Center | 5 | 248 | 1181 | 1064 |
| 277 | University of Parma | 1 | 247 | 3157 | 764 |
| 278 | Max Planck Institute for Biophysical Chemistry | 6 | 246 | 98 | 138 |
| 278 | University of North Carolina | 7 | 246 | 3951 | 3430 |
| 280 | University of Colorado | 5 | 245 | 3612 | 2598 |
| 281 | University of Miami | 4 | 242 | 3196 | 1920 |
| 282 | San Francisco State University | 4 | 241 | 512 | 343 |
| 283 | State University of New York | 6 | 240 | 3383 | 2621 |
| 284 | Geisel School of Medicine | 4 | 239 | 944 | 730 |
| 285 | Université libre de Bruxelles | 8 | 238 | 1156 | 1238 |
| 285 | University of Iowa | 8 | 238 | 2404 | 2431 |
| 287 | University of Duisburg-Essen | 7 | 237 | 1223 | 1047 |
| 288 | University of Pittsburgh | 11 | 235 | 2719 | 3164 |
| 288 | University of Arizona | 8 | 235 | 2957 | 2832 |
| 288 | University of Cincinnati Academic Health Center | 6 | 235 | 789 | 720 |
| 291 | Consiglio Nazionale delle Ricerche | 4 | 232 | 1292 | 868 |
| 292 | Instituto Gulbenkian de Ciência | 13 | 230 | 25 | 119 |
| 293 | Public Health Institute Ostrava | 3 | 228 | 4625 | 2219 |
| 294 | Texas A&M University | 8 | 227 | 3321 | 3096 |
| 295 | Cardiff University | 7 | 226 | 1880 | 1860 |
| 296 | Deaconess Medical Center | 4 | 224 | 1684 | 1249 |
| 297 | Erasmus University Rotterdam | 6 | 223 | 1645 | 1547 |
| 298 | University of Otago | 3 | 222 | 2993 | 1602 |
| 299 | Biotechnology Research Center | 3 | 221 | 890 | 435 |
| 299 | University of Tennessee | 2 | 221 | 5218 | 2130 |
| 301 | Durham University | 7 | 219 | 1298 | 1225 |
| 302 | West Virginia University | 5 | 218 | 1850 | 1232 |
| 302 | Biomedicum Helsinki | 5 | 218 | 90 | 148 |
| 302 | Robert Wood Johnson Medical School | 4 | 218 | 998 | 704 |
| 305 | University of Bern | 5 | 214 | 2685 | 2068 |
| 305 | Nanyang Technological University | 7 | 214 | 2115 | 1887 |
| 307 | Brown University | 9 | 213 | 1737 | 2072 |
| 307 | University of Copenhagen | 3 | 213 | 5249 | 3019 |
| 307 | Queen's Medical Centre | 6 | 213 | 457 | 493 |
| 307 | University of the Mediterranean - Aix Marseille II | 2 | 213 | 146 | 78 |
| 311 | Genentech | 6 | 211 | 419 | 522 |
| 312 | University of Massachusetts Boston | 2 | 210 | 843 | 270 |
| 312 | University of Tromsø – The Arctic University of Norway | 2 | 210 | 2015 | 861 |
| 314 | Keio University | 10 | 208 | 1388 | 1315 |
| 314 | Spanish National Research Council | 7 | 208 | 981 | 1078 |
| 316 | Children's Hospital of Philadelphia | 5 | 207 | 1098 | 917 |
| 317 | Wayne State University | 7 | 206 | 2050 | 1930 |
| 318 | University of Bremen | 5 | 205 | 1111 | 822 |
| 318 | University of Sussex | 8 | 205 | 1032 | 1339 |
| 320 | Centre for Life | 8 | 204 | 287 | 480 |
| 321 | Sapienza University of Rome | 7 | 203 | 3448 | 2932 |
| 322 | Beth Israel Deaconess Medical Center | 5 | 202 | 1600 | 1384 |
| 323 | Université catholique de Louvain | 2 | 201 | 3835 | 1807 |
| 323 | Tosoh | 1 | 201 | 140 | 19 |
| 323 | Research Triangle Park Foundation | 2 | 201 | 2303 | 1390 |
| 326 | University of Sydney | 10 | 198 | 3584 | 4096 |
| 326 | Claude Bernard University Lyon 1 | 4 | 198 | 2423 | 1542 |
| 328 | Radboud University Nijmegen | 9 | 197 | 1716 | 2167 |
| 329 | Shanghai Institutes for Biological Sciences | 5 | 195 | 382 | 429 |
| 330 | National Institute of Radiological Sciences | 6 | 194 | 266 | 233 |
| 330 | Tor Vergata University of Rome | 3 | 194 | 1190 | 633 |
| 330 | Okayama University | 6 | 194 | 1579 | 1115 |
| 333 | University of Saskatchewan | 3 | 193 | 2956 | 1632 |
| 334 | University of Louisville | 3 | 190 | 2617 | 1431 |
| 335 | Ludwig Maximilian University of Munich | 4 | 189 | 4886 | 3177 |
| 335 | University of Namur | 1 | 189 | 1093 | 311 |
| 337 | Chalmers University of Technology | 1 | 188 | 3491 | 1260 |
| 337 | University of Reading | 3 | 188 | 2333 | 1402 |
| 339 | Simon Fraser University | 7 | 186 | 1299 | 1528 |
| 339 | Stazione Zoologica Anton Dohrn | 7 | 186 | 132 | 191 |
| 339 | Eppendorf (Germany) | 3 | 186 | 1275 | 815 |
| 342 | Kyungpook National University | 13 | 185 | 740 | 1167 |
| 342 | Lariboisière Hospital | 3 | 185 | 512 | 348 |
| 344 | Pompeu Fabra University | 4 | 184 | 941 | 710 |
| 345 | University of Cincinnati | 5 | 183 | 2335 | 1914 |
| 345 | University of East Anglia | 13 | 183 | 599 | 1365 |
| 345 | National Institute of Agrobiological Sciences | 4 | 183 | 210 | 190 |
| 348 | University of Ioannina | 2 | 181 | 1715 | 741 |
| 348 | Southampton General Hospital | 3 | 181 | 587 | 487 |
| 350 | University of California, Los Angeles | 7 | 180 | 1644 | 2082 |
| 351 | London Research Institute | 10 | 178 | 57 | 179 |
| 352 | University of Hong Kong | 13 | 176 | 3127 | 4227 |
| 352 | Frederick National Laboratory for Cancer Research | 2 | 176 | 461 | 312 |
| 354 | University of Rennes 1 | 7 | 174 | 731 | 830 |
| 354 | Lawrence Berkeley National Laboratory | 3 | 174 | 1620 | 1285 |
| 356 | McGill University Health Centre | 4 | 172 | 977 | 761 |
| 357 | University of Barcelona | 14 | 170 | 1627 | 2691 |
| 357 | George Washington University | 3 | 170 | 3200 | 1892 |
| 357 | Anschutz Medical Campus | 3 | 170 | 1404 | 1049 |
| 357 | Max Planck Institute of Experimental Endocrinology | 4 | 170 | 2 | 6 |
| 357 | University of Rochester Medical Center | 4 | 170 | 1132 | 967 |
| 362 | Université de Lyon | 5 | 169 | 1201 | 1097 |
| 363 | University of Southampton | 3 | 167 | 4066 | 2616 |
| 363 | Oak Ridge National Laboratory | 1 | 167 | 2627 | 1192 |
