| 1 | Ecology (University of Lausanne) | 178 | 33.0K | 8 | 5 |
| 2 | Biology (University of Lausanne) | 10 | 22.7K | 100 | 4 |
| 3 | Ecology (University of Bern) | 94 | 20.9K | 11 | 5 |
| 4 | Evolutionary Biology (Uppsala University) | 202 | 20.6K | 9 | 11 |
| 5 | History (University of Oslo) | 76 | 17.6K | 11 | 3 |
| 6 | Biology (University of California, Berkeley) | 145 | 16.5K | 26 | 26 |
| 7 | Zoology (University of British Columbia) | 131 | 14.1K | 16 | 9 |
| 8 | Zoology (Brigham Young University) | 7 | 11.3K | 36 | 6 |
| 9 | Biology (Stanford University) | 62 | 10.7K | 33 | 26 |
| 10 | Plant Pathology (University of California, Berkeley) | 5 | 10.7K | 27 | 4 |
| 11 | Biology (University of Guelph) | 105 | 10.0K | 20 | 15 |
| 12 | Biology (Duke University) | 104 | 9.8K | 29 | 30 |
| 13 | Computer Science (Brigham Young University) | 2 | 9.3K | 59 | 2 |
| 14 | Ecology (University of California, Los Angeles) | 119 | 9.1K | 12 | 9 |
| 15 | Biology (University of Edinburgh) | 26 | 8.7K | 45 | 13 |
| 16 | Zoology (University of Queensland) | 37 | 8.6K | 22 | 8 |
| 17 | Zoology (University of Oxford) | 102 | 8.6K | 38 | 38 |
| 18 | Ecology (Cornell University) | 123 | 8.3K | 12 | 15 |
| 19 | Evolutionary Biology (Cornell University) | 124 | 8.2K | 8 | 11 |
| 20 | Evolutionary Biology (University of California, Los Angeles) | 109 | 8.0K | 12 | 9 |
| 21 | Evolutionary Biology (University of Edinburgh) | 117 | 7.8K | 8 | 15 |
| 22 | Biology (Indiana University Bloomington) | 73 | 7.7K | 33 | 23 |
| 23 | History (University of Copenhagen) | 70 | 7.6K | 15 | 12 |
| 24 | Botany (University of British Columbia) | 101 | 7.5K | 25 | 19 |
| 25 | Biology (Dalhousie University) | 92 | 7.5K | 18 | 12 |
| 26 | Ecology (Lund University) | 80 | 7.4K | 26 | 16 |
| 27 | Biology (University of New Mexico) | 55 | 7.4K | 40 | 16 |
| 28 | Zoology (University of Cambridge) | 89 | 7.4K | 35 | 32 |
| 29 | Evolutionary Biology (Harvard University) | 111 | 7.3K | 17 | 21 |
| 30 | Biology (Pennsylvania State University) | 70 | 7.0K | 31 | 22 |
| 31 | Ecology (University of Oregon) | 31 | 6.8K | 11 | 7 |
| 32 | Evolutionary Biology (University of Toronto) | 86 | 6.6K | 11 | 9 |
| 33 | Ecology (Uppsala University) | 102 | 6.5K | 11 | 12 |
| 34 | Ecology (University of Toronto) | 87 | 6.5K | 11 | 9 |
| 35 | Ecology (University of Tartu) | 30 | 6.4K | 45 | 10 |
| 36 | Evolutionary Biology (University of Zurich) | 78 | 6.3K | 14 | 11 |
| 37 | Ecology (University of Arizona) | 70 | 6.3K | 16 | 17 |
| 38 | Botany (Australian National University) | 37 | 6.2K | 16 | 9 |
| 38 | Zoology (Australian National University) | 37 | 6.2K | 19 | 10 |
| 40 | Biology (Australian National University) | 88 | 6.1K | 26 | 29 |
| 41 | Biology (Colorado State University) | 55 | 5.8K | 37 | 29 |
| 42 | Ecology (Montana State University) | 11 | 5.7K | 48 | 7 |
| 43 | Biology (University of Texas at Austin) | 89 | 5.6K | 20 | 20 |
| 44 | Botany (University of Vienna) | 54 | 5.6K | 17 | 8 |
| 45 | Evolutionary Biology (University of Arizona) | 66 | 5.6K | 16 | 23 |
| 45 | Zoology (University of Otago) | 87 | 5.6K | 18 | 18 |
| 47 | Biology (Ludwig-Maximilians-Universität München) | 40 | 5.6K | 39 | 16 |
| 48 | Zoology (University of California, Berkeley) | 99 | 5.5K | 24 | 21 |
| 49 | Biology (University of Helsinki) | 13 | 5.4K | 81 | 17 |
| 50 | Biology (ETH Zürich) | 74 | 5.3K | 28 | 27 |
| 51 | Genetics (University of Cambridge) | 36 | 5.3K | 36 | 31 |
| 52 | Genetics (Uppsala University) | 106 | 5.2K | 25 | 35 |
| 53 | Ecology (University of California, Santa Cruz) | 73 | 5.1K | 10 | 9 |
| 54 | Biology (University of Copenhagen) | 93 | 5.0K | 32 | 39 |
| 55 | History (University of Tartu) | 14 | 5.0K | 11 | 7 |
