| 1 | British Trust for Ornithology | 134 | 6.4K | 2 | 2 |
| 2 | Royal Society for the Protection of Birds | 111 | 5.2K | 1 | 4 |
| 3 | University of Oxford | 88 | 4.8K | 490 | 528 |
| 4 | University of Cape Town | 122 | 3.3K | 46 | 112 |
| 5 | Centre for Ecology & Hydrology | 74 | 3.2K | 20 | 50 |
| 6 | University of Glasgow | 77 | 2.8K | 196 | 289 |
| 7 | University of Cambridge | 70 | 2.8K | 528 | 727 |
| 8 | Durham University | 68 | 2.6K | 102 | 143 |
| 9 | University of Groningen | 72 | 2.5K | 195 | 417 |
| 10 | University of Sheffield | 45 | 2.3K | 385 | 371 |
| 11 | University of Aberdeen | 90 | 2.3K | 47 | 190 |
| 12 | British Antarctic Survey | 55 | 2.1K | 30 | 46 |
| 13 | Natural History Museum | 119 | 2.1K | 41 | 113 |
| 14 | Tel Aviv University | 28 | 2.0K | 942 | 476 |
| 15 | ECOLOG | 90 | 1.9K | 77 | 120 |
| 16 | University of British Columbia | 23 | 1.8K | 1832 | 1052 |
| 17 | University of California | 64 | 1.7K | 2582 | 3231 |
| 18 | University of East Anglia | 59 | 1.7K | 72 | 195 |
| 19 | Max Planck Society | 27 | 1.6K | 906 | 770 |
| 20 | Lund University | 54 | 1.5K | 428 | 738 |
| 21 | Game & Wildlife Conservation Trust | 41 | 1.5K | 1 | 3 |
| 22 | Natural Environment Research Council | 35 | 1.5K | 13 | 37 |
| 23 | University of Stirling | 37 | 1.4K | 49 | 80 |
| 24 | University of Pennsylvania | 8 | 1.3K | 4145 | 1440 |
| 25 | Phillips Exeter Academy | 36 | 1.3K | 319 | 419 |
| 26 | Uppsala University | 35 | 1.3K | 645 | 735 |
| 27 | Newcastle University | 23 | 1.2K | 623 | 538 |
| 28 | Canadian Wildlife Service | 52 | 1.2K | 7 | 22 |
| 29 | Museo Nacional de Ciencias Naturales | 42 | 1.1K | 21 | 50 |
| 30 | Penryn Campus | 19 | 1.0K | 27 | 31 |
| 31 | United States Geological Survey | 55 | 1.0K | 315 | 440 |
| 32 | University of Copenhagen | 38 | 974 | 827 | 1216 |
| 33 | Cornell University | 46 | 911 | 725 | 1617 |
| 34 | The Nature Conservancy | 30 | 909 | 32 | 66 |
| 35 | University of Birmingham | 29 | 829 | 815 | 1079 |
| 36 | Wildfowl & Wetlands Trust | 44 | 810 | 1 | 4 |
| 37 | Department of Conservation | 38 | 805 | 39 | 84 |
| 38 | Muséum national d'histoire naturelle | 31 | 786 | 90 | 137 |
| 39 | University of Barcelona | 23 | 766 | 1026 | 1016 |
| 40 | Bio-Medical Science (South Korean) | 17 | 751 | 2601 | 1902 |
| 41 | Norwegian University of Science and Technology | 23 | 736 | 121 | 168 |
| 42 | University of Sussex | 23 | 734 | 314 | 506 |
| 43 | Aarhus University | 32 | 732 | 589 | 1014 |
| 44 | University of St Andrews | 31 | 728 | 122 | 226 |
| 45 | Bernardo O'Higgins University | 33 | 722 | 2709 | 1921 |
| 46 | University of Nottingham | 22 | 716 | 1014 | 1159 |
| 47 | University of Canterbury | 29 | 715 | 122 | 236 |
| 47 | NatureScot | 21 | 715 | 4 | 5 |
| 49 | University of Helsinki | 27 | 666 | 936 | 1314 |
| 50 | University of Bristol | 21 | 660 | 1015 | 1222 |
| 51 | University of Bern | 20 | 654 | 924 | 994 |
| 52 | Patuxent Wildlife Research Center | 26 | 642 | 41 | 75 |
| 53 | Polish Academy of Sciences | 21 | 635 | 952 | 769 |
| 54 | Universidad Metropolitana | 21 | 632 | 539 | 436 |
| 55 | National Scientific and Technical Research Council | 44 | 628 | 575 | 920 |
| 56 | University of Oulu | 24 | 626 | 296 | 496 |
| 57 | Simon Fraser University | 20 | 624 | 377 | 531 |
| 58 | King's College | 21 | 610 | 540 | 738 |
| 59 | University of Otago | 31 | 605 | 303 | 726 |
| 60 | University of Gothenburg | 26 | 592 | 696 | 1167 |
| 61 | United States Fish and Wildlife Service | 31 | 586 | 60 | 93 |
| 62 | University of Reading | 13 | 580 | 652 | 620 |
| 63 | Massey University | 24 | 574 | 203 | 378 |
| 64 | University of Wrocław | 18 | 572 | 384 | 226 |
| 65 | Charles University | 14 | 564 | 1169 | 775 |
| 65 | University of London | 30 | 564 | 972 | 1801 |
| 67 | Bangor University | 9 | 559 | 352 | 234 |
| 68 | University of Antwerp | 23 | 556 | 394 | 696 |
| 69 | Swedish University of Agricultural Sciences | 22 | 552 | 337 | 509 |
| 70 | Complutense University of Madrid | 27 | 548 | 719 | 943 |
| 71 | University of Utah | 5 | 546 | 3671 | 1532 |
| 72 | Colorado State University | 12 | 541 | 1048 | 892 |
| 72 | Biology Institute of Shandong Academy of Sciences | 26 | 541 | 435 | 629 |
| 74 | Manchester Metropolitan University | 16 | 540 | 180 | 207 |
| 75 | Queen's Medical Centre | 15 | 536 | 183 | 246 |
| 76 | University of Greifswald | 8 | 531 | 445 | 208 |
| 77 | Audubon | 11 | 527 | 18 | 21 |
| 78 | University of Michigan | 23 | 524 | 2537 | 3141 |
| 79 | Monash University | 13 | 507 | 2375 | 1970 |
| 80 | The Peregrine Fund | 26 | 505 | 2 | 6 |
| 80 | Australian Museum | 7 | 505 | 138 | 75 |
| 82 | Natural England | 17 | 501 | 3 | 16 |
| 83 | Wildlife Conservation Society | 20 | 498 | 53 | 96 |
| 84 | University of Edinburgh | 19 | 497 | 1380 | 1781 |
| 85 | University of Washington | 19 | 496 | 2527 | 2999 |
| 86 | University of Queensland | 18 | 487 | 2037 | 2372 |
| 87 | Czech Academy of Sciences | 16 | 485 | 875 | 787 |
| 88 | University of Leeds | 16 | 473 | 1335 | 1471 |
| 89 | Norwich Research Park | 16 | 471 | 118 | 249 |
| 90 | University of Newcastle | 6 | 469 | 1877 | 949 |
| 91 | Department of Forestry | 11 | 460 | 356 | 275 |
| 92 | University of Tokyo | 11 | 458 | 3715 | 2377 |
| 93 | Imperial College London | 16 | 456 | 2284 | 2493 |
| 94 | Stockholm University | 21 | 453 | 574 | 1032 |
| 94 | Tanzania Wildlife Research Institute | 4 | 453 | 20 | 10 |
| 96 | Memorial University of Newfoundland | 26 | 451 | 187 | 472 |
| 97 | Swedish Museum of Natural History | 24 | 450 | 46 | 78 |
| 98 | University of Amsterdam | 18 | 447 | 1434 | 1994 |
| 98 | University of Tasmania | 13 | 447 | 588 | 584 |
| 100 | Korea University of Science and Technology | 18 | 442 | 4982 | 4043 |
| 101 | Autonomous University of Madrid | 9 | 440 | 1024 | 659 |
| 101 | University of Guelph | 11 | 440 | 875 | 823 |
| 103 | Bielefeld University | 11 | 424 | 268 | 272 |
| 103 | Department of Internal Affairs | 7 | 424 | 127 | 3 |
| 105 | University of Wisconsin–Madison | 18 | 400 | 2546 | 3146 |
| 106 | University of Zurich | 13 | 394 | 1291 | 1431 |
| 107 | University of Iceland | 16 | 392 | 175 | 336 |
| 108 | Environment and Climate Change Canada | 18 | 388 | 105 | 188 |
| 109 | University of Granada | 16 | 384 | 767 | 854 |
| 110 | Forest Research | 8 | 382 | 207 | 163 |
| 111 | Chinese Academy of Sciences | 19 | 377 | 3125 | 3122 |
| 112 | National University of Singapore | 13 | 373 | 2156 | 2222 |
| 113 | Smithsonian Institution | 37 | 369 | 77 | 234 |
| 114 | Albertina | 13 | 368 | 10 | 13 |
| 115 | University of Kansas | 11 | 366 | 1108 | 1073 |
| 116 | Swansea University | 9 | 365 | 685 | 605 |
| 117 | University of Saskatchewan | 19 | 362 | 568 | 1004 |
| 118 | Madras Christian College | 13 | 359 | 1187 | 1082 |
| 119 | University of Florida | 21 | 356 | 1945 | 2941 |
| 120 | Finnish Museum of Natural History | 11 | 346 | 34 | 61 |
| 121 | University of Tartu | 12 | 338 | 365 | 502 |
| 122 | The Rockefeller University | 6 | 335 | 579 | 571 |
| 123 | University of Milan | 7 | 333 | 2765 | 1894 |
| 124 | Marie Curie | 10 | 328 | 1361 | 1406 |
| 125 | Industrial Research Limited | 7 | 326 | 1112 | 785 |
| 126 | Museum and Institute of Zoology | 10 | 324 | 14 | 19 |
| 127 | University of Auckland | 16 | 322 | 1002 | 1703 |