| 363 | Department of Public Health | 3 | 167 | 5194 | 2915 |
| 366 | University of Salzburg | 2 | 165 | 1166 | 503 |
| 366 | Michigan State University | 3 | 165 | 5539 | 3446 |
| 368 | University of Tromso | 1 | 164 | 169 | 71 |
| 369 | Anthropologie | 2 | 162 | 349 | 164 |
| 370 | National Institute for Physiological Sciences | 2 | 161 | 252 | 162 |
| 371 | Federal University of Rio de Janeiro | 10 | 160 | 1897 | 1902 |
| 372 | University Hospital Zurich | 1 | 157 | 1947 | 793 |
| 372 | International Agency for Research on Cancer | 1 | 157 | 603 | 340 |
| 372 | Instituto Biológico | 2 | 157 | 488 | 220 |
| 372 | National Institutes of Natural Sciences, Japan | 5 | 157 | 106 | 124 |
| 376 | University of Tartu | 4 | 156 | 1210 | 961 |
| 377 | BC Cancer Agency | 2 | 155 | 614 | 433 |
| 378 | University of Oslo | 3 | 154 | 4570 | 2986 |
| 379 | Nagoya City University | 1 | 152 | 1829 | 594 |
| 380 | Semmelweis University | 3 | 151 | 1329 | 879 |
| 380 | WiCell | 1 | 151 | 15 | 20 |
| 382 | Atomic Energy Research Establishment | 11 | 149 | 259 | 499 |
| 382 | Maastricht University | 1 | 149 | 4424 | 1952 |
| 382 | MRC Institute of Hearing Research | 2 | 149 | 58 | 49 |
| 382 | Mammalian Genetics Unit | 7 | 149 | 13 | 59 |
| 386 | University of Notre Dame Australia | 3 | 147 | 2750 | 1740 |
| 386 | Centre for Cellular and Molecular Biology | 11 | 147 | 39 | 206 |
| 388 | École des Sciences de I'information | 1 | 146 | 796 | 177 |
| 389 | University of Hyogo | 5 | 145 | 487 | 314 |
| 390 | Nebraska Medical Center | 2 | 144 | 2152 | 1164 |
| 391 | University of Delaware | 2 | 143 | 3340 | 1861 |
| 391 | Cooperative Research Centre | 2 | 143 | 952 | 764 |
| 393 | Wistar Institute | 4 | 141 | 257 | 328 |
| 393 | interdisciplinarity | 7 | 141 | 4174 | 4078 |
| 395 | University of Algarve | 2 | 139 | 942 | 445 |
| 396 | University of Massachusetts Amherst | 3 | 138 | 1327 | 1008 |
| 396 | National Yang-Ming University | 6 | 138 | 936 | 1073 |
| 396 | Tottori University | 2 | 138 | 1458 | 617 |
| 399 | Cold Spring Harbor Laboratory | 2 | 137 | 300 | 294 |
| 399 | National Human Genome Research Institute | 7 | 137 | 175 | 369 |
| 399 | University of Kansas Medical Center | 2 | 137 | 1682 | 956 |
| 402 | Austrian Academy of Sciences | 7 | 135 | 311 | 593 |
| 402 | University of Alberta | 9 | 135 | 3238 | 4285 |
| 404 | University of Strasbourg | 4 | 134 | 1601 | 1310 |
| 404 | Necker–Enfants Malades Hospital | 5 | 134 | 579 | 595 |
| 404 | University of Nevada, Las Vegas | 3 | 134 | 436 | 314 |
| 407 | University of Jena | 2 | 133 | 2308 | 1296 |
| 407 | TU Dresden | 8 | 133 | 1588 | 2074 |
| 407 | Region of Southern Denmark | 1 | 133 | 4252 | 1772 |
| 410 | University of Buenos Aires | 8 | 132 | 1568 | 1586 |
| 410 | University of Houston | 3 | 132 | 2868 | 1951 |
| 410 | Peking Union Medical College | 3 | 132 | 2598 | 1810 |
| 413 | Food and Drug Administration | 1 | 130 | 2226 | 1076 |
| 413 | Institut Pasteur de Lille | 2 | 130 | 511 | 361 |
| 415 | Novartis | 3 | 129 | 1259 | 1109 |
| 415 | University of Regensburg | 4 | 129 | 1568 | 1416 |
| 417 | Royal Netherlands Academy of Arts and Sciences | 8 | 128 | 76 | 270 |
| 418 | Kansai Medical University | 2 | 127 | 826 | 404 |
| 418 | California State University, Chico | 2 | 127 | 59 | 37 |
| 418 | Cancer Hospital and Research Institute | 2 | 127 | 65 | 24 |
| 421 | National Centre for Biological Sciences | 8 | 126 | 50 | 146 |
| 421 | University of Florida | 4 | 126 | 6167 | 4913 |
| 421 | University of New South Wales | 6 | 126 | 4026 | 4287 |
| 421 | Victor Chang Cardiac Research Institute | 10 | 126 | 33 | 139 |
| 425 | Heriot-Watt University | 3 | 125 | 1345 | 965 |
| 425 | Aix-Marseille University | 3 | 125 | 2747 | 1867 |
| 427 | Kanazawa University | 3 | 124 | 2052 | 1276 |
| 427 | Max Planck Institute of Biochemistry | 1 | 124 | 369 | 237 |
| 429 | Jagiellonian University | 5 | 123 | 1968 | 1607 |
| 429 | National Autonomous University of Mexico | 8 | 123 | 2058 | 2091 |
| 431 | National Institute of Environmental Health Sciences | 1 | 122 | 1118 | 685 |
| 431 | Friedrich Miescher Institute for Biomedical Research | 6 | 122 | 80 | 188 |
| 431 | University Hospitals of Cleveland | 2 | 122 | 3447 | 2020 |
| 434 | Falkenried | 3 | 121 | 10 | 36 |
| 435 | University of Tübingen | 7 | 120 | 1510 | 2135 |
| 435 | NOVA University Lisbon | 3 | 120 | 1546 | 1109 |
| 435 | Vollum Institute | 2 | 120 | 131 | 157 |
| 438 | UCL Institute of Ophthalmology | 4 | 119 | 123 | 187 |
| 439 | Tokyo Women's Medical University | 7 | 118 | 493 | 596 |
| 440 | Vrije Universiteit Amsterdam | 3 | 117 | 5174 | 4025 |
| 440 | State University of New York Upstate Medical University | 3 | 117 | 1071 | 824 |
| 440 | Miami Project to Cure Paralysis | 1 | 117 | 178 | 110 |
| 443 | Francis Crick Institute | 18 | 116 | 56 | 273 |
| 443 | University of Texas System | 2 | 116 | 1994 | 1421 |
| 445 | University of Teramo | 3 | 115 | 202 | 207 |
| 445 | Radcliffe Infirmary | 1 | 115 | 531 | 327 |
| 445 | MetroHealth | 1 | 115 | 885 | 403 |
| 445 | Instituto de Biología Molecular y Celular de Rosario | 2 | 115 | 84 | 66 |
| 449 | University of Turin | 2 | 114 | 4075 | 2449 |
| 449 | Australian Museums and Galleries Association | 3 | 114 | 134 | 169 |
| 451 | Miguel Hernández University of Elche | 6 | 113 | 151 | 279 |
| 451 | Loyola University | 2 | 113 | 2541 | 1408 |