| 56 | Biology (University of California, Davis) | 54 | 5.0K | 40 | 34 |
| 57 | Biology (University of St Andrews) | 52 | 4.9K | 23 | 17 |
| 58 | Evolutionary Biology (University of California, Santa Cruz) | 71 | 4.9K | 11 | 9 |
| 59 | Biology (University of Turku) | 54 | 4.8K | 32 | 16 |
| 60 | Ecology (University of Michigan) | 92 | 4.7K | 13 | 16 |
| 61 | Evolutionary Biology (Yale University) | 51 | 4.6K | 17 | 13 |
| 62 | Ecology (Yale University) | 50 | 4.6K | 19 | 15 |
| 63 | Biology (University of Oslo) | 51 | 4.6K | 37 | 22 |
| 64 | Evolutionary Biology (University of Michigan) | 89 | 4.5K | 13 | 17 |
| 65 | Botany (University of Wyoming) | 53 | 4.5K | 6 | 11 |
| 66 | Environmental Science (University of California, Berkeley) | 85 | 4.4K | 33 | 53 |
| 67 | Biology (McGill University) | 54 | 4.3K | 35 | 29 |
| 67 | Plant Pathology (University of Florida) | 10 | 4.3K | 59 | 15 |
| 69 | Neuroscience (University of Oregon) | 4 | 4.3K | 93 | 17 |
| 70 | Biology (University of York) | 45 | 4.2K | 52 | 45 |
| 71 | Computational Biology (Cornell University) | 10 | 4.2K | 39 | 15 |
| 72 | Biology (Ohio State University) | 59 | 4.2K | 32 | 27 |
| 73 | Ecology (Bangor University) | 28 | 4.1K | 5 | 3 |
| 74 | Dynamics (University of Amsterdam) | 38 | 4.1K | 59 | 32 |
| 75 | Statistics (Cornell University) | 7 | 4.0K | 45 | 12 |
| 76 | Biology (Lund University) | 84 | 4.0K | 20 | 22 |
| 77 | Marine Ecology (University of Gothenburg) | 15 | 4.0K | 21 | 4 |
| 78 | Plant Pathology (Swedish University of Agricultural Sciences) | 10 | 4.0K | 37 | 8 |
| 79 | Genetics (Bangor University) | 18 | 3.9K | 7 | 3 |
| 80 | Human Genetics (University of Chicago) | 10 | 3.9K | 71 | 19 |
| 81 | Ecological Genetics (University of Helsinki) | 51 | 3.9K | 6 | 5 |
| 82 | Evolutionary Biology (University of St Andrews) | 10 | 3.8K | 16 | 6 |
| 83 | Genetics (Ludwig-Maximilians-Universität München) | 2 | 3.7K | 79 | 7 |
| 84 | Neuroscience (University of Gothenburg) | 2 | 3.7K | 590 | 54 |
| 84 | Physiology (University of Gothenburg) | 2 | 3.7K | 606 | 50 |
| 86 | Microbiology (Cornell University) | 7 | 3.7K | 160 | 36 |
| 87 | Biology (University of Tübingen) | 3 | 3.7K | 113 | 9 |
| 88 | Biology (Imperial College London) | 33 | 3.7K | 59 | 41 |
| 89 | Public Health (University of Toronto) | 1 | 3.6K | 1,103 | 66 |
| 89 | Public Health (Dalla Lana School of Public Health) | 1 | 3.6K | 1,064 | 58 |
| 91 | Biology (Uppsala University) | 61 | 3.6K | 11 | 15 |
| 92 | Ecology (University of California, Davis) | 64 | 3.5K | 18 | 28 |
| 93 | Ecology (Ohio State University) | 55 | 3.5K | 19 | 13 |
| 94 | Ecology (University of Colorado Boulder) | 49 | 3.5K | 17 | 22 |
| 95 | Biology (University of Western Australia) | 56 | 3.5K | 32 | 43 |
| 96 | Genetics (University of Georgia) | 65 | 3.4K | 21 | 27 |
| 97 | Ecology (University of Canberra) | 38 | 3.4K | 19 | 9 |
| 98 | Biology (University of Konstanz) | 60 | 3.4K | 29 | 37 |
| 99 | Statistics (University of Chicago) | 2 | 3.3K | 190 | 19 |
| 100 | Ecology (Tel Aviv University) | 2 | 3.3K | 53 | 8 |
| 100 | Molecular Biology (Tel Aviv University) | 2 | 3.3K | 109 | 9 |
| 102 | Microbiology (University of Chicago) | 2 | 3.3K | 85 | 18 |
| 103 | Human Genetics (Cornell University) | 1 | 3.3K | 48 | 3 |
| 103 | Statistics (University College Cork) | 1 | 3.3K | 82 | 3 |
| 103 | Microbiology (University College Cork) | 1 | 3.3K | 241 | 31 |
| 103 | Human Genetics (University College Cork) | 1 | 3.3K | 11 | 3 |
| 103 | History (University of Florida) | 30 | 3.3K | 42 | 15 |
| 103 | Computational Biology (University of Chicago) | 1 | 3.3K | 9 | 2 |
| 103 | Computational Biology (University College Cork) | 1 | 3.3K | 1 | 1 |
| 110 | Evolutionary Biology (University of Colorado Boulder) | 47 | 3.3K | 17 | 22 |