| 128 | University of Alicante | 7 | 321 | 629 | 373 |
| 129 | University of Buenos Aires | 17 | 314 | 711 | 877 |
| 130 | Boise State University | 13 | 312 | 78 | 146 |
| 131 | University of Lisbon | 14 | 307 | 1068 | 1304 |
| 132 | H.T. Harvey & Associates | 3 | 304 | 2 | 1 |
| 133 | University of Oslo | 16 | 303 | 1357 | 2009 |
| 134 | United States Forest Service | 17 | 301 | 338 | 520 |
| 135 | Harper Adams University | 3 | 286 | 101 | 43 |
| 136 | Rutgers University | 12 | 284 | 2323 | 2641 |
| 136 | University of Plymouth | 10 | 284 | 411 | 555 |
| 138 | Australian National University | 9 | 283 | 1766 | 1777 |
| 139 | Los Alamos National Laboratory | 2 | 281 | 1779 | 825 |
| 140 | University of Turku | 16 | 280 | 641 | 1161 |
| 141 | Max Planck Institute for Ornithology | 20 | 279 | 20 | 52 |
| 142 | University of Bath | 15 | 276 | 540 | 1094 |
| 143 | Charles Sturt University | 8 | 275 | 309 | 295 |
| 144 | Charles Darwin Foundation | 5 | 273 | 19 | 20 |
| 145 | University of Toronto | 8 | 272 | 5505 | 4475 |
| 145 | University of Maryland | 12 | 272 | 3015 | 3518 |
| 147 | Université de Montréal | 8 | 271 | 2304 | 1995 |
| 148 | University of Alberta | 17 | 268 | 1917 | 2973 |
| 149 | Institute for Biodiversity and Ecosystem Dynamics | 10 | 266 | 76 | 143 |
| 150 | Australian Antarctic Division | 7 | 263 | 62 | 84 |
| 150 | Zoological Society of London | 31 | 263 | 18 | 124 |
| 152 | University of Massachusetts | 9 | 262 | 2009 | 2260 |
| 153 | English Nature | 11 | 258 | 2 | 8 |
| 154 | American Museum of Natural History | 41 | 255 | 38 | 185 |
| 154 | Ohio State University | 7 | 255 | 5472 | 4247 |
| 154 | Museum of Vertebrate Zoology | 15 | 255 | 31 | 75 |
| 157 | Osaka City University | 14 | 254 | 463 | 594 |
| 157 | European Bird Census Council | 2 | 254 | 5 | 9 |
| 159 | University of Coimbra | 10 | 252 | 841 | 927 |
| 160 | University of Minnesota | 16 | 251 | 2699 | 3966 |
| 161 | Université du Québec à Rimouski | 7 | 250 | 110 | 98 |
| 162 | Indiana University | 5 | 246 | 4692 | 3035 |
| 163 | Princeton University | 12 | 244 | 1627 | 2281 |
| 163 | National Autonomous University of Mexico | 15 | 244 | 1188 | 1316 |
| 165 | Wageningen University & Research | 7 | 241 | 1340 | 1419 |
| 166 | University of Pavia | 5 | 240 | 1831 | 1258 |
| 166 | University of Manitoba | 12 | 240 | 1165 | 1678 |
| 168 | Halifax | 11 | 238 | 1339 | 1765 |
| 169 | University of Ibadan | 5 | 237 | 866 | 313 |
| 170 | Medway School of Science | 12 | 236 | 2934 | 2779 |
| 171 | McGill University | 11 | 235 | 2932 | 3469 |
| 171 | University of the Witwatersrand | 11 | 235 | 935 | 1036 |
| 171 | Dutch Centre for Field Ornithology | 8 | 235 | 4 | 8 |
| 174 | Polish Society for the Protection of Birds | 1 | 229 | 5 | 2 |
| 175 | Sustainability Institute | 7 | 228 | 100 | 150 |
| 176 | University of Malaya | 17 | 225 | 585 | 1100 |
| 177 | University College Cork | 12 | 224 | 440 | 863 |
| 178 | EcoHealth Alliance | 6 | 223 | 22 | 27 |
| 179 | PRBO Conservation Science | 9 | 222 | 3 | 23 |
| 180 | University of Nairobi | 10 | 221 | 182 | 234 |
| 180 | Falklands Conservation | 4 | 221 | 3 | 2 |
| 182 | University of Debrecen | 9 | 219 | 492 | 527 |
| 183 | Cornell Lab of Ornithology | 19 | 218 | 20 | 73 |
| 184 | Natura Cosméticos S.A. | 6 | 217 | 28 | 41 |
| 185 | Louisiana State University | 11 | 216 | 1458 | 1998 |
| 185 | Joint Nature Conservation Committee | 12 | 216 | 3 | 13 |
| 185 | Royal Holloway, University of London | 15 | 216 | 183 | 517 |
| 188 | University of Pretoria | 11 | 215 | 796 | 930 |
| 189 | Hungarian Academy of Sciences | 2 | 213 | 2380 | 1085 |
| 190 | Dorset Green Technology Park | 7 | 209 | 12 | 30 |
| 191 | Queen's University Belfast | 5 | 208 | 1828 | 1406 |
| 191 | Society for the Protection of Nature in Israel | 4 | 208 | 1 | 1 |
| 193 | Rikkyo University | 10 | 207 | 109 | 99 |
| 194 | Norwegian Institute of Marine Research | 5 | 206 | 208 | 199 |
| 194 | Lone Star College System | 8 | 206 | 2740 | 2987 |
| 196 | Adam Mickiewicz University in Poznań | 8 | 203 | 1015 | 813 |
| 196 | Smithsonian Tropical Research Institute | 6 | 203 | 194 | 231 |
| 198 | University of Pisa | 7 | 199 | 1895 | 1810 |
| 199 | Ahmadu Bello University | 6 | 193 | 286 | 132 |
| 200 | University of Bergen | 11 | 190 | 1044 | 1756 |
| 201 | University of Connecticut | 4 | 189 | 3414 | 2367 |
| 201 | Spanish National Research Council | 11 | 189 | 673 | 1140 |
| 203 | University of Aveiro | 11 | 188 | 631 | 909 |
| 203 | University of Turin | 11 | 188 | 1175 | 1805 |
| 203 | Royal Ontario Museum | 8 | 188 | 47 | 65 |
| 206 | University of Montpellier | 9 | 186 | 1143 | 1484 |
| 207 | National Zoological Park | 6 | 184 | 97 | 116 |
| 208 | Johannes Gutenberg University Mainz | 2 | 179 | 2427 | 1220 |
| 209 | University of the West of Scotland | 4 | 178 | 279 | 212 |
| 209 | Archaeological Institute of Kashihara | 17 | 178 | 9 | 38 |
| 211 | Correos | 5 | 177 | 151 | 209 |
| 212 | Cardiff University | 9 | 176 | 1511 | 2130 |
| 212 | University of California, Davis | 10 | 176 | 1232 | 2105 |
| 214 | Edinburgh Napier University | 3 | 174 | 519 | 288 |
| 215 | Yamashina Institute for Ornithology | 6 | 173 | 6 | 6 |
| 215 | Broadway | 3 | 173 | 1723 | 1273 |
| 217 | Mississippi State University | 8 | 172 | 725 | 908 |
| 217 | Fera Science | 4 | 172 | 55 | 66 |
| 217 | Royal Veterinary College | 4 | 172 | 356 | 324 |
| 220 | Field Museum of Natural History | 14 | 171 | 68 | 158 |
| 221 | Marine and Freshwater Research Institute | 5 | 170 | 136 | 129 |
| 221 | Centre for Life | 3 | 170 | 776 | 557 |
| 223 | University of Missouri | 8 | 169 | 3004 | 3285 |
| 224 | British Ornithologists' Union | 15 | 168 | 1 | 1 |
| 225 | Manaaki Whenua – Landcare Research | 7 | 167 | 148 | 238 |
| 226 | Victoria University | 12 | 166 | 603 | 1286 |
| 226 | University of Vigo | 5 | 166 | 663 | 584 |
| 226 | Victoria University of Wellington | 11 | 166 | 347 | 781 |
| 226 | University of New Mexico | 7 | 166 | 1668 | 2007 |
| 230 | Université de La Rochelle | 8 | 164 | 58 | 134 |
| 231 | Pennsylvania State University | 4 | 162 | 5772 | 4209 |
| 231 | Carleton University | 10 | 162 | 636 | 1151 |
| 233 | University of Valencia | 10 | 161 | 869 | 1468 |
| 234 | University of Giessen | 2 | 160 | 2773 | 1461 |
| 234 | Forest Research Institute | 7 | 160 | 341 | 404 |
| 236 | Konrad Lorenz Institute for Evolution and Cognition Research | 10 | 158 | 5 | 46 |
| 236 | University College London | 14 | 158 | 2495 | 4172 |
| 236 | Deakin University | 16 | 158 | 637 | 1811 |
| 236 | University of Leicester | 4 | 158 | 2276 | 1753 |
| 236 | Museo delle Scienze | 3 | 158 | 33 | 27 |
| 241 | University of Melbourne | 5 | 157 | 5170 | 4352 |
| 241 | Fudan University | 3 | 157 | 4337 | 2617 |
| 243 | Kristianstad University | 4 | 156 | 88 | 67 |
| 244 | IPB University | 5 | 155 | 870 | 525 |
| 245 | University of Liverpool | 10 | 154 | 1933 | 2804 |
| 245 | Leibniz Association | 5 | 154 | 1132 | 1142 |
| 247 | Yale University | 11 | 153 | 2552 | 3843 |
| 247 | University of Jyväskylä | 7 | 153 | 670 | 869 |
| 247 | Czech University of Life Sciences Prague | 4 | 153 | 304 | 210 |
| 250 | Thomson Reuters | 8 | 152 | 19 | 50 |
| 250 | University of Costa Rica | 4 | 152 | 486 | 310 |
| 252 | University of Burgundy | 9 | 151 | 511 | 918 |