| 451 | Monash University | 16 | 113 | 1960 | 4578 |
| 451 | Veterans Health Administration | 2 | 113 | 2580 | 1902 |
| 451 | National Institute of Diabetes and Digestive and Kidney Diseases | 2 | 113 | 621 | 570 |
| 456 | Northumbria University | 1 | 112 | 2691 | 911 |
| 456 | Cochin Institute | 2 | 112 | 35 | 51 |
| 458 | Kaohsiung Medical University | 1 | 111 | 2106 | 895 |
| 458 | National Academy of Medicine | 1 | 111 | 2701 | 1128 |
| 460 | University of Auckland | 5 | 110 | 2986 | 3287 |
| 461 | University of Bristol | 10 | 108 | 1953 | 3445 |
| 461 | Yamaguchi University | 4 | 108 | 1215 | 906 |
| 463 | Indian Institute of Science | 10 | 107 | 911 | 1449 |
| 464 | Instituto de Medicina Molecular | 9 | 106 | 78 | 279 |
| 464 | Harvard Stem Cell Institute | 2 | 106 | 183 | 204 |
| 466 | University of Giessen | 4 | 105 | 1849 | 1870 |
| 466 | Shiga University of Medical Science | 2 | 105 | 872 | 498 |
| 466 | Rollins College | 1 | 105 | 1855 | 929 |
| 466 | Ottawa Hospital Research Institute | 2 | 105 | 988 | 729 |
| 466 | Weatherall Institute of Molecular Medicine | 6 | 105 | 139 | 334 |
| 466 | Université de Paris | 2 | 105 | 2625 | 1512 |
| 472 | Université de Montréal | 3 | 102 | 4285 | 3387 |
| 472 | University of Medicine and Dentistry of New Jersey | 1 | 102 | 1502 | 991 |
| 472 | Bio-oriented Technology Research Advancement Institution | 1 | 102 | 46 | 34 |
| 472 | Food Research Institute | 1 | 102 | 1125 | 637 |
| 476 | Purdue University | 3 | 101 | 5670 | 4392 |
| 476 | Okinawa Institute of Science and Technology | 3 | 101 | 274 | 258 |
| 476 | National Institute of Child Health | 3 | 101 | 515 | 589 |
| 476 | Royal Women's Hospital | 2 | 101 | 437 | 346 |
| 476 | Okinawa Prefecture | 3 | 101 | 1345 | 981 |
| 481 | Russian Academy of Sciences | 5 | 100 | 4327 | 3155 |
| 481 | Chonbuk National University | 1 | 100 | 2073 | 989 |
| 481 | University of Lisbon | 6 | 100 | 2366 | 2706 |
| 481 | MEDICA | 2 | 100 | 2358 | 1558 |
| 485 | University of Kansas | 4 | 99 | 2644 | 2562 |
| 485 | Western Washington University | 2 | 99 | 657 | 404 |
| 485 | San Diego State University | 2 | 99 | 2549 | 1681 |
| 485 | Temple University | 2 | 99 | 3861 | 2498 |
| 485 | Monash Medical Centre | 2 | 99 | 521 | 427 |
| 485 | Centre for Genomic Regulation | 4 | 99 | 116 | 204 |
| 491 | University of Gothenburg | 2 | 98 | 4764 | 3382 |
| 491 | Kawasaki Medical School | 4 | 98 | 432 | 387 |
| 491 | USC Norris Cancer Hospital | 1 | 98 | 7 | 3 |
| 491 | Chinese University of Hong Kong | 11 | 98 | 1731 | 3439 |
| 495 | University of Málaga | 2 | 97 | 1875 | 933 |
| 495 | University of Leicester | 3 | 97 | 2744 | 2339 |
| 495 | Robarts Research Institute | 2 | 97 | 439 | 380 |
| 495 | Halifax | 5 | 97 | 2495 | 3014 |
| 499 | University of Mississippi | 1 | 95 | 2659 | 1099 |
| 499 | Nihon University | 3 | 95 | 1986 | 1273 |
| 499 | MRC Laboratory for Molecular Cell Biology | 3 | 95 | 55 | 106 |
| 502 | University of São Paulo | 11 | 94 | 3839 | 4810 |
| 503 | Kindai University | 1 | 93 | 2631 | 1034 |
| 503 | Laval University | 4 | 93 | 2961 | 3036 |
| 505 | Erasmus MC | 2 | 92 | 2423 | 1957 |
| 505 | National Institute of Allergy and Infectious Diseases | 2 | 92 | 831 | 757 |
| 505 | University of Nevada, Reno | 2 | 92 | 3464 | 2181 |
| 508 | University of Illinois at Urbana–Champaign | 2 | 91 | 5030 | 3567 |
| 508 | Hong Kong Polytechnic University | 1 | 91 | 4848 | 2525 |
| 510 | University of California, Irvine | 3 | 90 | 1777 | 1830 |
| 511 | Hospital Real | 1 | 89 | 1423 | 695 |
| 512 | University of Central Florida | 3 | 88 | 2301 | 1984 |
| 512 | Chungnam National University | 2 | 88 | 2139 | 1366 |
| 512 | University of Putra Malaysia | 1 | 88 | 3692 | 1471 |
| 512 | Herman Ostrow School of Dentistry of USC | 2 | 88 | 47 | 62 |
| 512 | Houston Methodist Hospital | 4 | 88 | 307 | 545 |
| 512 | University Health Network | 2 | 88 | 2090 | 1634 |
| 518 | Royal Veterinary College | 4 | 85 | 356 | 499 |
| 518 | Biology Institute of Shandong Academy of Sciences | 7 | 85 | 1354 | 1920 |
| 518 | Scuola Normale Superiore di Pisa | 4 | 85 | 359 | 419 |
| 521 | Huntsman Cancer Institute | 1 | 84 | 694 | 433 |
| 521 | Ottawa Civic Hospital | 1 | 84 | 198 | 133 |
| 523 | Aarhus University | 2 | 83 | 5163 | 3686 |
| 523 | Cleveland Clinic | 2 | 83 | 2906 | 2147 |
| 525 | National Institute on Aging | 5 | 82 | 442 | 889 |
| 525 | list of women's colleges | 3 | 82 | 1408 | 1110 |
| 527 | Osaka City University | 3 | 81 | 1557 | 1226 |
| 527 | St George's, University of London | 1 | 81 | 561 | 399 |
| 529 | Reichman University | 6 | 80 | 540 | 1243 |
| 529 | Luther University | 2 | 80 | 1076 | 925 |
| 531 | Barcelona Institute for Science and Technology | 1 | 79 | 679 | 432 |
| 531 | Ajuntament de L’Hospitalet | 2 | 79 | 993 | 808 |
| 533 | Charité | 3 | 78 | 2707 | 2556 |
| 533 | University of Siena | 3 | 78 | 1630 | 1659 |
| 533 | University of Portsmouth | 8 | 78 | 415 | 1230 |
| 533 | Department of Biotechnology | 4 | 78 | 3203 | 3062 |
| 537 | Taipei Medical University | 5 | 77 | 971 | 1262 |
| 537 | University of Yamanashi | 2 | 77 | 1259 | 778 |
| 537 | Indian Institutes of Science Education and Research | 11 | 77 | 179 | 460 |
| 537 | Murdoch Children's Research Institute | 9 | 77 | 351 | 884 |
| 541 | Queen Mary University of London | 8 | 76 | 1227 | 2556 |
| 541 | Hôpital Saint Vincent de Paul | 1 | 76 | 111 | 85 |
| 541 | Universidad Nacional de Rosario | 1 | 76 | 1328 | 399 |