| 111 | Biology (University of Maryland) | 43 | 3.2K | 22 | 28 |
| 112 | Zoology (Stockholm University) | 53 | 3.1K | 24 | 19 |
| 113 | Genetics (University College London) | 52 | 3.1K | 28 | 32 |
| 114 | Ecology (Australian National University) | 51 | 3.1K | 15 | 15 |
| 115 | Biology (James Cook University) | 40 | 3.0K | 27 | 27 |
| 116 | Ecology (Princeton University) | 47 | 2.9K | 25 | 37 |
| 117 | Biology (University of Oregon) | 12 | 2.9K | 54 | 19 |
| 118 | Biology (San Francisco State University) | 31 | 2.9K | 17 | 13 |
| 119 | Biology (University of Washington) | 41 | 2.9K | 53 | 55 |
| 120 | Biology (University of Alaska Fairbanks) | 41 | 2.8K | 33 | 30 |
| 121 | Zoology (Harvard University) | 47 | 2.8K | 25 | 26 |
| 122 | Evolutionary Biology (University of Adelaide) | 31 | 2.8K | 14 | 7 |
| 123 | Ecology (Aarhus University) | 36 | 2.8K | 20 | 13 |
| 124 | Genetics (Aarhus University) | 43 | 2.8K | 35 | 29 |
| 125 | Evolutionary Biology (Princeton University) | 46 | 2.8K | 28 | 37 |
| 126 | Botany (University of Zurich) | 19 | 2.7K | 23 | 12 |
| 127 | Biology (University of Antwerp) | 35 | 2.7K | 54 | 40 |
| 128 | Biology (University of Leeds) | 40 | 2.6K | 45 | 47 |
| 129 | Zoology (Oregon State University) | 27 | 2.6K | 33 | 15 |
| 130 | Oceanography (University of California, San Diego) | 44 | 2.6K | 115 | 111 |
| 131 | Biology (University of New Brunswick) | 16 | 2.6K | 59 | 22 |
| 132 | Ecology (University of Chicago) | 39 | 2.6K | 17 | 27 |
| 133 | Ecology (University of Innsbruck) | 47 | 2.6K | 6 | 12 |
| 134 | Biology (University of Montana) | 31 | 2.6K | 17 | 16 |
| 135 | Botany (University of Toronto) | 11 | 2.5K | 46 | 23 |
| 136 | Ecology (Swedish University of Agricultural Sciences) | 43 | 2.5K | 47 | 45 |
| 137 | Biomedical Sciences (Cardiff University) | 11 | 2.5K | 7 | 4 |
| 138 | Ecology (Katholieke Universiteit Leuven) | 39 | 2.5K | 18 | 11 |
| 139 | Biology (Boston University) | 37 | 2.5K | 49 | 54 |
| 140 | Epidemiology (Imperial College London) | 25 | 2.5K | 111 | 95 |
| 141 | Ecology (University of Wyoming) | 33 | 2.4K | 6 | 9 |
| 142 | Genomics (University of Copenhagen) | 34 | 2.4K | 8 | 11 |
| 143 | Biology (Western University) | 28 | 2.4K | 49 | 19 |
| 144 | Biology (Drexel University) | 19 | 2.4K | 11 | 7 |
| 145 | Ecology (University of East Anglia) | 23 | 2.4K | 16 | 14 |
| 146 | Biology (University of Rochester) | 33 | 2.4K | 50 | 58 |
| 147 | Biology (McMaster University) | 40 | 2.4K | 54 | 34 |
| 148 | Ecology (University of Minnesota) | 36 | 2.4K | 33 | 42 |
| 149 | Biology (University of Oklahoma) | 30 | 2.3K | 30 | 29 |
| 150 | Botany (Oregon State University) | 19 | 2.3K | 60 | 35 |
| 150 | Plant Pathology (Oregon State University) | 19 | 2.3K | 60 | 35 |
| 152 | Plant Pathology (North Carolina State University) | 23 | 2.3K | 44 | 20 |
| 153 | Biology (Michigan State University) | 35 | 2.3K | 37 | 37 |
| 154 | Ecology (Rutgers University) | 35 | 2.3K | 20 | 23 |
| 155 | Conservation Biology (Uppsala University) | 32 | 2.3K | 4 | 3 |
| 156 | Biology (Utah State University) | 46 | 2.3K | 19 | 20 |
| 157 | Ecology (University of California, Irvine) | 39 | 2.3K | 33 | 51 |
| 158 | Ecology (Iowa State University) | 38 | 2.3K | 34 | 30 |
| 159 | Marine Biology (University of Hawaiʻi at Mānoa) | 32 | 2.3K | 14 | 13 |
| 160 | Ecology (University of Kaiserslautern) | 8 | 2.3K | 14 | 8 |
| 161 | Biology (University of Minnesota) | 25 | 2.2K | 60 | 44 |
| 162 | Evolutionary Biology (University of California, Irvine) | 38 | 2.2K | 26 | 35 |
| 163 | Biology (University of California, Los Angeles) | 20 | 2.2K | 80 | 57 |
| 164 | Zoology (Michigan State University) | 13 | 2.2K | 81 | 25 |