| 253 | James Cook University | 4 | 150 | 1192 | 1036 |
| 253 | National Institute of Polar Research | 5 | 150 | 108 | 105 |
| 253 | University of Ghana | 11 | 150 | 226 | 436 |
| 256 | Prince George's Community College | 1 | 149 | 352 | 25 |
| 257 | University of Göttingen | 4 | 148 | 2470 | 2063 |
| 257 | Ball State University | 1 | 148 | 1483 | 283 |
| 257 | Birds Canada | 5 | 148 | 8 | 15 |
| 257 | Madison University | 1 | 148 | 1207 | 251 |
| 261 | Evolution Group | 4 | 147 | 44 | 89 |
| 262 | Livingston College | 5 | 146 | 7 | 14 |
| 263 | National Park Service | 4 | 145 | 228 | 174 |
| 263 | University of Basel | 4 | 145 | 2311 | 1979 |
| 263 | University of Montana | 7 | 145 | 332 | 557 |
| 263 | National Parks and Wildlife Service | 7 | 145 | 12 | 30 |
| 267 | University of Extremadura | 11 | 144 | 310 | 628 |
| 267 | University of Nevada, Reno | 10 | 144 | 813 | 1637 |
| 269 | Ghana Wildlife Society | 3 | 143 | 3 | 1 |
| 270 | Bedford Institute of Oceanography | 4 | 142 | 156 | 171 |
| 270 | Universitat Politècnica de Catalunya | 1 | 142 | 2571 | 913 |
| 272 | University of Southampton | 8 | 141 | 2052 | 2864 |
| 273 | University of Vienna | 8 | 140 | 3190 | 3664 |
| 273 | University of Tromsø – The Arctic University of Norway | 6 | 140 | 874 | 1114 |
| 273 | Scotland's Rural College | 2 | 140 | 315 | 218 |
| 276 | Wuhan University | 7 | 138 | 2409 | 2413 |
| 277 | Agropolis International | 1 | 137 | 362 | 184 |
| 278 | Barnard College | 7 | 136 | 94 | 262 |
| 279 | Hebrew University of Jerusalem | 9 | 135 | 1586 | 2385 |
| 279 | Hellenic Ornithological Society | 2 | 135 | 6 | 3 |
| 281 | Hawk Mountain Sanctuary | 6 | 134 | 4 | 10 |
| 282 | Norwegian Polar Institute | 3 | 133 | 130 | 128 |
| 282 | Murdoch University | 5 | 133 | 796 | 954 |
| 284 | Moscow State University | 12 | 132 | 1456 | 1711 |
| 284 | Russian Academy of Sciences | 8 | 132 | 3437 | 2826 |
| 286 | University of Dundee | 4 | 131 | 1496 | 1508 |
| 287 | Swiss Ornithological Institute Sempach | 8 | 130 | 6 | 11 |
| 287 | National Museums of Kenya | 12 | 130 | 24 | 91 |
| 289 | University of Oviedo | 9 | 129 | 655 | 1123 |
| 290 | Oregon State University | 8 | 128 | 1430 | 2200 |
| 290 | The Institute of Conservation | 8 | 128 | 28 | 65 |
| 292 | University of Rennes 1 | 8 | 127 | 640 | 1054 |
| 292 | Hokkaido University | 10 | 127 | 2093 | 2561 |
| 292 | State University of New York at Albany | 3 | 127 | 536 | 465 |
| 295 | University of Ioannina | 1 | 125 | 2514 | 980 |
| 296 | St Bartholomew's Hospital | 4 | 124 | 566 | 590 |
| 296 | Smithsonian Conservation Biology Institute | 8 | 124 | 69 | 126 |
| 296 | Universidad de Córdoba | 2 | 124 | 1134 | 462 |
| 299 | Eötvös Loránd University | 10 | 123 | 412 | 754 |
| 300 | West Virginia University | 6 | 122 | 1573 | 1873 |
| 300 | Durrell Wildlife Conservation Trust | 8 | 122 | 11 | 25 |
| 302 | University of Freiburg | 6 | 121 | 1848 | 2238 |
| 302 | Ghent University | 8 | 121 | 2680 | 3649 |
| 302 | Macaulay Institute | 4 | 121 | 131 | 183 |
| 305 | National Taiwan Normal University | 4 | 120 | 610 | 505 |
| 305 | Kyushu University | 4 | 120 | 3514 | 2667 |
| 307 | Queen Mary University of London | 6 | 119 | 1590 | 2012 |
| 308 | UNESCO | 2 | 118 | 736 | 398 |
| 308 | Johns Hopkins University | 6 | 118 | 4055 | 4364 |
| 310 | University of Cologne | 2 | 117 | 3784 | 2249 |
| 310 | University of Haifa | 9 | 117 | 663 | 1288 |
| 310 | University of KwaZulu-Natal | 6 | 117 | 1100 | 1220 |
| 310 | Tshwane University of Technology | 5 | 117 | 255 | 239 |
| 310 | Medical Research Council | 2 | 117 | 2069 | 1626 |
| 315 | Universidad Nacional del Comahue | 6 | 114 | 131 | 192 |
| 316 | Brock University | 4 | 113 | 681 | 683 |
| 316 | University of Gloucestershire | 5 | 113 | 93 | 148 |
| 318 | Sun Yat-sen University | 8 | 112 | 2978 | 3425 |
| 318 | University of Wales | 7 | 112 | 874 | 1559 |
| 318 | University of the Highlands and Islands | 3 | 112 | 72 | 81 |
| 318 | University of California, Los Angeles | 5 | 112 | 2120 | 2660 |
| 322 | Mahidol University | 4 | 111 | 1852 | 1603 |
| 322 | Environmental Research Institute | 4 | 111 | 112 | 175 |
| 322 | Environment Agency | 4 | 111 | 224 | 274 |
| 325 | Jagiellonian University | 6 | 110 | 1708 | 1718 |
| 325 | University of Gdańsk | 8 | 110 | 621 | 822 |
| 325 | Scottish Universities Environmental Research Centre | 5 | 110 | 65 | 108 |
| 328 | United States Department of Agriculture | 3 | 109 | 1252 | 1209 |
| 328 | Museu Paraense Emílio Goeldi | 5 | 109 | 88 | 95 |
| 330 | University of South Florida | 6 | 108 | 2402 | 2952 |
| 330 | Beijing Normal University | 8 | 108 | 1146 | 1543 |
| 330 | North Highland College | 2 | 108 | 20 | 25 |
| 333 | University of Science and Technology of China | 4 | 107 | 2298 | 1921 |
| 333 | Pierre and Marie Curie University | 3 | 107 | 1749 | 1676 |
| 333 | Palacký University Olomouc | 7 | 107 | 290 | 509 |
| 333 | National Trust for Scotland | 4 | 107 | 1 | 2 |
| 337 | University of Pittsburgh | 3 | 106 | 5680 | 4532 |
| 337 | La Trobe University | 5 | 106 | 1731 | 2078 |
| 337 | Canterbury Museum | 4 | 106 | 12 | 18 |
| 340 | California Academy of Sciences | 5 | 105 | 90 | 127 |
| 341 | United Nations | 4 | 104 | 613 | 658 |
| 341 | University of La Laguna | 4 | 104 | 923 | 776 |
| 341 | Allegheny College | 2 | 104 | 100 | 60 |
| 341 | Fram Centre | 2 | 104 | 129 | 106 |
| 341 | Natural History Museum of Los Angeles County | 4 | 104 | 61 | 65 |
| 341 | Food and Agriculture Organization of the United Nations | 4 | 104 | 182 | 248 |
| 347 | Academy of Natural Sciences of Drexel University | 6 | 103 | 52 | 110 |
| 348 | City University of New York | 5 | 102 | 2058 | 2324 |
| 348 | Liverpool John Moores University | 5 | 102 | 719 | 1002 |
| 350 | Harvard University | 7 | 101 | 6182 | 7113 |
| 350 | Virginia Tech | 5 | 101 | 2906 | 3258 |
| 350 | Microsoft Research | 1 | 101 | 705 | 449 |
| 350 | National Institute for Education | 3 | 101 | 134 | 21 |
| 354 | University of Alaska Fairbanks | 8 | 100 | 253 | 501 |
| 354 | Foreign and Commonwealth Office | 2 | 100 | 17 | 2 |
| 356 | University of Adelaide | 5 | 99 | 2846 | 3319 |
| 356 | Iowa State University | 4 | 99 | 2898 | 2834 |
| 356 | Griffith University | 5 | 99 | 2081 | 2585 |
| 356 | Bombay Natural History Society | 8 | 99 | 5 | 14 |
| 360 | University of Caen Normandy | 1 | 98 | 1742 | 753 |
| 360 | Institut für Meereskunde Kiel | 1 | 98 | 4 | 5 |
| 362 | Autonomous University of Tlaxcala | 2 | 97 | 119 | 42 |
| 362 | New Zealand Broadcasting Corporation | 1 | 97 | 1 | 1 |
| 364 | Stanford University | 2 | 96 | 8467 | 6107 |
| 364 | University of Chicago | 5 | 96 | 4228 | 4386 |
| 364 | Stellenbosch University | 7 | 96 | 1211 | 1825 |
| 364 | Heriot-Watt University | 4 | 96 | 1068 | 1129 |
| 368 | Qinetiq (United Kingdom) | 1 | 95 | 179 | 86 |
| 369 | Acadia University | 5 | 94 | 124 | 248 |
| 369 | University of Arizona | 5 | 94 | 4079 | 4464 |
| 371 | Federal University of Minas Gerais | 6 | 93 | 2273 | 2097 |
| 371 | University of Lausanne | 7 | 93 | 1101 | 1966 |
| 371 | University of Maine | 4 | 93 | 838 | 1045 |
| 371 | Westfield College | 1 | 93 | 819 | 409 |
| 375 | Charles Darwin University | 3 | 92 | 534 | 521 |
| 375 | Laval University | 5 | 92 | 2550 | 3056 |