| 544 | Westfield College | 2 | 75 | 523 | 495 |
| 544 | Cancer Research Institute | 3 | 75 | 1203 | 1223 |
| 546 | University of Bonn | 3 | 74 | 3192 | 3052 |
| 546 | University of Szeged | 4 | 74 | 1086 | 1259 |
| 546 | Iwate University | 1 | 74 | 1136 | 459 |
| 549 | Georgetown University | 2 | 73 | 3106 | 2256 |
| 549 | Tampere University Hospital | 1 | 73 | 1314 | 894 |
| 551 | Tsinghua University | 12 | 72 | 2365 | 4031 |
| 551 | Moorfields Eye Hospital | 1 | 72 | 434 | 275 |
| 551 | Universidad Nacional del Sur | 1 | 72 | 1200 | 393 |
| 554 | Wellcome Sanger Institute | 6 | 71 | 236 | 520 |
| 554 | University of California, Berkeley | 4 | 71 | 1859 | 2778 |
| 554 | Korea University of Science and Technology | 14 | 71 | 5754 | 8491 |
| 554 | University of Maryland, Baltimore County | 2 | 71 | 657 | 679 |
| 558 | University of Extremadura | 3 | 70 | 1188 | 1065 |
| 558 | Sophia University | 2 | 70 | 581 | 355 |
| 558 | University of Alaska Anchorage | 2 | 70 | 247 | 231 |
| 558 | Goodwin College | 2 | 70 | 493 | 598 |
| 558 | Wellcome Genome Campus | 1 | 70 | 495 | 445 |
| 558 | Institute For Ecology of Industrial Areas | 1 | 70 | 45 | 30 |
| 564 | University of Leeds | 9 | 69 | 2265 | 4369 |
| 564 | Medical University of Innsbruck | 1 | 69 | 1042 | 760 |
| 564 | Oita University | 1 | 69 | 1432 | 662 |
| 564 | United States Department of Veterans Affairs | 2 | 69 | 2891 | 2509 |
| 568 | University of Paris 1 Pantheon-Sorbonne | 3 | 68 | 1160 | 1374 |
| 569 | University of Milan | 5 | 67 | 3428 | 4188 |
| 569 | Nippon Medical School | 3 | 67 | 939 | 807 |
| 569 | Universidad de Cantabria | 3 | 67 | 600 | 609 |
| 572 | Amgen | 1 | 66 | 706 | 577 |
| 572 | SUNY Downstate Medical Center | 1 | 66 | 1771 | 1035 |
| 572 | Downing Site | 3 | 66 | 34 | 134 |
| 572 | Life Sciences Institute | 5 | 66 | 260 | 632 |
| 572 | Asahikawa Medical College | 1 | 66 | 518 | 328 |
| 577 | Carnegie Mellon University | 3 | 65 | 2390 | 2624 |
| 577 | Waseda University | 3 | 65 | 1999 | 1656 |
| 577 | Biozentrum University of Basel | 1 | 65 | 52 | 69 |
| 577 | Florey Institute of Neuroscience and Mental Health | 3 | 65 | 413 | 599 |
| 581 | Charles University | 4 | 64 | 2990 | 3092 |
| 581 | University of Crete | 5 | 64 | 882 | 1626 |
| 581 | Kitasato University | 3 | 64 | 1361 | 1211 |
| 581 | Methodist University Hospital | 3 | 64 | 859 | 918 |
| 581 | art school | 1 | 64 | 2568 | 959 |
| 586 | Ochanomizu University | 3 | 63 | 356 | 343 |
| 587 | Addenbrooke's Hospital | 3 | 62 | 1154 | 1415 |
| 587 | Universidad Nacional del Litoral | 1 | 62 | 919 | 374 |
| 587 | Estonian Biocentre | 1 | 62 | 83 | 83 |
| 587 | Tartu University Clinic | 1 | 62 | 335 | 218 |
| 587 | Kurume University | 1 | 62 | 1367 | 834 |
| 587 | Instituto de Biología y Genética Molecular | 3 | 62 | 45 | 85 |
| 587 | Instituto de Agrobiotecnología del Litoral | 1 | 62 | 47 | 38 |
| 594 | Wyeth | 1 | 61 | 258 | 236 |
| 594 | Jacobi Medical Center | 1 | 61 | 429 | 243 |
| 594 | Robert H. Lurie Comprehensive Cancer Center | 2 | 61 | 359 | 479 |
| 594 | Broadway | 3 | 61 | 1723 | 2305 |
| 598 | Polish Academy of Sciences | 5 | 60 | 2702 | 3256 |
| 598 | Shizuoka University | 5 | 60 | 693 | 864 |
| 598 | University of Utah Hospital | 1 | 60 | 382 | 268 |
| 598 | Gembloux Agro-Bio Tech | 1 | 60 | 498 | 336 |
| 598 | Sorbonne Universités | 7 | 60 | 520 | 1478 |
| 603 | University of Texas at San Antonio | 1 | 59 | 2625 | 1503 |
| 603 | Stockholm University | 2 | 59 | 3756 | 3227 |
| 603 | University of Cantabria | 1 | 59 | 1009 | 665 |
| 603 | Montana State University | 3 | 59 | 1191 | 1533 |
| 603 | Royal Brisbane and Women's Hospital | 1 | 59 | 1428 | 1011 |
| 608 | National Taiwan University | 9 | 58 | 2365 | 4278 |
| 608 | Max Planck Institute for Medical Research | 1 | 58 | 164 | 150 |
| 608 | University of Turku | 2 | 58 | 3405 | 2892 |
| 608 | California State University | 3 | 58 | 3171 | 3030 |
| 608 | Observatoire océanologique de Banyuls-sur-Mer | 1 | 58 | 132 | 110 |
| 608 | Ripon College | 1 | 58 | 49 | 26 |
| 608 | Royal Institute of Public Health | 2 | 58 | 770 | 761 |
| 608 | Rodale, Inc. | 3 | 58 | 1652 | 1652 |
| 616 | Pablo de Olavide University | 6 | 57 | 237 | 634 |
| 616 | Max Planck Institute of Psychiatry | 3 | 57 | 279 | 469 |
| 616 | MRC Toxicology Unit | 1 | 57 | 202 | 186 |
| 619 | Nagoya Institute of Technology | 1 | 56 | 1325 | 670 |
| 619 | University of Toledo | 1 | 56 | 2839 | 1693 |
| 619 | The Ottawa Hospital | 1 | 56 | 1307 | 879 |
| 619 | Sunnybrook Health Sciences Centre | 1 | 56 | 1858 | 1364 |
| 619 | Michaels | 2 | 56 | 327 | 403 |
| 619 | Penn State College of Medicine | 2 | 56 | 690 | 672 |
| 625 | Seoul National University | 4 | 55 | 4544 | 4914 |
| 625 | Hungarian Academy of Sciences | 1 | 55 | 3231 | 2225 |
| 625 | Karolinska University Hospital | 2 | 55 | 2292 | 2262 |
| 625 | Autonomous University of Queretaro | 3 | 55 | 308 | 287 |
| 625 | Ninewells Hospital | 1 | 55 | 1190 | 923 |
| 630 | University of Halle-Wittenberg | 1 | 54 | 1572 | 1043 |
| 630 | University of Valencia | 3 | 54 | 2578 | 2715 |
| 632 | Central University of Venezuela | 1 | 53 | 1292 | 614 |
| 632 | University of California, Davis | 2 | 53 | 3868 | 3670 |
| 634 | University of Connecticut | 5 | 52 | 2947 | 4435 |
| 634 | University of Vienna | 7 | 52 | 3483 | 5512 |
| 634 | Teikyo University | 2 | 52 | 1404 | 1139 |
| 634 | Mount Saint Vincent University | 3 | 52 | 132 | 181 |