| 165 | Biomedical Sciences (University of Queensland) | 7 | 2.2K | 196 | 36 |
| 166 | Botany (University of Oslo) | 7 | 2.2K | 15 | 5 |
| 167 | Ecology (Colorado State University) | 27 | 2.2K | 70 | 62 |
| 168 | Genetics (La Trobe University) | 31 | 2.2K | 18 | 16 |
| 169 | Biology (Norwegian University of Science and Technology) | 47 | 2.1K | 21 | 32 |
| 170 | Genetics (Australian National University) | 26 | 2.1K | 19 | 19 |
| 171 | Biology (University of Victoria) | 24 | 2.1K | 47 | 25 |
| 172 | Zoology (University of Michigan) | 35 | 2.1K | 28 | 20 |
| 172 | Population Genetics (University of Bern) | 22 | 2.1K | 5 | 11 |
| 174 | Botany (University of Texas at Austin) | 3 | 2.1K | 69 | 14 |
| 175 | History (University of Vienna) | 2 | 2.1K | 32 | 3 |
| 176 | Ecology (University of Exeter) | 48 | 2.0K | 23 | 27 |
| 177 | Biology (University of Oulu) | 35 | 2.0K | 19 | 21 |
| 178 | Biology (University of Fribourg) | 37 | 2.0K | 18 | 23 |
| 179 | Biology (Aarhus University) | 37 | 2.0K | 24 | 26 |
| 180 | Biology (Kansas State University) | 33 | 2.0K | 45 | 43 |
| 181 | Genetics (University of Geneva) | 22 | 2.0K | 35 | 35 |
| 182 | Ecology (Brown University) | 21 | 2.0K | 21 | 18 |
| 183 | Ecology (University of Groningen) | 30 | 2.0K | 25 | 21 |
| 184 | Zoology (University of Melbourne) | 21 | 1.9K | 41 | 22 |
| 185 | Marine Biology (University of Miami) | 22 | 1.9K | 20 | 13 |
| 186 | Ecology (Rice University) | 23 | 1.9K | 11 | 8 |
| 186 | Evolutionary Biology (Rice University) | 23 | 1.9K | 11 | 8 |
| 188 | Zoology (University of Aberdeen) | 31 | 1.9K | 40 | 30 |
| 189 | Biology (University of Louisiana at Lafayette) | 21 | 1.9K | 30 | 14 |
| 190 | Biology (University of Queensland) | 16 | 1.9K | 45 | 37 |
| 191 | Biology (Virginia Commonwealth University) | 23 | 1.9K | 13 | 10 |
| 192 | Biology (Iowa State University) | 29 | 1.9K | 27 | 29 |
| 193 | Microbiology (University of Oklahoma) | 17 | 1.9K | 79 | 60 |
| 194 | Genomics (University of Oklahoma) | 16 | 1.9K | 24 | 26 |
| 194 | Conservation Biology (University of Queensland) | 13 | 1.9K | 3 | 6 |
| 196 | Marine Biology (University of Hawaii) | 14 | 1.9K | 25 | 13 |
| 197 | Genomics (University of Guelph) | 35 | 1.9K | 5 | 8 |
| 198 | Biology (Leiden University) | 29 | 1.8K | 50 | 63 |
| 199 | Evolutionary Biology (Brown University) | 13 | 1.8K | 31 | 19 |
| 200 | Biology (University of Georgia) | 43 | 1.8K | 22 | 41 |
| 201 | Ecology (University of Helsinki) | 20 | 1.8K | 23 | 17 |
| 202 | Marine Biology (University of California, Santa Barbara) | 27 | 1.8K | 37 | 39 |
| 203 | Biology (University of Texas at Arlington) | 30 | 1.8K | 17 | 14 |
| 204 | History (University of Minnesota) | 13 | 1.8K | 23 | 14 |
| 205 | Animal Science (University of California, Davis) | 32 | 1.8K | 40 | 33 |
| 206 | Microbiology (University of Exeter) | 1 | 1.8K | 20 | 1 |
| 206 | Computer Science (University of Kaiserslautern) | 1 | 1.8K | 95 | 1 |
| 208 | Biology (University of South Florida) | 17 | 1.8K | 60 | 31 |
| 208 | Zoology (University of Guelph) | 24 | 1.8K | 42 | 26 |
| 210 | Atmospheric Science (University of Miami) | 18 | 1.7K | 89 | 67 |
| 211 | Biology (University of Toronto) | 20 | 1.7K | 66 | 44 |
| 212 | Zoology (University of Wyoming) | 22 | 1.7K | 48 | 25 |
| 212 | Physiology (University of Wyoming) | 22 | 1.7K | 47 | 25 |
| 214 | Biology (University of California, Riverside) | 27 | 1.7K | 46 | 51 |
| 214 | Crop Science (Swedish University of Agricultural Sciences) | 1 | 1.7K | 60 | 5 |
| 216 | Biology (University of Ljubljana) | 18 | 1.7K | 49 | 20 |
| 217 | Zoology (Charles University) | 40 | 1.7K | 18 | 15 |
| 218 | Biology (Marine Biological Laboratory) | 22 | 1.7K | 60 | 57 |
| 219 | Chemistry (University of Colorado Boulder) | 4 | 1.7K | 217 | 93 |