| 375 | University of the Basque Country | 5 | 92 | 1800 | 1981 |
| 375 | Trent University | 4 | 92 | 1175 | 1415 |
| 375 | Colorado Parks and Wildlife | 2 | 92 | 68 | 55 |
| 380 | University of Miami | 5 | 91 | 2731 | 3254 |
| 380 | Umeå University | 6 | 91 | 1516 | 2319 |
| 382 | Leiden University | 8 | 90 | 2155 | 3448 |
| 382 | Wetlands International | 4 | 90 | 6 | 36 |
| 384 | University of Strasbourg | 3 | 89 | 1922 | 1637 |
| 384 | University of Hull | 1 | 89 | 2965 | 1351 |
| 384 | Scottish Association for Marine Science | 3 | 89 | 108 | 156 |
| 387 | Purdue University | 5 | 88 | 4280 | 4671 |
| 387 | Ferdowsi University of Mashhad | 2 | 88 | 1794 | 976 |
| 387 | Leuphana University of Lüneburg | 1 | 88 | 813 | 376 |
| 387 | Florida State University | 1 | 88 | 5302 | 2665 |
| 387 | University of Putra Malaysia | 1 | 88 | 3692 | 1477 |
| 392 | University of Murcia | 3 | 87 | 1830 | 1444 |
| 392 | University of Natal | 5 | 87 | 322 | 452 |
| 394 | University of Jos | 4 | 86 | 110 | 77 |
| 395 | Mauritian Wildlife Foundation | 6 | 85 | 2 | 14 |
| 396 | Rhodes University | 10 | 84 | 244 | 594 |
| 396 | University of North Dakota | 3 | 84 | 886 | 908 |
| 396 | University of Prince Edward Island | 2 | 84 | 405 | 294 |
| 399 | Jean Monnet University | 3 | 83 | 265 | 278 |
| 399 | Academia Sinica | 4 | 83 | 2038 | 2033 |
| 401 | Humboldt State University | 3 | 82 | 172 | 225 |
| 401 | University of Cape Verde | 3 | 82 | 3112 | 2405 |
| 401 | Linnaeus University | 7 | 82 | 829 | 1629 |
| 401 | Animal and Plant Health Agency | 2 | 82 | 128 | 129 |
| 405 | Auburn University | 6 | 81 | 1476 | 2153 |
| 405 | College of William & Mary | 1 | 81 | 1901 | 906 |
| 405 | Carnegie Museum of Natural History | 3 | 81 | 49 | 64 |
| 408 | Pacific Science Center | 1 | 80 | 25 | 21 |
| 408 | Havstovan | 2 | 80 | 15 | 19 |
| 410 | Kyoto University | 5 | 79 | 4258 | 4166 |
| 410 | Washington State University | 6 | 79 | 1933 | 3007 |
| 410 | Naturalis Biodiversity Center | 8 | 79 | 107 | 233 |
| 410 | University of Eswatini | 3 | 79 | 55 | 57 |
| 414 | Utrecht University | 7 | 78 | 2658 | 4172 |
| 414 | Michigan State University | 8 | 78 | 2995 | 4873 |
| 414 | University of New England | 7 | 78 | 569 | 1341 |
| 414 | University of the South Pacific | 2 | 78 | 307 | 224 |
| 414 | University of Liège | 1 | 78 | 3982 | 2284 |
| 414 | Joetsu University of Education | 6 | 78 | 34 | 46 |
| 420 | University of the Faroe Islands | 1 | 77 | 104 | 33 |
| 420 | Keldur | 1 | 77 | 64 | 38 |
| 422 | Bournemouth University | 5 | 76 | 376 | 734 |
| 422 | Hostelling International USA | 2 | 76 | 104 | 105 |
| 424 | King's Buildings | 5 | 75 | 427 | 877 |
| 425 | Nagoya University | 4 | 74 | 3331 | 3093 |
| 425 | University of Lapland | 2 | 74 | 187 | 128 |
| 427 | Claude Bernard University Lyon 1 | 7 | 73 | 1706 | 2641 |
| 427 | Agroecology | 3 | 73 | 662 | 664 |
| 429 | University of Marburg | 2 | 72 | 1856 | 1541 |
| 429 | University of Calgary | 2 | 72 | 6064 | 4521 |
| 429 | University of South Bohemia in České Budějovice | 5 | 72 | 355 | 570 |
| 429 | University of Cyprus | 3 | 72 | 846 | 914 |
| 429 | Marine Institute of Memorial University of Newfoundland | 1 | 72 | 428 | 255 |
| 429 | National University of Mar del Plata | 5 | 72 | 296 | 406 |
| 435 | Generalitat of Catalonia | 1 | 71 | 329 | 181 |
| 435 | National Biodiversity Institute | 2 | 71 | 130 | 114 |
| 435 | California Department of Parks and Recreation | 1 | 71 | 9 | 2 |
| 435 | Allan Wilson Centre | 3 | 71 | 44 | 89 |
| 439 | Radboud University Nijmegen | 6 | 70 | 2314 | 3581 |
| 439 | Royal Society | 7 | 70 | 37 | 130 |
| 439 | College of Nyíregyháza | 3 | 70 | 14 | 30 |
| 442 | Goethe University Frankfurt | 3 | 69 | 2575 | 2505 |
| 442 | University of Castile-La Mancha | 3 | 69 | 742 | 684 |
| 442 | University of York | 6 | 69 | 1494 | 2641 |
| 442 | University of Kentucky | 1 | 69 | 7704 | 4415 |
| 442 | University of Iowa | 3 | 69 | 4427 | 4238 |
| 447 | University of Potsdam | 4 | 68 | 838 | 1298 |
| 447 | University of Sydney | 4 | 68 | 6267 | 6464 |
| 447 | University of Missouri–St. Louis | 1 | 68 | 137 | 78 |
| 447 | Pontifícia Universidade Católica de Minas Gerais | 2 | 68 | 477 | 169 |
| 447 | University of North Carolina | 5 | 68 | 4782 | 5755 |
| 447 | University of Kent | 5 | 68 | 1019 | 1689 |
| 447 | Thomas Jefferson University | 3 | 68 | 2760 | 2759 |
| 447 | University of Salford | 2 | 68 | 1496 | 1164 |
| 447 | Saint Louis Zoo | 1 | 68 | 47 | 28 |
| 447 | Lancaster Environment Centre | 9 | 68 | 144 | 470 |
| 447 | state health agency | 1 | 68 | 1170 | 821 |
| 447 | Wadsworth Center | 1 | 68 | 700 | 587 |
| 459 | University of Hawaii | 3 | 67 | 1759 | 2179 |
| 459 | University of Western Brittany | 3 | 67 | 608 | 692 |
| 459 | Lancaster University | 8 | 67 | 807 | 2196 |
| 462 | Nippon Telegraph and Telephone | 2 | 66 | 396 | 280 |
| 462 | National Institute of Water and Atmospheric Research | 1 | 66 | 455 | 352 |
| 462 | University of Paris-Sud | 4 | 66 | 1957 | 2569 |
| 462 | Macdonald Campus | 2 | 66 | 383 | 402 |
| 462 | Institute for Nature Management | 1 | 66 | 160 | 70 |
| 467 | Duke University | 3 | 65 | 4895 | 4818 |
| 467 | Hofstra University | 1 | 65 | 1868 | 803 |
| 467 | Pacific University | 2 | 65 | 599 | 497 |
| 467 | Ontario Veterinary College | 1 | 65 | 433 | 316 |
| 471 | Kansas State University | 5 | 64 | 1459 | 2184 |
| 471 | University of Oklahoma | 8 | 64 | 1668 | 3531 |
| 471 | University of Georgia | 4 | 64 | 3549 | 4071 |
| 471 | University of North Texas | 3 | 64 | 1942 | 2003 |
| 471 | Indian Institute of Science | 3 | 64 | 2019 | 1881 |
| 471 | University of Victoria | 4 | 64 | 1698 | 2344 |
| 471 | Estonian Institute of Zoology and Botany | 2 | 64 | 18 | 37 |
| 471 | Intermountain Healthcare | 2 | 64 | 442 | 406 |
| 479 | University of Waikato | 3 | 63 | 1028 | 1203 |
| 479 | University of Colorado | 5 | 63 | 3612 | 4898 |
| 479 | Nelson Mandela University | 4 | 63 | 200 | 199 |
| 479 | Sultan Qaboos University | 4 | 63 | 694 | 1058 |
| 483 | University of New South Wales | 8 | 62 | 3338 | 5866 |
| 483 | Tulane University | 3 | 62 | 2497 | 2717 |
| 483 | University of Windsor | 3 | 62 | 1220 | 1290 |
| 483 | Punjab Agricultural University | 3 | 62 | 440 | 376 |
| 483 | Research Center for Eco-Environmental Sciences | 2 | 62 | 592 | 613 |
| 488 | Addenbrooke's Hospital | 1 | 61 | 1934 | 1422 |
| 488 | Ohio University | 3 | 61 | 1534 | 1744 |
| 488 | Western University | 3 | 61 | 2048 | 2034 |
| 488 | University of the Azores | 1 | 61 | 326 | 177 |
| 488 | IFREMER | 3 | 61 | 573 | 724 |
| 488 | Konrad Lorenz Forschungsstelle | 1 | 61 | 20 | 18 |
| 488 | Ludwig Boltzmann Gesellschaft | 1 | 61 | 868 | 670 |
| 488 | Foundation for Advancement of International Medical Education and Research | 1 | 61 | 40 | 29 |
| 488 | Sea Mammal Research Unit | 1 | 61 | 112 | 116 |
| 497 | University of Hong Kong | 4 | 60 | 6248 | 6505 |
| 497 | University of Oldenburg | 3 | 60 | 872 | 1060 |
| 497 | Hunter College | 2 | 60 | 1098 | 1004 |
| 500 | Makerere University | 6 | 59 | 482 | 948 |
| 500 | Anglia Ruskin University | 3 | 59 | 493 | 699 |
| 502 | Boston University | 3 | 58 | 4515 | 4735 |
| 502 | National Trust | 2 | 58 | 24 | 24 |
| 502 | University of Nicosia | 1 | 58 | 840 | 457 |