| 634 | House Ear Institute | 2 | 52 | 63 | 117 |
| 634 | Holdsworth | 1 | 52 | 195 | 208 |
| 634 | Iona College | 1 | 52 | 185 | 109 |
| 641 | University of Bologna | 2 | 51 | 5647 | 4773 |
| 641 | University of Toulouse | 2 | 51 | 3099 | 2753 |
| 641 | Florida State University | 3 | 51 | 3124 | 3432 |
| 641 | Royal College of Physicians and Surgeons of Canada | 1 | 51 | 826 | 548 |
| 641 | National Cancer Center | 2 | 51 | 1768 | 1516 |
| 641 | Kanagawa Cancer Center | 1 | 51 | 315 | 190 |
| 647 | Institute for Research in Biomedicine | 4 | 50 | 467 | 803 |
| 647 | University of California, Santa Barbara | 2 | 50 | 1267 | 1470 |
| 649 | Ocean University of China | 3 | 49 | 1342 | 1485 |
| 649 | Deakin University | 2 | 49 | 3652 | 3498 |
| 649 | Oklahoma Medical Research Foundation | 1 | 49 | 526 | 477 |
| 649 | Robert Larner College of Medicine | 1 | 49 | 1066 | 797 |
| 653 | Wellesley College | 1 | 48 | 701 | 537 |
| 653 | Texas Tech University | 2 | 48 | 3172 | 2819 |
| 653 | Genzyme | 1 | 48 | 380 | 407 |
| 653 | Joslin Diabetes Center | 1 | 48 | 447 | 474 |
| 657 | Istituto Giannina Gaslini | 1 | 47 | 664 | 531 |
| 657 | University Hospital Heidelberg | 2 | 47 | 1184 | 1199 |
| 657 | Bar-Ilan University | 3 | 47 | 2014 | 2335 |
| 657 | National University of Córdoba | 1 | 47 | 350 | 204 |
| 657 | Lexicon Pharmaceuticals | 1 | 47 | 82 | 95 |
| 657 | Showa University | 1 | 47 | 1723 | 1108 |
| 657 | Doheny Eye Institute | 1 | 47 | 162 | 179 |
| 664 | Florida Atlantic University | 3 | 46 | 1309 | 1752 |
| 664 | University of Guelph | 3 | 46 | 2435 | 2973 |
| 664 | Middlesex Hospital | 1 | 46 | 584 | 493 |
| 664 | Memorial University of Newfoundland | 3 | 46 | 2011 | 2545 |
| 664 | Atlantic University | 3 | 46 | 1345 | 1783 |
| 664 | Dupont Hospital | 1 | 46 | 243 | 230 |
| 670 | Saint Louis University | 1 | 45 | 3157 | 2186 |
| 670 | Gregor Mendel Institute | 1 | 45 | 82 | 100 |
| 670 | Royal Military College | 2 | 45 | 457 | 411 |
| 670 | Royal Military College of Canada | 2 | 45 | 372 | 357 |
| 670 | Seto Marine Biological Laboratory | 1 | 45 | 37 | 23 |
| 670 | Wellcome Trust Centre for Cell-Matrix Research | 2 | 45 | 71 | 200 |
| 676 | Technical University of Munich | 1 | 44 | 2378 | 1848 |
| 676 | University of Kassel | 2 | 44 | 1140 | 1123 |
| 676 | Kwansei Gakuin University | 4 | 44 | 282 | 406 |
| 679 | Tel Aviv University | 4 | 43 | 4202 | 5209 |
| 679 | University of Montpellier 1 | 1 | 43 | 225 | 321 |
| 679 | University of New Brunswick | 1 | 43 | 2313 | 1558 |
| 679 | Southern Methodist University | 1 | 43 | 1850 | 1326 |
| 679 | Methodist University | 1 | 43 | 1975 | 1376 |
| 684 | University of British Columbia | 7 | 42 | 4581 | 7577 |
| 684 | University of Detroit Mercy | 1 | 42 | 463 | 233 |
| 684 | National Centre of Scientific Research "DEMOKRITOS" | 1 | 42 | 1237 | 927 |
| 684 | RIKEN Quantitative Biology Center | 3 | 42 | 15 | 64 |
| 688 | Duke University | 4 | 41 | 4238 | 5639 |
| 688 | Australian Regenerative Medicine Institute | 7 | 41 | 18 | 152 |
| 688 | Georgetown University Medical Center | 2 | 41 | 952 | 1259 |
| 688 | The Alfred Hospital | 2 | 41 | 1626 | 1790 |
| 692 | University of Glasgow | 3 | 40 | 4369 | 5077 |
| 692 | University of Alabama | 1 | 40 | 5487 | 3939 |
| 692 | Banaras Hindu University | 7 | 40 | 1057 | 2243 |
| 692 | University of Arkansas for Medical Sciences | 1 | 40 | 2601 | 2052 |
| 692 | Gifu University | 1 | 40 | 2724 | 1800 |
| 692 | University of Paris-Sud | 3 | 40 | 2293 | 3034 |
| 692 | Republic of China National Defense Medical Center | 2 | 40 | 1123 | 1166 |
| 699 | Central Research Laboratories | 1 | 39 | 732 | 600 |
| 699 | Section of Painting and Sculpture | 1 | 39 | 574 | 559 |
| 701 | Virginia Tech | 3 | 38 | 3977 | 4844 |
| 701 | Astellas Pharma | 1 | 38 | 658 | 562 |
| 701 | Hamamatsu University | 2 | 38 | 1018 | 993 |
| 701 | Wolfson Centre for Age-Related Diseases | 1 | 38 | 137 | 195 |
| 705 | University of Vermont | 1 | 37 | 3713 | 2831 |
| 705 | Sophia Children's Hospital | 1 | 37 | 876 | 826 |
| 705 | Istituto per le Applicazioni del Calcolo Mauro Picone | 1 | 37 | 469 | 344 |
| 705 | Princess Margaret Hospital | 1 | 37 | 1029 | 977 |
| 705 | Nara Medical University | 2 | 37 | 963 | 937 |
| 705 | Ontario Cancer Institute | 1 | 37 | 522 | 581 |
| 705 | Population Council | 1 | 37 | 398 | 367 |
| 705 | The Neurosciences Institute | 2 | 37 | 293 | 488 |
| 713 | University of Kiel | 2 | 36 | 1860 | 2290 |
| 713 | Creighton University | 2 | 36 | 1152 | 1399 |
| 713 | University of the Sacred Heart | 1 | 36 | 1025 | 894 |
| 713 | University of Maine | 2 | 36 | 1431 | 1727 |
| 713 | Kyoto Pharmaceutical University | 1 | 36 | 438 | 378 |
| 713 | Wellcome Trust Centre for Stem Cell Research | 2 | 36 | 16 | 53 |
| 713 | MRC London Institute of Medical Sciences | 7 | 36 | 89 | 457 |
| 713 | Babcock University | 1 | 36 | 576 | 414 |
| 713 | Instituto Tecnológico de Chascomús | 1 | 36 | 147 | 177 |
| 722 | Akdeniz University | 1 | 35 | 2044 | 1325 |
| 722 | Witten/Herdecke University | 2 | 35 | 784 | 1010 |
| 722 | University of Liverpool | 6 | 35 | 2944 | 5452 |
| 722 | Galton Laboratory | 1 | 35 | 126 | 144 |
| 722 | Istanbul Medipol University | 1 | 35 | 1073 | 491 |
| 722 | Université de Rouen | 1 | 35 | 1098 | 896 |
| 722 | Universidad de Estudios Avanzados | 2 | 35 | 43 | 85 |
| 729 | University of Manchester | 8 | 34 | 834 | 3139 |