| 219 | Biochemistry (University of Colorado Boulder) | 4 | 1.7K | 176 | 80 |
| 221 | Biology (Texas A&M University) | 32 | 1.6K | 42 | 48 |
| 222 | Genetics (University of Melbourne) | 20 | 1.6K | 31 | 31 |
| 223 | Biology (University College Dublin) | 20 | 1.6K | 61 | 37 |
| 224 | Ecology (University of Antwerp) | 30 | 1.6K | 26 | 25 |
| 225 | Anatomy (University of Otago) | 39 | 1.6K | 20 | 23 |
| 226 | Conservation Biology (University of Toronto) | 13 | 1.6K | 16 | 8 |
| 227 | Genetics (University of Oslo) | 19 | 1.6K | 29 | 24 |
| 228 | Genetics (University of Wisconsin–Madison) | 15 | 1.6K | 67 | 51 |
| 229 | Biology (University of Nevada, Reno) | 32 | 1.6K | 20 | 20 |
| 230 | Ecology (University of Kansas) | 37 | 1.6K | 36 | 49 |
| 230 | Zoology (Stellenbosch University) | 35 | 1.6K | 45 | 50 |
| 232 | Ecology (Lanzhou University) | 25 | 1.6K | 26 | 14 |
| 233 | Ecology (University of Connecticut) | 25 | 1.6K | 45 | 38 |
| 233 | Evolutionary Biology (University of Connecticut) | 25 | 1.6K | 44 | 38 |
| 235 | Ecology (University of Georgia) | 45 | 1.6K | 49 | 75 |
| 236 | Biology (Texas State University) | 17 | 1.6K | 33 | 8 |
| 237 | Genetics (Norwegian University of Life Sciences) | 16 | 1.5K | 13 | 7 |
| 238 | Environmental Health (University of Georgia) | 18 | 1.5K | 22 | 20 |
| 239 | Ecology (University of California, Santa Barbara) | 27 | 1.5K | 37 | 43 |
| 240 | Biology (Georgia Tech) | 14 | 1.5K | 59 | 42 |
| 241 | Genetics (University of Turku) | 19 | 1.5K | 15 | 10 |
| 242 | History (University of Helsinki) | 31 | 1.5K | 35 | 29 |
| 243 | Biology (Brigham Young University) | 26 | 1.5K | 25 | 25 |
| 244 | Biology (University of Potsdam) | 41 | 1.5K | 24 | 38 |
| 244 | Biochemistry (University of Potsdam) | 41 | 1.5K | 25 | 37 |
| 246 | Physiology (University of Turku) | 18 | 1.5K | 63 | 45 |
| 247 | Evolutionary Biology (University of Kansas) | 35 | 1.5K | 34 | 40 |
| 248 | Biology (City University of New York) | 27 | 1.4K | 28 | 30 |
| 249 | Genomics (University of Montana) | 20 | 1.4K | 3 | 3 |
| 250 | Ecology (University of Fribourg) | 23 | 1.4K | 7 | 8 |
| 251 | Ecology (University of Queensland) | 11 | 1.4K | 34 | 22 |
| 252 | Genetics (University of Nottingham) | 18 | 1.4K | 25 | 26 |
| 253 | Biology (University of Virginia) | 29 | 1.4K | 49 | 67 |
| 254 | Botany (Charles University) | 22 | 1.4K | 30 | 23 |
| 255 | Zoology (University of Leicester) | 11 | 1.4K | 20 | 7 |
| 256 | Biology (Carleton University) | 18 | 1.4K | 87 | 51 |
| 257 | Ecology (University of Copenhagen) | 28 | 1.4K | 24 | 30 |
| 258 | Mathematics (University of Helsinki) | 10 | 1.4K | 63 | 19 |
| 258 | Statistics (University of Helsinki) | 10 | 1.4K | 54 | 17 |
| 260 | Education (University of Florida) | 12 | 1.4K | 247 | 102 |
| 261 | Genetics (Cornell University) | 22 | 1.3K | 63 | 86 |
| 262 | Genetics (Wageningen University & Research) | 20 | 1.3K | 39 | 25 |
| 263 | Environmental Science (University of Jyväskylä) | 35 | 1.3K | 47 | 49 |
| 264 | Public Health (Imperial College London) | 9 | 1.3K | 297 | 150 |
| 265 | Biology (San Diego State University) | 23 | 1.3K | 54 | 61 |
| 266 | Botany (Stellenbosch University) | 32 | 1.3K | 43 | 48 |
| 267 | Zoology (University of Pretoria) | 18 | 1.3K | 81 | 44 |
| 268 | Evolutionary Biology (Pompeu Fabra University) | 19 | 1.3K | 17 | 13 |
| 269 | Zoology (University of Tasmania) | 20 | 1.3K | 32 | 19 |
| 270 | History (University of Toronto) | 24 | 1.3K | 25 | 18 |
| 271 | Biology (Ghent University) | 36 | 1.3K | 38 | 66 |
| 272 | Ecology (Université libre de Bruxelles) | 35 | 1.3K | 13 | 16 |
| 273 | Plant Ecology (Uppsala University) | 21 | 1.3K | 13 | 11 |
| 274 | Biomedical Sciences (Durham University) | 22 | 1.3K | 20 | 35 |