| 502 | Lafayette College | 1 | 58 | 567 | 268 |
| 502 | Université de Moncton | 3 | 58 | 259 | 361 |
| 502 | interdisciplinarity | 4 | 58 | 5977 | 6110 |
| 502 | Canadian Museum of Nature | 2 | 58 | 134 | 140 |
| 509 | Norwegian University of Life Sciences | 3 | 57 | 820 | 1080 |
| 509 | Aves Argentinas | 3 | 57 | 4 | 4 |
| 509 | California State University, Long Beach | 1 | 57 | 561 | 325 |
| 509 | British Museum | 32 | 57 | 9 | 75 |
| 509 | Queen's University Kingston | 6 | 57 | 129 | 373 |
| 514 | University of Málaga | 2 | 56 | 1875 | 1349 |
| 514 | University of Western Australia | 6 | 56 | 2649 | 4401 |
| 514 | Ulster University | 1 | 56 | 2957 | 1846 |
| 514 | Federal University of Rio Grande | 2 | 56 | 851 | 409 |
| 514 | Tyndall Centre | 2 | 56 | 133 | 214 |
| 519 | Vanderbilt University | 1 | 55 | 6887 | 4674 |
| 519 | Estonian University of Life Sciences | 3 | 55 | 258 | 361 |
| 519 | North Carolina State University | 4 | 55 | 3265 | 4147 |
| 519 | Royal Holloway University of London | 5 | 55 | 168 | 574 |
| 523 | Meijo University | 2 | 54 | 624 | 445 |
| 523 | University College Dublin | 7 | 54 | 1385 | 3269 |
| 523 | University of Białystok | 4 | 54 | 633 | 783 |
| 523 | Arthur Rylah Institute for Environmental Research | 3 | 54 | 60 | 90 |
| 523 | Catalan Ornithological Institute | 2 | 54 | 8 | 17 |
| 523 | Royal Navy | 1 | 54 | 44 | 11 |
| 529 | California State University | 3 | 53 | 3171 | 3179 |
| 529 | Universidade Estadual Paulista | 3 | 53 | 3238 | 2433 |
| 529 | Heritage College of Osteopathic Medicine | 2 | 53 | 182 | 184 |
| 529 | Icelandic Institute of Natural History | 3 | 53 | 16 | 46 |
| 529 | Marine Biological Association of the United Kingdom | 2 | 53 | 160 | 232 |
| 534 | Arizona State University | 1 | 52 | 7411 | 4854 |
| 534 | PSL University | 5 | 52 | 774 | 1296 |
| 534 | Universidade de Santiago de Compostela | 2 | 52 | 1033 | 850 |
| 537 | Brigham Young University | 4 | 51 | 1486 | 2400 |
| 537 | Université du Québec à Montréal | 2 | 51 | 1710 | 1613 |
| 539 | Columbia University | 7 | 50 | 4368 | 6929 |
| 539 | University of Ferrara | 2 | 50 | 1922 | 1892 |
| 539 | East Carolina University | 2 | 50 | 2139 | 1979 |
| 539 | United States Air Force Academy | 4 | 50 | 1376 | 1844 |
| 539 | Guyra Paraguay | 1 | 50 | 11 | 11 |
| 539 | Plymouth Marine Laboratory | 1 | 50 | 306 | 320 |
| 539 | Universidad de Navarra | 5 | 50 | 944 | 1825 |
| 539 | Carolina University | 2 | 50 | 2651 | 2302 |
| 547 | IE University | 2 | 49 | 47 | 82 |
| 547 | Wildlife Institute of India | 3 | 49 | 84 | 104 |
| 547 | Keio University Shonan Fujisawa Campus | 1 | 49 | 14 | 8 |
| 547 | Wuhan Institute of Virology | 1 | 49 | 288 | 257 |
| 547 | Conservation International | 1 | 49 | 213 | 205 |
| 547 | National University of Austral Patagonia | 2 | 49 | 54 | 43 |
| 547 | City College | 2 | 49 | 1029 | 970 |
| 554 | Flinders University | 7 | 48 | 1115 | 2820 |
| 554 | Humboldt University of Berlin | 2 | 48 | 7197 | 5955 |
| 554 | Bowdoin College | 3 | 48 | 167 | 330 |
| 554 | Slovak Academy of Sciences | 3 | 48 | 1505 | 1710 |
| 554 | Southwest Forestry University | 1 | 48 | 484 | 278 |
| 554 | Evolution Institute | 4 | 48 | 28 | 51 |
| 554 | Edwards | 2 | 48 | 481 | 692 |
| 554 | Museu de Ciências Naturais da Fundação Zoobotânica do Rio Grande do Sul | 4 | 48 | 62 | 143 |
| 562 | Heidelberg University | 4 | 47 | 2708 | 3766 |
| 562 | University of Valladolid | 3 | 47 | 1383 | 1477 |
| 562 | Vytautas Magnus University Agriculture Academy | 1 | 47 | 120 | 73 |
| 562 | Sorbonne Universités | 3 | 47 | 1114 | 1657 |
| 566 | Sapienza University of Rome | 2 | 46 | 6467 | 5575 |
| 566 | Trinity College Dublin | 3 | 46 | 2110 | 2818 |
| 566 | Florida Atlantic University | 2 | 46 | 1858 | 1755 |
| 566 | Museums Victoria | 2 | 46 | 103 | 116 |
| 566 | University of Sunderland | 1 | 46 | 776 | 469 |
| 566 | Atlantic University | 2 | 46 | 1900 | 1786 |
| 566 | University of California, Santa Barbara | 2 | 46 | 1267 | 1522 |
| 566 | Northeast Catholic College | 2 | 46 | 79 | 86 |
| 574 | University of Chile | 5 | 45 | 1977 | 3127 |
| 574 | University of Ottawa | 3 | 45 | 4424 | 4941 |
| 574 | University of Puerto Rico at Mayaguez | 3 | 45 | 744 | 722 |
| 574 | University of Tennessee | 2 | 45 | 5218 | 4830 |
| 574 | list of agriculture ministries | 4 | 45 | 249 | 550 |
| 574 | Galapagos (Netherlands) | 1 | 45 | 137 | 124 |
| 574 | Museum of New Zealand Te Papa Tongarewa | 3 | 45 | 39 | 93 |
| 581 | University of North Carolina at Chapel Hill | 5 | 44 | 4500 | 6251 |
| 581 | University of León | 4 | 44 | 455 | 821 |
| 581 | University of Eastern Piedmont | 1 | 44 | 1310 | 994 |
| 581 | Rothamsted Research | 2 | 44 | 564 | 694 |
| 585 | Bay de Noc Community College | 1 | 43 | 3 | 3 |
| 585 | Rural Development Service | 1 | 43 | 5 | 5 |
| 585 | Australian Museums and Galleries Association | 4 | 43 | 97 | 319 |
| 585 | Natural History Museum of Geneva | 4 | 43 | 19 | 40 |
| 585 | University of Agriculture | 2 | 43 | 3082 | 2378 |
| 590 | Wellesley College | 1 | 42 | 701 | 592 |
| 590 | University of Antananarivo | 1 | 42 | 325 | 214 |
| 590 | onet.pl | 3 | 42 | 20 | 32 |
| 590 | Society for Science and the Public | 2 | 42 | 120 | 170 |
| 590 | Burke Museum of Natural History and Culture | 4 | 42 | 55 | 144 |
| 590 | South Florida Water Management District | 1 | 42 | 101 | 107 |
| 590 | Université de Lyon | 2 | 42 | 2200 | 2351 |
| 597 | Junta de Extremadura | 1 | 41 | 75 | 63 |
| 597 | Hasselt University | 1 | 41 | 1715 | 1350 |
| 597 | Nottingham Trent University | 2 | 41 | 1456 | 1658 |
| 597 | Glendon College | 1 | 41 | 109 | 68 |
| 597 | Denver Museum of Nature and Science | 1 | 41 | 308 | 178 |
| 597 | Ministry of Natural Resources and Forestry | 1 | 41 | 120 | 93 |
| 597 | National University of La Pampa | 1 | 41 | 264 | 123 |
| 597 | Paris-Saclay Institute of Neuroscience | 2 | 41 | 33 | 80 |
| 597 | Poznań University of Life Sciences | 4 | 41 | 177 | 327 |
| 606 | Royal Netherlands Meteorological Institute | 1 | 40 | 210 | 179 |
| 606 | Syracuse University | 2 | 40 | 2573 | 2599 |
| 606 | University of Bradford | 1 | 40 | 2121 | 1573 |
| 606 | University of Akron | 1 | 40 | 2060 | 1493 |
| 606 | University of Tulsa | 2 | 40 | 580 | 724 |
| 606 | China Institute | 1 | 40 | 3694 | 2726 |
| 606 | James Hutton Institute | 4 | 40 | 273 | 624 |
| 606 | Universidad de Oriente | 3 | 40 | 182 | 191 |
| 606 | Bernice P. Bishop Museum | 1 | 40 | 110 | 92 |
| 615 | Lanzhou University | 5 | 39 | 1588 | 2477 |
| 615 | Vietnam Academy of Science and Technology | 2 | 39 | 835 | 725 |
| 615 | Natural Resources Research Institute | 1 | 39 | 102 | 110 |
| 615 | Commonwealth Scientific and Industrial Research Organisation | 2 | 39 | 598 | 967 |
| 615 | British Science Association | 3 | 39 | 39 | 50 |
| 620 | International Livestock Research Institute | 6 | 38 | 165 | 450 |
| 620 | Daugavpils University | 1 | 38 | 139 | 53 |
| 620 | Aix-Marseille University | 2 | 38 | 3419 | 3317 |
| 620 | National University of Tucumán | 1 | 38 | 772 | 476 |
| 624 | Ben-Gurion University of the Negev | 6 | 37 | 1664 | 3677 |
| 624 | Aston University | 1 | 37 | 2377 | 1844 |
| 624 | Universidad Nacional de Cuyo | 3 | 37 | 336 | 487 |
| 624 | Roskilde University | 2 | 37 | 688 | 916 |
| 624 | Lactalis | 2 | 37 | 826 | 951 |