| 729 | Helmholtz Zentrum München | 2 | 34 | 882 | 1269 |
| 729 | Hong Kong University of Science and Technology | 2 | 34 | 1297 | 1610 |
| 729 | University of Campinas | 2 | 34 | 5455 | 4275 |
| 729 | Virginia Commonwealth University | 1 | 34 | 5111 | 4029 |
| 729 | Wakayama Medical University | 2 | 34 | 734 | 783 |
| 729 | UC San Diego School of Medicine | 1 | 34 | 763 | 813 |
| 729 | Research Institute of Evolutionary Biology | 1 | 34 | 16 | 14 |
| 729 | Chiba Cancer Center | 1 | 34 | 279 | 231 |
| 738 | Universidad Mayor | 1 | 33 | 602 | 403 |
| 738 | Daegu University | 1 | 33 | 817 | 686 |
| 738 | University of Florence | 1 | 33 | 5158 | 4028 |
| 738 | University Hospital Erlangen | 1 | 33 | 890 | 803 |
| 738 | National Institute of Dental and Craniofacial Research | 1 | 33 | 584 | 677 |
| 738 | Santos | 1 | 33 | 2233 | 1431 |
| 744 | Lehigh University | 1 | 32 | 2275 | 1791 |
| 744 | Franklin & Marshall College | 2 | 32 | 275 | 432 |
| 744 | Marshall University | 2 | 32 | 796 | 1125 |
| 744 | University of Texas Health Science Center at Houston | 1 | 32 | 2807 | 2348 |
| 744 | Instituto de Química y Fisicoquímica Biológicas | 1 | 32 | 164 | 166 |
| 744 | Institute for Biomedical Research and Innovation | 1 | 32 | 95 | 93 |
| 750 | University of Paris | 1 | 31 | 4164 | 3416 |
| 751 | University of Stuttgart | 1 | 30 | 3059 | 2512 |
| 751 | North Carolina Central University | 1 | 30 | 511 | 441 |
| 751 | New College of Florida | 1 | 30 | 122 | 130 |
| 751 | Savitribai Phule Pune University | 1 | 30 | 1659 | 1227 |
| 751 | Rigshospitalet | 1 | 30 | 2231 | 2031 |
| 751 | Center for Research and Advanced Studies of the National Polytechnic Institute | 2 | 30 | 1370 | 1610 |
| 751 | Abasaheb Garware College | 1 | 30 | 83 | 74 |
| 758 | Hochschule Hannover | 1 | 29 | 615 | 537 |
| 758 | University of Caen Normandy | 1 | 29 | 1742 | 1574 |
| 758 | Ben-Gurion University of the Negev | 2 | 29 | 3545 | 4040 |
| 758 | Ghent University | 2 | 29 | 5993 | 6241 |
| 758 | Akita University | 2 | 29 | 1161 | 1152 |
| 758 | Gakushuin University | 2 | 29 | 279 | 350 |
| 758 | IFREMER | 1 | 29 | 934 | 940 |
| 758 | Bharathidasan University | 1 | 29 | 999 | 780 |
| 758 | Benaroya Research Institute | 1 | 29 | 204 | 275 |
| 758 | Chubu University | 2 | 29 | 509 | 589 |
| 768 | University of Georgia | 1 | 28 | 6660 | 5445 |
| 768 | University of León | 1 | 28 | 1325 | 1059 |
| 768 | Kyorin University | 3 | 28 | 597 | 889 |
| 768 | Broad Institute | 1 | 28 | 951 | 1014 |
| 768 | Imperial College School of Medicine | 1 | 28 | 887 | 1010 |
| 768 | Pirbright Institute | 1 | 28 | 627 | 640 |
| 768 | California State Polytechnic University, Pomona | 1 | 28 | 1693 | 998 |
| 775 | University of Zaragoza | 2 | 27 | 2988 | 3128 |
| 775 | National Chung Cheng University | 1 | 27 | 1347 | 1098 |
| 775 | Scripps Research | 1 | 27 | 1549 | 1627 |
| 775 | Konan University | 3 | 27 | 223 | 361 |
| 775 | Bennett College | 1 | 27 | 597 | 644 |
| 775 | Hyogo College of Medicine | 2 | 27 | 837 | 980 |
| 775 | National University Hospital | 1 | 27 | 2903 | 2354 |
| 775 | Roy J. and Lucille A. Carver College of Medicine | 1 | 27 | 1426 | 1433 |
| 775 | Simpson College | 1 | 27 | 27 | 20 |
| 775 | Megmilk Snow Brand | 1 | 27 | 126 | 125 |
| 775 | Yong Loo Lin School of Medicine | 2 | 27 | 1541 | 1976 |
| 775 | Smith-Kettlewell Eye Research Institute | 1 | 27 | 836 | 950 |
| 775 | Centre for Biomedical Network Research on Rare Diseases | 2 | 27 | 482 | 722 |
| 788 | Scripps Institution of Oceanography | 2 | 26 | 738 | 1075 |
| 788 | University of Cyprus | 1 | 26 | 1859 | 1669 |
| 788 | University of Windsor | 2 | 26 | 1628 | 2115 |
| 788 | Miyagi Prefectural Hospital Organization | 1 | 26 | 130 | 150 |
| 788 | University at Albany | 1 | 26 | 2263 | 2160 |
| 793 | Sun Yat-sen University | 2 | 25 | 5727 | 5791 |
| 793 | University of Colorado Boulder | 1 | 25 | 3353 | 3136 |
| 793 | National Health Research Institutes | 3 | 25 | 505 | 1003 |
| 796 | Complutense University of Madrid | 2 | 24 | 5240 | 5104 |
| 796 | Max Planck Institute for Molecular Biomedicine | 4 | 24 | 42 | 179 |
| 796 | Universiti Malaysia Terengganu | 1 | 24 | 817 | 666 |
| 796 | Boston Biomedical Research Institute | 1 | 24 | 129 | 183 |
| 796 | Protein Sciences | 3 | 24 | 135 | 377 |
| 796 | UNC Lineberger Comprehensive Cancer Center | 1 | 24 | 848 | 987 |
| 802 | KAIST | 1 | 23 | 2728 | 2515 |
| 802 | University of Louisiana at Lafayette | 1 | 23 | 1208 | 1141 |
| 802 | St Bartholomew's Hospital | 1 | 23 | 1230 | 1245 |
| 802 | German Primate Center | 2 | 23 | 157 | 347 |
| 802 | Sichuan University | 2 | 23 | 4739 | 4872 |
| 802 | University of America | 1 | 23 | 919 | 806 |
| 802 | Georgetown Lombardi Comprehensive Cancer Center | 1 | 23 | 595 | 758 |
| 802 | Translationale Onkologie an der Universitätsmedizin der Johannes Gutenberg-Universität Mainz | 1 | 23 | 534 | 582 |
| 810 | Almirall | 1 | 22 | 131 | 162 |
| 810 | Beijing University of Chinese Medicine | 1 | 22 | 995 | 956 |
| 810 | Saga University | 1 | 22 | 1737 | 1414 |
| 810 | San Raffaele Hospital | 1 | 22 | 1131 | 1126 |
| 810 | University of Campania "Luigi Vanvitelli" | 1 | 22 | 1500 | 1430 |
| 810 | University of Texas Health Science Center at San Antonio | 1 | 22 | 2316 | 2354 |
| 810 | Becker College | 1 | 22 | 2 | 4 |
| 810 | Fujita Health University | 1 | 22 | 1409 | 1228 |
| 810 | Susquehanna University | 2 | 22 | 68 | 191 |