| 275 | Biotechnology (Tabriz University of Medical Sciences) | 15 | 1.3K | 271 | 154 |
| 276 | Genomics (New York University) | 18 | 1.3K | 28 | 36 |
| 276 | Systems Biology (New York University) | 18 | 1.3K | 25 | 31 |
| 278 | Biology (Katholieke Universiteit Leuven) | 27 | 1.3K | 61 | 100 |
| 279 | Genetics (North Carolina State University) | 22 | 1.3K | 30 | 30 |
| 280 | Ecology (Umeå University) | 28 | 1.3K | 47 | 67 |
| 280 | Biology (University of Crete) | 12 | 1.3K | 76 | 34 |
| 282 | Ecology (University of California, San Diego) | 18 | 1.3K | 15 | 13 |
| 283 | Ecology (Stony Brook University) | 20 | 1.2K | 31 | 56 |
| 284 | Zoology (Occidental College) | 11 | 1.2K | 8 | 5 |
| 285 | Biotechnology (University of Helsinki) | 23 | 1.2K | 76 | 86 |
| 286 | Evolutionary Biology (Ludwig-Maximilians-Universität München) | 32 | 1.2K | 9 | 9 |
| 287 | Zoology (University of Florida) | 13 | 1.2K | 58 | 39 |
| 288 | Biology (University of Florida) | 28 | 1.2K | 74 | 86 |
| 289 | Marine Biology (University of California, San Diego) | 18 | 1.2K | 37 | 34 |
| 290 | Biology (University of Leicester) | 16 | 1.2K | 40 | 33 |
| 291 | Zoology (University of Greifswald) | 7 | 1.2K | 13 | 4 |
| 292 | Evolutionary Biology (University of Oregon) | 7 | 1.2K | 12 | 14 |
| 293 | Genetics (Rutgers University) | 15 | 1.2K | 32 | 37 |
| 294 | Ecology (University of New South Wales) | 26 | 1.2K | 25 | 31 |
| 294 | Environmental Science (Umeå University) | 25 | 1.2K | 48 | 61 |
| 296 | Zoology (University of New Hampshire) | 8 | 1.2K | 34 | 16 |
| 297 | Molecular Biology (James Cook University) | 12 | 1.2K | 9 | 5 |
| 298 | Genetics (University of Oulu) | 39 | 1.2K | 8 | 16 |
| 299 | Biology (University of Barcelona) | 19 | 1.2K | 75 | 58 |
| 300 | Biology (University College London) | 13 | 1.2K | 72 | 69 |
| 301 | Plant Pathology (University of Minnesota) | 8 | 1.2K | 64 | 33 |
| 302 | Zoology (University of Alberta) | 3 | 1.2K | 112 | 15 |
| 303 | Evolutionary Biology (University of Chicago) | 19 | 1.2K | 19 | 20 |
| 304 | Biology (University of Kentucky) | 26 | 1.2K | 28 | 32 |
| 304 | Genomics (University of Georgia) | 5 | 1.2K | 52 | 15 |
| 306 | Biology (Yale University) | 6 | 1.2K | 186 | 94 |
| 307 | Ecology (Complutense University of Madrid) | 10 | 1.1K | 22 | 6 |
| 308 | Evolutionary Biology (University of Helsinki) | 40 | 1.1K | 10 | 12 |
| 309 | Zoology (University of Oslo) | 13 | 1.1K | 22 | 10 |
| 310 | Ecology (Tulane University of Louisiana) | 23 | 1.1K | 18 | 18 |
| 310 | Evolutionary Biology (Tulane University of Louisiana) | 23 | 1.1K | 16 | 16 |
| 312 | Ecology (University of Wisconsin–Madison) | 12 | 1.1K | 83 | 48 |
| 313 | Biology (Cornell University) | 22 | 1.1K | 108 | 119 |
| 313 | Marine Biology (University of Groningen) | 11 | 1.1K | 14 | 10 |
| 315 | Zoology (Lund University) | 2 | 1.1K | 80 | 10 |
| 316 | Biology (Laval University) | 23 | 1.1K | 11 | 15 |
| 317 | Biology (University of California, San Diego) | 11 | 1.1K | 121 | 115 |
| 318 | Botany (University of Florida) | 5 | 1.1K | 58 | 27 |
| 319 | Ecology (Goethe University Frankfurt) | 22 | 1.1K | 31 | 26 |
| 320 | History (Norwegian University of Science and Technology) | 22 | 1.1K | 12 | 17 |
| 321 | Oceanography (University of Southampton) | 16 | 1.1K | 96 | 74 |
| 322 | Biology (University of Miami) | 26 | 1.1K | 34 | 40 |
| 323 | Biochemistry (James Cook University) | 7 | 1.1K | 21 | 7 |
| 324 | Ecology (University of Southampton) | 7 | 1.1K | 15 | 5 |
| 325 | Environmental Science (University of California, Santa Barbara) | 6 | 1.1K | 99 | 67 |
| 326 | Biology (Concordia University) | 20 | 1.1K | 40 | 27 |
| 327 | Molecular Biology (Cornell University) | 16 | 1.1K | 69 | 80 |