| 624 | Université Officielle de Bukavu | 1 | 37 | 43 | 26 |
| 624 | Jacob Blaustein Institutes for Desert Research | 5 | 37 | 129 | 416 |
| 624 | Federal Government of Belgium | 3 | 37 | 104 | 210 |
| 632 | University of Pannonia | 3 | 36 | 343 | 552 |
| 632 | University of Louisville | 1 | 36 | 4738 | 3608 |
| 632 | University of La Rochelle | 5 | 36 | 64 | 212 |
| 632 | Royal College of Surgeons of England | 4 | 36 | 739 | 1450 |
| 632 | Lawrence University | 1 | 36 | 509 | 368 |
| 632 | Penn State Beaver | 1 | 36 | 7 | 7 |
| 632 | Theodore Roosevelt High School | 5 | 36 | 1118 | 2637 |
| 632 | The Honourable Society of Lincoln's Inn | 3 | 36 | 235 | 483 |
| 640 | Benemérita Universidad Autónoma de Puebla | 3 | 35 | 506 | 525 |
| 640 | San José State University | 1 | 35 | 1755 | 1260 |
| 640 | Texas Tech University | 3 | 35 | 2397 | 3307 |
| 640 | Marine Scotland | 2 | 35 | 152 | 203 |
| 640 | University of California, Santa Cruz | 1 | 35 | 1180 | 1113 |
| 645 | Aberystwyth University | 2 | 34 | 977 | 1207 |
| 645 | Babeș-Bolyai University | 1 | 34 | 1944 | 1340 |
| 645 | McMaster University | 4 | 34 | 3885 | 5563 |
| 645 | University of Northern Colorado | 2 | 34 | 462 | 595 |
| 645 | University of Algarve | 2 | 34 | 942 | 1191 |
| 645 | Bath Spa University | 1 | 34 | 249 | 150 |
| 645 | Scientific University of the South | 1 | 34 | 351 | 117 |
| 652 | University of Padua | 2 | 33 | 5576 | 5586 |
| 652 | Nanyang Technological University | 1 | 33 | 5863 | 4532 |
| 652 | Masaryk University | 2 | 33 | 2110 | 2242 |
| 652 | University of Veterinary Medicine Vienna | 4 | 33 | 216 | 501 |
| 652 | University of Roehampton | 2 | 33 | 309 | 460 |
| 652 | Montana State University | 2 | 33 | 1595 | 2016 |
| 652 | University of Brighton | 1 | 33 | 1554 | 1320 |
| 652 | Hainan Normal University | 2 | 33 | 309 | 357 |
| 652 | list of universities in China | 3 | 33 | 2731 | 3189 |
| 652 | Laidlaw | 1 | 33 | 11 | 10 |
| 652 | Department of the Environment | 3 | 33 | 141 | 337 |
| 663 | Miguel Hernández University of Elche | 2 | 32 | 547 | 739 |
| 663 | Dartmouth College | 1 | 32 | 3086 | 2659 |
| 663 | University of Bonn | 2 | 32 | 3899 | 4141 |
| 663 | University of Neuchâtel | 1 | 32 | 1117 | 1048 |
| 663 | Kumamoto University | 1 | 32 | 2968 | 2209 |
| 663 | National University of Colombia | 2 | 32 | 2211 | 1827 |
| 663 | University of Calabria | 1 | 32 | 2297 | 1944 |
| 663 | Montclair State University | 2 | 32 | 647 | 885 |
| 663 | Kunming Institute of Zoology | 3 | 32 | 245 | 488 |
| 663 | Robert Wood Johnson Medical School | 1 | 32 | 2137 | 1864 |
| 663 | South Australian Research and Development Institute | 1 | 32 | 254 | 278 |
| 663 | University of California, Berkeley | 4 | 32 | 1859 | 3790 |
| 663 | Douglas | 4 | 32 | 433 | 1297 |
| 663 | International Union for Conservation of Nature | 1 | 32 | 136 | 158 |
| 663 | Fauna and Flora International | 3 | 32 | 24 | 73 |
| 678 | Smith College | 1 | 31 | 892 | 764 |
| 678 | Bureau of Land Management | 2 | 31 | 102 | 157 |
| 678 | University of New Brunswick | 2 | 31 | 1513 | 1857 |
| 678 | Life Sciences Institute | 2 | 31 | 537 | 880 |
| 678 | Akvaplan-niva | 1 | 31 | 104 | 110 |
| 678 | aid agency | 1 | 31 | 1336 | 980 |
| 684 | Kyung Hee University | 2 | 30 | 2773 | 2964 |
| 684 | Fryske Akademy | 1 | 30 | 26 | 9 |
| 684 | Université Cheikh Anta Diop | 1 | 30 | 664 | 470 |
| 684 | University of Porto | 3 | 30 | 2301 | 3417 |
| 684 | Maria Curie-Skłodowska University | 2 | 30 | 903 | 762 |
| 684 | North Dakota State University | 2 | 30 | 1585 | 2059 |
| 684 | Dakota State University | 2 | 30 | 2169 | 2702 |
| 684 | Federal University of Mato Grosso | 3 | 30 | 702 | 710 |
| 684 | University of Maryland, Baltimore County | 2 | 30 | 657 | 1156 |
| 684 | Universiti Sains Malaysia | 4 | 30 | 3551 | 4945 |
| 684 | St Andrews University | 1 | 30 | 81 | 114 |
| 695 | Texas A&M University | 2 | 29 | 7252 | 7216 |
| 695 | Graduate Institute of International and Development Studies | 3 | 29 | 3450 | 4415 |
| 695 | National Chung Hsing University | 2 | 29 | 1727 | 1989 |
| 695 | University of Franche-Comté | 1 | 29 | 390 | 474 |
| 695 | Norwegian Ornithological Society | 2 | 29 | 1 | 8 |
| 695 | Ohio Northern University | 1 | 29 | 169 | 158 |
| 695 | Grenfell Campus | 1 | 29 | 99 | 105 |
| 695 | Northern University | 1 | 29 | 295 | 235 |
| 695 | Universidad Politécnica de Valencia | 1 | 29 | 1166 | 1117 |
| 695 | National Pingtung University of Science & Technology | 2 | 29 | 12 | 41 |
| 705 | Wake Forest University | 2 | 28 | 1717 | 2390 |
| 705 | Oberlin College | 2 | 28 | 288 | 445 |
| 705 | University of São Paulo | 3 | 28 | 7598 | 7148 |
| 705 | Graduate University for Advanced Studies | 3 | 28 | 474 | 795 |
| 705 | Walla Walla University | 1 | 28 | 30 | 14 |
| 705 | Wrocław University of Environmental and Life Sciences | 1 | 28 | 660 | 540 |
| 705 | National University of Jujuy | 2 | 28 | 82 | 99 |
| 705 | Yukon College | 1 | 28 | 9 | 30 |
| 705 | Western Foundation of Vertebrate Zoology | 1 | 28 | 3 | 7 |
| 705 | The Graduate University for Advanced Studies | 3 | 28 | 336 | 568 |
| 715 | Université catholique de Louvain | 1 | 27 | 5174 | 4459 |
| 715 | Drexel University | 2 | 27 | 3126 | 3858 |
| 715 | Montpellier 2 University | 1 | 27 | 1698 | 1779 |
| 715 | Royal Zoological Society of Scotland | 1 | 27 | 48 | 54 |
| 715 | Universidade Federal do Pará | 3 | 27 | 1438 | 1329 |
| 715 | Universidad de Los Lagos | 1 | 27 | 393 | 219 |
| 715 | Institut national d'études supérieures agronomiques de Montpellier | 1 | 27 | 474 | 538 |
| 715 | University of Nevada, Las Vegas | 1 | 27 | 1190 | 1079 |
| 715 | Generalitat Valenciana | 1 | 27 | 178 | 201 |
| 715 | Universidade do Porto | 5 | 27 | 1273 | 2891 |
| 725 | Waseda University | 1 | 26 | 3554 | 2617 |
| 725 | Kyungsung University | 1 | 26 | 312 | 292 |
| 725 | University of Naples Federico II | 1 | 26 | 6797 | 5827 |
| 725 | Cardif | 1 | 26 | 13 | 28 |
| 725 | University of Essex | 2 | 26 | 1790 | 2349 |
| 725 | Nihon University | 1 | 26 | 3375 | 2486 |
| 725 | Tokyo Medical and Dental University | 1 | 26 | 2549 | 2121 |
| 725 | Universidade Federal de São Carlos | 5 | 26 | 1348 | 2062 |
| 725 | Smithsonian Migratory Bird Center | 2 | 26 | 53 | 89 |
| 725 | Statistics New Zealand | 3 | 26 | 4512 | 5663 |
| 725 | Island Conservation Society | 3 | 26 | 1 | 13 |
| 736 | King Mongkut's University of Technology Thonburi | 1 | 25 | 971 | 839 |
| 736 | University of Pau and Pays de l'Adour | 2 | 25 | 476 | 703 |
| 736 | Aristotle University of Thessaloniki | 1 | 25 | 5577 | 4641 |
| 736 | Banaras Hindu University | 1 | 25 | 3368 | 2707 |
| 736 | University of Technology Sydney | 1 | 25 | 3669 | 3271 |
| 736 | Portland State University | 1 | 25 | 2323 | 2156 |
| 736 | Federal University of Viçosa | 1 | 25 | 943 | 760 |
| 736 | Jahangirnagar University | 1 | 25 | 702 | 668 |
| 736 | Universidad Internacional SEK | 1 | 25 | 94 | 43 |
| 736 | Kharazmi University | 1 | 25 | 1607 | 1222 |
| 736 | Rocky Mountain Research Station | 2 | 25 | 254 | 389 |
| 736 | St. Mary's College of Maryland | 1 | 25 | 150 | 169 |
| 736 | autonomous university | 1 | 25 | 3294 | 2655 |
| 736 | Nordisk Fond for Miljø og Udvikling | 2 | 25 | 3 | 10 |
| 736 | Universitas Halu Oleo Kendari | 2 | 25 | 24 | 1 |