| 819 | Fudan University | 4 | 21 | 3813 | 5413 |
| 819 | Inselspital | 1 | 21 | 2025 | 1930 |
| 819 | Nanjing University | 3 | 21 | 3967 | 5053 |
| 819 | University of Veterinary Medicine Vienna | 1 | 21 | 539 | 625 |
| 819 | Pharmacia | 2 | 21 | 400 | 728 |
| 819 | Coventry University | 1 | 21 | 6801 | 5282 |
| 819 | University of North Carolina at Greensboro | 1 | 21 | 1903 | 1906 |
| 819 | New York Medical College | 1 | 21 | 1823 | 1776 |
| 819 | Instituto de Biomedicina y Biotecnología de Cantabria | 1 | 21 | 83 | 114 |
| 819 | Mackay Memorial Hospital | 1 | 21 | 1002 | 1030 |
| 819 | Earlham Institute | 2 | 21 | 80 | 187 |
| 819 | Universidad Austral | 1 | 21 | 353 | 385 |
| 819 | Upjohn | 2 | 21 | 422 | 673 |
| 819 | Norwich Research Park | 2 | 21 | 702 | 1338 |
| 833 | Moscow State University | 3 | 20 | 3245 | 3737 |
| 833 | Oakland University | 1 | 20 | 2157 | 2090 |
| 833 | Universidad Veracruzana | 1 | 20 | 1336 | 899 |
| 833 | Odense University Hospital | 1 | 20 | 1611 | 1661 |
| 833 | Chinese Academy of Fishery Sciences | 1 | 20 | 559 | 598 |
| 833 | Devi Ahilya Vishwavidyalaya | 1 | 20 | 384 | 373 |
| 833 | Advanced Medical Research Center | 1 | 20 | 111 | 181 |
| 840 | Dartmouth College | 2 | 19 | 2073 | 3170 |
| 840 | Amherst College | 2 | 19 | 304 | 644 |
| 840 | Illinois State University | 1 | 19 | 2203 | 2093 |
| 840 | University of North Carolina at Charlotte | 1 | 19 | 2498 | 2412 |
| 840 | Vassar College | 2 | 19 | 353 | 768 |
| 840 | University of Shizuoka | 1 | 19 | 794 | 820 |
| 840 | Kyoto Sangyo University | 3 | 19 | 166 | 380 |
| 840 | Columbia University Irving Medical Center | 1 | 19 | 1395 | 1189 |
| 840 | Northeastern State University | 1 | 19 | 109 | 158 |
| 840 | Teikyo Heisei University | 1 | 19 | 187 | 151 |
| 840 | Anand Agricultural University | 1 | 19 | 173 | 178 |
| 840 | BioGeM | 1 | 19 | 135 | 197 |
| 840 | Saitama Medical University | 1 | 19 | 1435 | 1314 |
| 840 | Schneider Children's Medical Center of Israel | 1 | 19 | 1353 | 1599 |
| 840 | Graduate Center, CUNY | 2 | 19 | 1854 | 2885 |
| 855 | Sheffield Medical School | 1 | 18 | 327 | 437 |
| 855 | Colby College | 1 | 18 | 418 | 515 |
| 855 | University Medical Center Utrecht | 4 | 18 | 1686 | 2912 |
| 855 | UPMC Children's Hospital of Pittsburgh | 4 | 18 | 450 | 1172 |
| 855 | Osaka Women's and Children's Hospital | 2 | 18 | 102 | 242 |
| 855 | Harvey Mudd College | 1 | 18 | 298 | 371 |
| 855 | Japan Agency for Medical Research and Development | 1 | 18 | 120 | 204 |
| 862 | University of Warsaw | 1 | 17 | 4283 | 3449 |
| 862 | University of Iceland | 1 | 17 | 2219 | 2481 |
| 862 | University of Belgrade | 1 | 17 | 4765 | 4218 |
| 862 | University of Delhi | 3 | 17 | 2131 | 3450 |
| 862 | University of Innsbruck | 1 | 17 | 4125 | 4000 |
| 862 | Iwate Medical University | 5 | 17 | 380 | 1029 |
| 862 | Landspítali hospital | 1 | 17 | 533 | 685 |
| 862 | National Centre for Cell Science | 1 | 17 | 284 | 412 |
| 870 | University of Concepción | 2 | 16 | 311 | 807 |
| 870 | Free University | 1 | 16 | 3415 | 3667 |
| 870 | Chungbuk National University | 1 | 16 | 2275 | 2318 |
| 870 | Georgia Southern University | 1 | 16 | 1521 | 1551 |
| 870 | Southern University | 1 | 16 | 3966 | 3655 |
| 870 | Pontifical Catholic University of Chile | 4 | 16 | 1465 | 3440 |
| 870 | Janelia Research Campus | 5 | 16 | 70 | 302 |
| 870 | Pontificia Universidad Católica del Ecuador | 1 | 16 | 559 | 420 |
| 870 | Max Planck Institute for Molecular Genetics | 1 | 16 | 278 | 451 |
| 870 | Maharaja Sayajirao University of Baroda | 1 | 16 | 960 | 951 |
| 870 | LSU Health Sciences Center New Orleans | 2 | 16 | 1187 | 1922 |
| 870 | Brain and Mind Centre | 2 | 16 | 317 | 664 |
| 870 | Sydney Medical School | 3 | 16 | 1137 | 2317 |
| 870 | Wenzhou Medical University | 1 | 16 | 2322 | 2296 |
| 884 | Yonsei University | 4 | 15 | 3405 | 5619 |
| 884 | Sejong University | 1 | 15 | 1655 | 1841 |
| 884 | Universidad de Guanajuato | 1 | 15 | 1258 | 1151 |
| 884 | Industrial Research Limited | 1 | 15 | 3525 | 3614 |
| 884 | Linnaeus University | 4 | 15 | 1346 | 3544 |
| 884 | Hôtel-Dieu de Québec | 2 | 15 | 114 | 316 |
| 884 | National University of Tucumán | 1 | 15 | 772 | 763 |
| 884 | Department of Physiological Sciences | 1 | 15 | 1089 | 1363 |
| 884 | Instituto Superior de Investigaciones Biológicas | 1 | 15 | 125 | 174 |
| 884 | National Heart, Lung, and Blood Institute | 1 | 15 | 548 | 664 |
| 884 | Theodore Roosevelt High School | 1 | 15 | 3185 | 3465 |
| 895 | University of Tasmania | 1 | 14 | 3820 | 4217 |
| 895 | Missouri State University | 1 | 14 | 1557 | 1642 |
| 895 | National Defense University | 1 | 14 | 529 | 601 |
| 895 | Maine Medical Center | 2 | 14 | 467 | 908 |
| 895 | Temasek Life Sciences Laboratory | 18 | 14 | 14 | 277 |
| 895 | Tsuyama National College of Technology | 1 | 14 | 48 | 34 |
| 895 | Metropolitan Police Service | 1 | 14 | 64 | 87 |
| 895 | Royal Holloway University of London | 2 | 14 | 434 | 1084 |
| 903 | University of London | 5 | 13 | 4950 | 9459 |
| 903 | Brigham Young University | 4 | 13 | 1486 | 3922 |
| 903 | China Medical University | 2 | 13 | 2842 | 3647 |
| 903 | Ibaraki University | 1 | 13 | 1265 | 1217 |
| 903 | Centre for DNA Fingerprinting and Diagnostics | 2 | 13 | 84 | 230 |
| 903 | Herbert Irving Comprehensive Cancer Center | 1 | 13 | 313 | 478 |
| 903 | Centro Universitário de Rio Preto | 1 | 13 | 22 | 25 |