| 328 | Ecology (University of Oulu) | 36 | 1.1K | 8 | 13 |
| 329 | Genomics (University of Hull) | 5 | 1.1K | 2 | 1 |
| 330 | Ecology (Wageningen University & Research) | 19 | 1.1K | 92 | 78 |
| 331 | Evolutionary Biology (Université libre de Bruxelles) | 28 | 1.0K | 12 | 10 |
| 332 | Biology (Queen's University Belfast) | 14 | 1.0K | 68 | 50 |
| 333 | Biotechnology (University of New South Wales) | 17 | 1.0K | 74 | 72 |
| 334 | Earth Science (University of Hawaiʻi at Mānoa) | 13 | 1.0K | 37 | 32 |
| 335 | Computer Engineering (Drexel University) | 1 | 1.0K | 218 | 22 |
| 336 | Biology (University of Mississippi) | 11 | 1.0K | 45 | 14 |
| 337 | Biology (University of North Carolina at Chapel Hill) | 18 | 1.0K | 136 | 163 |
| 338 | Biology (University of Central Florida) | 20 | 1.0K | 17 | 25 |
| 339 | Biology (Columbia University) | 21 | 1.0K | 42 | 69 |
| 340 | Biology (University of Bergen) | 9 | 1.0K | 101 | 51 |
| 340 | Microbiology (University of Victoria) | 16 | 1.0K | 39 | 40 |
| 340 | Biochemistry (University of Victoria) | 16 | 1.0K | 39 | 44 |
| 343 | Microbiology (University of California, Riverside) | 7 | 1.0K | 53 | 38 |
| 343 | Plant Pathology (University of California, Riverside) | 7 | 1.0K | 70 | 48 |
| 345 | Environmental Science (University of Adelaide) | 11 | 1.0K | 5 | 9 |
| 346 | Education (Ocean University of China) | 17 | 1.0K | 169 | 103 |
| 347 | Ecology (University of Leeds) | 3 | 997 | 14 | 9 |
| 347 | Biochemistry (Queen's University Belfast) | 13 | 997 | 33 | 25 |
| 349 | Zoology (Complutense University of Madrid) | 2 | 994 | 37 | 5 |
| 350 | Anthropology (Duke University) | 11 | 990 | 42 | 27 |
| 351 | Zoology (University of Bern) | 9 | 981 | 13 | 12 |
| 352 | Evolutionary Biology (University of Oslo) | 14 | 980 | 9 | 9 |
| 353 | Population Genetics (Stockholm University) | 16 | 969 | 6 | 14 |
| 354 | Environmental Science (Sun Yat-sen University) | 6 | 967 | 131 | 58 |
| 355 | Botany (University of Tartu) | 10 | 957 | 32 | 26 |
| 355 | Genomics (Laval University) | 11 | 957 | 20 | 13 |
| 357 | Earth Science (University of Southampton) | 13 | 950 | 82 | 54 |
| 357 | Microbial Ecology (Michigan State University) | 10 | 950 | 23 | 27 |
| 359 | Ecology (Columbia University) | 20 | 945 | 27 | 38 |
| 360 | Biology (University of North Dakota) | 11 | 940 | 27 | 18 |
| 360 | Dynamics (Norwegian University of Science and Technology) | 29 | 940 | 12 | 12 |
| 362 | Paleontology (University of Zurich) | 1 | 931 | 19 | 3 |
| 363 | History (University of Washington) | 16 | 929 | 17 | 18 |
| 364 | Genomics (Wageningen University & Research) | 18 | 928 | 38 | 54 |
| 365 | Ecology (Ghent University) | 28 | 926 | 47 | 53 |
| 365 | Botany (Osnabrück University) | 10 | 926 | 10 | 8 |
| 367 | Biology (University of Utah) | 13 | 923 | 111 | 113 |
| 368 | Ecology (University of Edinburgh) | 18 | 920 | 28 | 29 |
| 368 | Genomics (Forest Research) | 10 | 920 | 4 | 4 |
| 370 | Ecology (University of Münster) | 3 | 915 | 25 | 3 |
| 371 | Genetics (University of Pretoria) | 24 | 914 | 31 | 30 |
| 372 | Zoology (University of Tartu) | 13 | 911 | 29 | 17 |
| 373 | Ecology (University of Florida) | 13 | 907 | 81 | 40 |
| 373 | Pathology (Swedish University of Agricultural Sciences) | 16 | 907 | 31 | 33 |
| 375 | Neurobiology (Cornell University) | 17 | 904 | 64 | 71 |
| 376 | Biology (University of Liverpool) | 29 | 901 | 58 | 97 |
| 377 | Genetics (Swedish University of Agricultural Sciences) | 17 | 895 | 66 | 71 |
| 378 | Ecology (Flinders University) | 20 | 890 | 8 | 7 |
| 379 | Biology (Oregon State University) | 19 | 886 | 22 | 31 |
| 380 | Ecology (Deakin University) | 20 | 879 | 43 | 45 |
| 381 | Biology (University of Padua) | 30 | 876 | 58 | 82 |