| 751 | Free University | 4 | 24 | 1466 | 3259 |
| 751 | Lincoln University | 3 | 24 | 533 | 1107 |
| 751 | University of Sassari | 1 | 24 | 1791 | 1816 |
| 751 | Max Planck Institute for Human Development | 1 | 24 | 365 | 493 |
| 751 | University of Milan-Bicocca | 1 | 24 | 1374 | 1473 |
| 751 | Koç University | 2 | 24 | 688 | 1101 |
| 751 | Forestry Commission Scotland | 1 | 24 | 4 | 7 |
| 751 | University North | 2 | 24 | 347 | 574 |
| 751 | Department of Biotechnology | 2 | 24 | 4294 | 4463 |
| 751 | New College | 3 | 24 | 221 | 521 |
| 751 | Università degli Studi di Milano Bicocca | 2 | 24 | 1585 | 2282 |
| 762 | University of Massachusetts Boston | 1 | 23 | 1415 | 1481 |
| 762 | University of Limpopo | 1 | 23 | 776 | 658 |
| 762 | National Marine Fisheries Service | 2 | 23 | 393 | 588 |
| 762 | Research Institute of Crop Production | 1 | 23 | 654 | 701 |
| 762 | The Citadel, The Military College of South Carolina | 1 | 23 | 206 | 183 |
| 762 | United States Army Corps of Engineers | 2 | 23 | 244 | 404 |
| 768 | Seoul National University | 1 | 22 | 7318 | 6323 |
| 768 | Concordia University | 1 | 22 | 4012 | 3435 |
| 768 | Osnabrück University | 1 | 22 | 1010 | 1137 |
| 768 | University of Konstanz | 3 | 22 | 1360 | 2411 |
| 768 | Universidad de Antioquia | 1 | 22 | 2310 | 1750 |
| 768 | University of Skövde | 1 | 22 | 440 | 565 |
| 768 | University of the Philippines Mindanao | 1 | 22 | 31 | 31 |
| 768 | Beijing Forestry University | 2 | 22 | 926 | 1227 |
| 768 | Stranmillis University College | 2 | 22 | 24 | 42 |
| 768 | Maria Mitchell Association | 1 | 22 | 2 | 4 |
| 768 | Bedford College | 2 | 22 | 135 | 206 |
| 768 | Forestry Commission England | 1 | 22 | 3 | 7 |
| 768 | Florida Fish and Wildlife Conservation Commission | 1 | 22 | 204 | 214 |
| 781 | Austrian Academy of Sciences | 2 | 21 | 920 | 1489 |
| 781 | German Cancer Research Center | 1 | 21 | 1073 | 1149 |
| 781 | University of South Africa | 2 | 21 | 1372 | 1684 |
| 781 | University of Illinois at Chicago | 1 | 21 | 6342 | 5950 |
| 781 | Curtin University | 1 | 21 | 5108 | 4840 |
| 781 | Shinshu University | 1 | 21 | 2907 | 2507 |
| 781 | University of Palermo | 2 | 21 | 2447 | 3341 |
| 781 | Weber State University | 1 | 21 | 377 | 381 |
| 781 | seznam.cz | 1 | 21 | 59 | 55 |
| 781 | Inland Norway University of Applied Sciences | 1 | 21 | 380 | 320 |
| 781 | Stockholm Resilience Centre | 2 | 21 | 123 | 277 |
| 781 | Emmanuel College | 4 | 21 | 43 | 195 |
| 781 | Royal Belgian Institute of Natural Sciences | 2 | 21 | 227 | 347 |
| 781 | United States Department of Veterans Affairs | 1 | 21 | 3644 | 3516 |
| 781 | Denver Zoo | 1 | 21 | 34 | 39 |
| 796 | Natural Resources Canada | 1 | 20 | 895 | 937 |
| 796 | Instituto Butantan | 1 | 20 | 636 | 674 |
| 796 | University of Alcalá | 2 | 20 | 1644 | 2194 |
| 796 | Trinity University | 1 | 20 | 641 | 667 |
| 796 | University of Minnesota Duluth | 1 | 20 | 726 | 833 |
| 796 | BirdLife Australia | 3 | 20 | 5 | 27 |
| 796 | Minnesota Department of Natural Resources | 1 | 20 | 118 | 138 |
| 796 | New York Genome Center | 1 | 20 | 101 | 140 |
| 804 | Niigata University | 2 | 19 | 2263 | 2607 |
| 804 | Ministry of Environment | 4 | 19 | 167 | 628 |
| 804 | University of Tsukuba | 2 | 19 | 3997 | 4429 |
| 804 | University of Southern Queensland | 1 | 19 | 1616 | 1726 |
| 804 | State University of New York | 2 | 19 | 6357 | 7461 |
| 804 | Japan Women's University | 1 | 19 | 343 | 366 |
| 804 | Hokkaido University of Education | 1 | 19 | 257 | 229 |
| 804 | University of Wollongong | 2 | 19 | 2972 | 4138 |
| 804 | University of Canberra | 1 | 19 | 1618 | 1774 |
| 804 | Omsk State University | 1 | 19 | 201 | 93 |
| 804 | A Rocha | 2 | 19 | 7 | 32 |
| 804 | Alexander von Humboldt Biological Resources Research Institute | 1 | 19 | 85 | 92 |
| 804 | American Bird Conservancy | 1 | 19 | 23 | 38 |
| 804 | The Nippon Dental University | 2 | 19 | 226 | 368 |
| 804 | University of Seychelles | 2 | 19 | 5 | 28 |
| 804 | ComUE Paris-Saclay University | 2 | 19 | 2982 | 3417 |
| 804 | The Rowett Institute | 1 | 19 | 466 | 657 |
| 804 | Axact | 2 | 19 | 926 | 1463 |
| 804 | Met Office | 3 | 19 | 241 | 451 |
| 804 | National Science Museum | 1 | 19 | 142 | 169 |
| 804 | Oeko Institut | 1 | 19 | 235 | 328 |
| 804 | Natural History Society of Northumbria | 1 | 19 | 1 | 1 |
| 826 | World Bank | 1 | 18 | 1101 | 1167 |
| 826 | University of Würzburg | 1 | 18 | 3868 | 3924 |
| 826 | Minzu University of China | 1 | 18 | 2157 | 1787 |
| 826 | Maputo University | 1 | 18 | 2438 | 1326 |
| 826 | GEOMAR Helmholtz Centre for Ocean Research Kiel | 1 | 18 | 486 | 551 |
| 826 | University of Illinois at Urbana–Champaign | 3 | 18 | 4121 | 6262 |
| 826 | Illinois State University | 2 | 18 | 1486 | 2136 |
| 826 | Santa Fe Institute | 1 | 18 | 471 | 661 |
| 826 | University of Idaho | 2 | 18 | 1534 | 2330 |
| 826 | University of Memphis | 2 | 18 | 1819 | 2790 |
| 826 | Primate Research Institute | 1 | 18 | 310 | 364 |
| 826 | Sichuan University | 1 | 18 | 5750 | 5187 |
| 826 | College of Staten Island | 1 | 18 | 1414 | 1520 |
| 826 | Guizhou Normal University | 1 | 18 | 641 | 588 |
| 826 | National Institute for Environmental Studies | 1 | 18 | 1022 | 1155 |
| 826 | Henan Normal University | 1 | 18 | 1186 | 1120 |
| 826 | National Dong Hwa University | 1 | 18 | 874 | 928 |
| 826 | Auckland Council | 1 | 18 | 31 | 46 |
| 826 | Keele University | 1 | 18 | 2239 | 2447 |
| 845 | Anheuser-Busch | 1 | 17 | 92 | 143 |
| 845 | University of Warsaw | 2 | 17 | 3119 | 3455 |
| 845 | University of Trnava | 1 | 17 | 157 | 167 |
| 845 | University of Bremen | 1 | 17 | 3167 | 3214 |
| 845 | University of Innsbruck | 2 | 17 | 2977 | 4007 |
| 845 | Bulgarian Academy of Sciences | 1 | 17 | 2583 | 2364 |
| 845 | Vrije Universiteit Amsterdam | 2 | 17 | 6235 | 7547 |
| 845 | Tokyo University of Agriculture and Technology | 2 | 17 | 1349 | 1734 |
| 845 | King Juan Carlos University | 1 | 17 | 2329 | 2072 |
| 845 | Finnish Meteorological Institute | 1 | 17 | 367 | 450 |
| 845 | Universidad de Cantabria | 1 | 17 | 1326 | 1313 |
| 856 | University of A Coruña | 1 | 16 | 1575 | 1524 |
| 856 | Calvin University | 2 | 16 | 142 | 347 |
| 856 | Fukushima University | 1 | 16 | 374 | 343 |
| 856 | University of Seville | 2 | 16 | 3680 | 4381 |
| 856 | Idaho State University | 1 | 16 | 1083 | 1307 |
| 856 | Hammersmith Hospital | 1 | 16 | 1429 | 1598 |
| 856 | Indiana Wesleyan University | 1 | 16 | 76 | 114 |
| 856 | Maryland Department of Natural Resources | 2 | 16 | 52 | 101 |
| 856 | Monell Chemical Senses Center | 1 | 16 | 255 | 410 |
| 856 | Prairie Research Institute | 3 | 16 | 100 | 295 |
| 856 | Sorbonne University | 5 | 16 | 1705 | 3345 |
| 856 | Colegio de Postgraduados | 1 | 16 | 507 | 474 |
| 856 | National Museums Scotland | 3 | 16 | 45 | 122 |
| 869 | Tsinghua University | 2 | 15 | 5628 | 6433 |
| 869 | University of Tübingen | 1 | 15 | 4561 | 4663 |
| 869 | University of Szeged | 1 | 15 | 2504 | 2606 |
| 869 | Arcadis | 1 | 15 | 147 | 206 |
| 869 | Wellcome | 1 | 15 | 1720 | 2014 |
| 869 | College of Wooster | 1 | 15 | 320 | 387 |
| 869 | Federation University Australia | 1 | 15 | 782 | 959 |
| 869 | San Francisco State University | 1 | 15 | 1756 | 1950 |
| 869 | National University of Malaysia | 1 | 15 | 3681 | 3468 |