| 903 | Shanda | 1 | 13 | 284 | 397 |
| 903 | Brazilian Agricultural Research Corporation | 1 | 13 | 1880 | 1891 |
| 912 | Kagoshima University | 1 | 12 | 2495 | 2529 |
| 912 | University of Indonesia | 1 | 12 | 4428 | 2468 |
| 912 | Rice University | 1 | 12 | 3437 | 3964 |
| 912 | Jawaharlal Nehru University | 6 | 12 | 504 | 2106 |
| 916 | Cairo University | 1 | 11 | 5641 | 5430 |
| 916 | University of Southern Mississippi | 1 | 11 | 1709 | 2073 |
| 916 | University of York | 8 | 11 | 1175 | 5077 |
| 916 | Animal Health Trust | 1 | 11 | 146 | 224 |
| 916 | National Institute of Technology | 1 | 11 | 4299 | 4041 |
| 916 | Kanagawa Dental College | 1 | 11 | 204 | 284 |
| 916 | National Institute of Plant Genome Research | 1 | 11 | 157 | 228 |
| 923 | Meiji University | 1 | 10 | 1137 | 1200 |
| 923 | Linköping University | 1 | 10 | 4358 | 5147 |
| 923 | University of Porto | 3 | 10 | 2301 | 4899 |
| 923 | University of Rhode Island | 1 | 10 | 2681 | 3387 |
| 923 | National Institute on Deafness and Other Communication Disorders | 1 | 10 | 203 | 389 |
| 923 | Hollins University | 1 | 10 | 67 | 85 |
| 929 | James Madison University | 1 | 9 | 1192 | 1657 |
| 929 | City University of New York | 1 | 9 | 5580 | 5963 |
| 929 | Ritsumeikan University | 1 | 9 | 1750 | 1979 |
| 929 | East Carolina University | 1 | 9 | 3231 | 3888 |
| 929 | Queens College | 1 | 9 | 1433 | 1882 |
| 929 | Madison University | 1 | 9 | 1207 | 1681 |
| 929 | Symbiosis International University | 1 | 9 | 852 | 1020 |
| 929 | Taipei Veterans General Hospital | 1 | 9 | 2331 | 2553 |
| 929 | Graduate School USA | 1 | 9 | 2183 | 2914 |
| 929 | Carolina University | 1 | 9 | 3908 | 4506 |
| 939 | Nankai University | 3 | 8 | 1999 | 3716 |
| 939 | Uppsala University Hospital | 1 | 8 | 1135 | 1615 |
| 939 | Gyeongsang National University | 1 | 8 | 2320 | 2807 |
| 939 | Washington State University | 1 | 8 | 5329 | 6233 |
| 939 | Shantou University | 1 | 8 | 2001 | 2513 |
| 939 | Fujian Normal University | 1 | 8 | 1343 | 1681 |
| 939 | Central South University | 1 | 8 | 2835 | 3175 |
| 939 | South University | 2 | 8 | 4590 | 6248 |
| 939 | Amsterdam University Medical Centers | 1 | 8 | 2218 | 2464 |
| 948 | Trinity College Dublin | 9 | 7 | 757 | 5187 |
| 948 | University of Coimbra | 1 | 7 | 4659 | 5238 |
| 948 | Anhui Normal University | 1 | 7 | 989 | 1266 |
| 948 | Ministry of Education of the People's Republic of China | 1 | 7 | 2467 | 3117 |
| 948 | Guangxi University | 1 | 7 | 2243 | 2564 |
| 948 | Andrés Bello University | 2 | 7 | 359 | 830 |
| 948 | Federal University of ABC | 1 | 7 | 1534 | 1779 |
| 948 | Chicago Medical School | 1 | 7 | 742 | 1098 |
| 948 | Guangxi Medical University | 1 | 7 | 1467 | 1810 |
| 948 | Rosalind Franklin University of Medicine and Science | 1 | 7 | 665 | 1088 |
| 948 | Royal Hallamshire Hospital | 1 | 7 | 856 | 1196 |
| 948 | Second Military Medical University | 1 | 7 | 1995 | 2391 |
| 948 | Chinese National Human Genome Center, Beijing | 1 | 7 | 362 | 640 |
| 948 | École Normale Supérieure de Lyon | 4 | 7 | 305 | 1211 |
| 948 | Laboratoire de Reproduction et Développement des Plantes | 1 | 7 | 35 | 70 |
| 948 | Universidade do Porto | 1 | 7 | 3577 | 4242 |
| 964 | Tongji University | 3 | 6 | 3744 | 6090 |
| 964 | Australian National University | 2 | 6 | 4933 | 7724 |
| 964 | Reed College | 1 | 6 | 414 | 685 |
| 964 | Colgate University | 1 | 6 | 646 | 1001 |
| 964 | Villanova University | 1 | 6 | 1615 | 2318 |
| 964 | Toronto General Hospital | 1 | 6 | 1380 | 1912 |
| 964 | Indian Agricultural Research Institute | 1 | 6 | 819 | 1076 |
| 964 | Chinese Academy of Agricultural Sciences | 2 | 6 | 1340 | 2129 |
| 964 | Peter MacCallum Cancer Centre | 5 | 6 | 379 | 1246 |
| 964 | Southwest Medical University | 1 | 6 | 1143 | 1535 |
| 964 | McGowan Institute for Regenerative Medicine | 1 | 6 | 242 | 494 |
| 964 | Oregon National Primate Research Center | 1 | 6 | 384 | 640 |
| 964 | Institut des Sciences du Végétal | 1 | 6 | 99 | 188 |
| 964 | Madras Christian College | 1 | 6 | 3837 | 4307 |
| 978 | College of William & Mary | 1 | 5 | 1901 | 2686 |
| 978 | Fukuoka Prefectural University | 1 | 5 | 24 | 55 |
| 978 | University of the Basque Country | 1 | 5 | 4694 | 5590 |
| 978 | Manipal University | 4 | 5 | 343 | 1828 |
| 978 | Nagasaki International University | 1 | 5 | 122 | 257 |
| 978 | Oswaldo Cruz Foundation | 1 | 5 | 2321 | 2764 |
| 978 | Agency for Science, Technology and Research | 23 | 5 | 128 | 2402 |
| 985 | University of Texas Medical Branch | 1 | 4 | 3217 | 4117 |
| 985 | PSL University | 1 | 4 | 2044 | 2985 |
| 985 | University of Insubria | 1 | 4 | 1847 | 2828 |
| 985 | European Institute of Oncology | 1 | 4 | 868 | 1373 |
| 985 | University of Victoria | 1 | 4 | 4017 | 5457 |
| 985 | University of Dayton | 1 | 4 | 1226 | 2019 |
| 985 | Universidad Tecnológica | 1 | 4 | 737 | 965 |
| 985 | Tata Institute of Fundamental Research Hyderabad | 1 | 4 | 80 | 143 |
| 985 | Leeds Dental Institute | 1 | 4 | 117 | 231 |
| 985 | Oslo Metropolitan University | 1 | 4 | 4249 | 4895 |
| 985 | University of Wales, Newport | 1 | 4 | 244 | 554 |
| 985 | European Molecular Biology Organization | 1 | 4 | 10 | 31 |
| 997 | Goethe University Frankfurt | 1 | 3 | 4411 | 5783 |
| 997 | University of Lausanne | 2 | 3 | 2655 | 5206 |
| 997 | National Taiwan Normal University | 3 | 3 | 766 | 2537 |
| 997 | University of Missouri–Kansas City | 1 | 3 | 2334 | 3612 |