| 869 | French National Institute of Health and Medical Research | 2 | 15 | 5152 | 5812 |
| 869 | University of Szczecin | 2 | 15 | 611 | 688 |
| 869 | University of Crete | 2 | 15 | 1786 | 3019 |
| 869 | Ducks Unlimited | 1 | 15 | 59 | 86 |
| 869 | Field Studies Council | 1 | 15 | 13 | 19 |
| 869 | Golden Gate National Parks Conservancy | 2 | 15 | 1 | 3 |
| 869 | Golden Gate Raptor Observatory | 2 | 15 | 2 | 9 |
| 869 | Indonesian Institute of Sciences | 2 | 15 | 456 | 738 |
| 869 | Natural History Museum of Crete | 2 | 15 | 23 | 80 |
| 869 | Harvard Graduate School of Education | 1 | 15 | 336 | 455 |
| 869 | The James Cancer Hospital | 2 | 15 | 948 | 1769 |
| 869 | Nord University | 1 | 15 | 664 | 778 |
| 869 | SMEC Holdings | 1 | 15 | 17 | 33 |
| 869 | Museum für Naturkunde | 1 | 15 | 365 | 439 |
| 869 | Volda University College | 1 | 15 | 88 | 82 |
| 869 | Robert Gordon's College | 1 | 15 | 66 | 65 |
| 894 | Kazan Federal University | 2 | 14 | 1182 | 1498 |
| 894 | Technion – Israel Institute of Technology | 1 | 14 | 5041 | 5190 |
| 894 | University of Zielona Góra | 2 | 14 | 346 | 554 |
| 894 | Metsähallitus | 1 | 14 | 43 | 81 |
| 894 | Binghamton University | 2 | 14 | 1407 | 2342 |
| 894 | Tohoku University | 1 | 14 | 6163 | 5749 |
| 894 | Hungarian Ornithological and Nature Conservation Society | 2 | 14 | 9 | 31 |
| 894 | University of Lleida | 2 | 14 | 434 | 982 |
| 894 | Konrad Lorenz Institute of Ethology | 2 | 14 | 31 | 63 |
| 894 | Universidade do Estado do Rio de Janeiro | 1 | 14 | 2928 | 2100 |
| 894 | Limnological Institute | 1 | 14 | 349 | 416 |
| 894 | Charing Cross Hospital Medical School | 2 | 14 | 68 | 142 |
| 894 | East–West Center | 1 | 14 | 153 | 227 |
| 894 | Federal University of Uberlândia | 1 | 14 | 2426 | 1720 |
| 894 | Universidad Nacional del Sur | 1 | 14 | 1200 | 1120 |
| 894 | Fujita Health University | 1 | 14 | 1409 | 1427 |
| 894 | Pacific Adventist University | 1 | 14 | 8 | 10 |
| 894 | Southern Research | 2 | 14 | 451 | 878 |
| 894 | Western Transportation Institute | 1 | 14 | 37 | 76 |
| 894 | Women's Health Research Institute | 1 | 14 | 147 | 219 |
| 894 | Florida Museum of Natural History | 2 | 14 | 239 | 403 |
| 915 | University of Kiel | 1 | 13 | 2630 | 3178 |
| 915 | Utah State University | 2 | 13 | 2222 | 3620 |
| 915 | New Mexico State University | 2 | 13 | 1543 | 2662 |
| 915 | Fairfield University | 1 | 13 | 472 | 606 |
| 915 | Loughborough University | 1 | 13 | 3743 | 3972 |
| 915 | Pontificia Universidad Católica del Ecuador | 1 | 13 | 559 | 480 |
| 915 | Queens College | 2 | 13 | 858 | 1679 |
| 915 | Rollins College | 1 | 13 | 1855 | 2325 |
| 915 | University of Valle | 1 | 13 | 1532 | 1409 |
| 915 | Bayero University Kano | 1 | 13 | 823 | 813 |
| 915 | Millikin University | 1 | 13 | 67 | 101 |
| 915 | Instituto Biológico | 1 | 13 | 728 | 876 |
| 915 | East Bay Regional Park District | 1 | 13 | 6 | 12 |
| 915 | Archbold Biological Station | 1 | 13 | 97 | 109 |
| 915 | Australian Marine Mammal Research Centre | 1 | 13 | 6 | 16 |
| 915 | National Center for Ecological Analysis and Synthesis | 1 | 13 | 108 | 196 |
| 915 | Graduate Center, CUNY | 2 | 13 | 1854 | 3264 |
| 915 | Université de La Réunion | 2 | 13 | 340 | 755 |
| 915 | Manchester University | 3 | 13 | 184 | 952 |
| 934 | Institute for Advanced Study | 1 | 12 | 3456 | 3813 |
| 934 | Irkutsk State University | 1 | 12 | 450 | 369 |
| 934 | Mount Allison University | 2 | 12 | 258 | 608 |
| 934 | Sciences Po | 1 | 12 | 1058 | 1127 |
| 934 | Woods Hole Oceanographic Institution | 4 | 12 | 394 | 968 |
| 934 | Old Dominion University | 1 | 12 | 2557 | 3017 |
| 934 | Universidad San Francisco de Quito | 1 | 12 | 551 | 657 |
| 934 | Central Research Institute of Electric Power Industry | 1 | 12 | 632 | 707 |
| 934 | Escuela Superior Politécnica del Litoral | 1 | 12 | 336 | 453 |
| 934 | Universidade Federal de Mato Grosso do Sul | 1 | 12 | 1692 | 1196 |
| 934 | Jožef Stefan Institute | 1 | 12 | 1028 | 1284 |
| 934 | Anhanguera-Uniderp University | 1 | 12 | 39 | 29 |
| 934 | Universidade Federal da Bahia | 1 | 12 | 3507 | 2751 |
| 934 | Queensland Museum | 2 | 12 | 111 | 221 |
| 934 | Brazilian Agricultural Research Corporation | 1 | 12 | 1880 | 1921 |
| 934 | University of the West of England | 1 | 12 | 1990 | 2432 |
| 950 | Wesleyan University | 3 | 11 | 578 | 1590 |
| 950 | Ohio Wesleyan University | 3 | 11 | 46 | 220 |
| 950 | University of Latvia | 1 | 11 | 1273 | 1533 |
| 950 | Southern University | 2 | 11 | 2728 | 4181 |
| 950 | National University of Misiones | 1 | 11 | 211 | 256 |
| 950 | Linköping University | 1 | 11 | 4358 | 5059 |
| 950 | University of Basilicata | 1 | 11 | 995 | 1379 |
| 950 | Villanova University | 1 | 11 | 1615 | 1975 |
| 950 | Hendrix College | 1 | 11 | 108 | 183 |
| 950 | Institut universitaire de France | 3 | 11 | 657 | 1734 |
| 950 | University of Lethbridge | 1 | 11 | 1241 | 1656 |
| 950 | University of South Wales | 2 | 11 | 307 | 788 |
| 950 | California Department of Fish and Wildlife | 1 | 11 | 158 | 226 |
| 950 | Campus university | 1 | 11 | 500 | 841 |
| 950 | Instituto de Biología Subtropical | 1 | 11 | 79 | 123 |
| 950 | Nature Conservation Foundation | 2 | 11 | 47 | 124 |
| 950 | Universidade Federal de Goiás | 1 | 11 | 2957 | 2255 |
| 950 | University Centre of the Westfjords | 1 | 11 | 10 | 12 |
| 950 | Old Road Campus | 1 | 11 | 951 | 1350 |
| 950 | The Stables | 1 | 11 | 8 | 24 |
| 950 | Christian Albrechts Universität zu Kiel | 1 | 11 | 1074 | 1500 |
| 950 | Universidad de la República | 1 | 11 | 2368 | 2273 |
| 972 | University of Lancaster | 1 | 10 | 894 | 1262 |
| 972 | University of Salamanca | 1 | 10 | 3343 | 3435 |
| 972 | Tours University | 1 | 10 | 1405 | 1777 |
| 972 | Alfred Wegener Institute for Polar and Marine Research | 1 | 10 | 625 | 809 |
| 972 | Middle East Technical University | 2 | 10 | 1903 | 3227 |
| 972 | Austral University of Chile | 1 | 10 | 1315 | 1557 |
| 972 | Tokai University | 1 | 10 | 2603 | 2740 |
| 972 | Hatfield Marine Science Center | 1 | 10 | 151 | 236 |
| 972 | University of the Free State | 2 | 10 | 1356 | 2352 |
| 972 | Long Island University | 1 | 10 | 854 | 1230 |
| 972 | Kenyon College | 3 | 10 | 81 | 410 |
| 972 | Mammal Research Institute of the Polish Academy of Sciences | 1 | 10 | 248 | 343 |
| 972 | Temple College | 1 | 10 | 52 | 71 |
| 972 | Gurukul Kangri Vishwavidyalaya | 1 | 10 | 100 | 203 |
| 972 | University of California, San Francisco | 1 | 10 | 3776 | 4287 |
| 972 | Royal Museum for Central Africa | 1 | 10 | 241 | 364 |
| 972 | Okinawa Prefecture | 1 | 10 | 2413 | 2763 |
| 972 | Ben Gurion University of the Negev | 1 | 10 | 1626 | 2224 |
| 972 | Colorado State University Pueblo | 1 | 10 | 133 | 230 |
| 991 | Bordeaux Montaigne University | 1 | 9 | 161 | 183 |
| 991 | University of Bordeaux | 1 | 9 | 3958 | 4524 |
| 991 | National Centre for Biological Sciences | 1 | 9 | 347 | 569 |
| 991 | Verizon | 1 | 9 | 130 | 225 |
| 991 | American University of Beirut | 1 | 9 | 2869 | 3523 |
| 991 | University of Huelva | 1 | 9 | 1137 | 1385 |
| 991 | Bates College | 1 | 9 | 321 | 538 |
| 991 | Hongdu | 1 | 9 | 4819 | 5024 |
| 991 | Eastern Connecticut State University | 1 | 9 | 148 | 291 |
| 991 | National Polytechnic Institute | 1 | 9 | 3060 | 3215 |