| 382 | Biology (New York University) | 18 | 872 | 97 | 118 |
| 383 | Biology (Emory University) | 12 | 865 | 85 | 71 |
| 384 | Zoology (University of Oklahoma) | 11 | 864 | 64 | 43 |
| 385 | Biology (Zhejiang University) | 12 | 863 | 84 | 68 |
| 386 | Biology (Northwest A&F University) | 9 | 859 | 150 | 68 |
| 386 | Biology (East Carolina University) | 14 | 859 | 46 | 37 |
| 386 | Ecology (Norwegian University of Life Sciences) | 10 | 859 | 50 | 35 |
| 389 | Genetics (Trinity College, Dublin) | 8 | 858 | 74 | 64 |
| 390 | Plant Pathology (Cornell University) | 12 | 853 | 58 | 46 |
| 390 | Biology (University of Vermont) | 19 | 853 | 25 | 27 |
| 392 | Ecology (University of Tübingen) | 8 | 851 | 48 | 23 |
| 392 | Ecology (Johannes Gutenberg University Mainz) | 13 | 851 | 8 | 7 |
| 394 | Zoology (University of Toronto) | 8 | 844 | 122 | 77 |
| 395 | Biology (Aristotle University of Thessaloniki) | 12 | 841 | 100 | 61 |
| 396 | Conservation Biology (University of Minnesota) | 11 | 839 | 56 | 37 |
| 397 | Ecology (Massey University) | 24 | 837 | 25 | 29 |
| 398 | Ecology (Michigan State University) | 10 | 827 | 52 | 25 |
| 399 | Molecular Biology (University of Vienna) | 1 | 820 | 76 | 17 |
| 400 | Biology (New Mexico State University) | 13 | 818 | 39 | 28 |
| 400 | Evolutionary Biology (University of Western Australia) | 21 | 818 | 14 | 13 |
| 402 | Biology (Environment and Climate Change Canada) | 8 | 815 | 23 | 13 |
| 402 | Genomics (University of Helsinki) | 3 | 815 | 47 | 15 |
| 404 | Evolutionary Biology (University of Glasgow) | 9 | 813 | 38 | 45 |
| 405 | Biology (University of Neuchâtel) | 28 | 812 | 19 | 43 |
| 406 | Biology (University of Porto) | 10 | 808 | 81 | 40 |
| 407 | Microbiology (Katholieke Universiteit Leuven) | 13 | 803 | 124 | 106 |
| 407 | Agricultural Science (University of Kentucky) | 7 | 803 | 20 | 9 |
| 409 | Biology (University of Ferrara) | 7 | 800 | 42 | 21 |
| 410 | Biology (Sciences Po) | 2 | 791 | 20 | 11 |
| 411 | Environmental Science (King Abdullah University of Science and Technology) | 18 | 788 | 45 | 49 |
| 412 | Biology (Washington State University) | 10 | 785 | 58 | 58 |
| 413 | Mathematics (University of Montana) | 3 | 781 | 7 | 2 |
| 414 | Zoology (University of Graz) | 16 | 779 | 13 | 10 |
| 414 | Agricultural Biotechnology (University of Pretoria) | 16 | 779 | 39 | 33 |
| 416 | Biology (University of Bath) | 16 | 778 | 67 | 88 |
| 417 | Ecology (University of Otago) | 11 | 777 | 15 | 12 |
| 417 | Biology (University of Puerto Rico) | 18 | 777 | 29 | 34 |
| 419 | Ecology (Friedrich Schiller University Jena) | 15 | 775 | 28 | 25 |
| 419 | Biology (University of Hawaiʻi at Mānoa) | 12 | 775 | 24 | 24 |
| 419 | Conservation Biology (Colorado State University) | 16 | 775 | 42 | 45 |
| 422 | Education (Sichuan University) | 27 | 774 | 178 | 182 |
| 423 | Zoology (Jagiellonian University) | 11 | 772 | 78 | 31 |
| 424 | Biology (University of Oldenburg) | 18 | 768 | 62 | 72 |
| 425 | Biology (University of Massachusetts Boston) | 13 | 766 | 24 | 25 |
| 426 | Biochemistry (University of Bath) | 15 | 765 | 69 | 92 |
| 427 | Ecological Engineering (Beijing Normal University) | 12 | 761 | 30 | 13 |
| 428 | Biology (University of Maine) | 15 | 760 | 33 | 24 |
| 428 | Ecology (University of Maine) | 15 | 760 | 44 | 33 |
| 428 | Molecular Biology (Babeș-Bolyai University) | 2 | 760 | 71 | 6 |
| 431 | Geography (University of Southampton) | 4 | 757 | 177 | 75 |
| 432 | Biogeography (Trier University) | 23 | 756 | 9 | 17 |
| 433 | Ecology (Radboud University) | 9 | 750 | 81 | 62 |
| 434 | Zoology (University of Innsbruck) | 14 | 749 | 20 | 14 |
| 434 | Biology (Vancouver Island University) | 3 | 749 | 8 | 4 |
| 436 | Biology (Rutgers University) | 11 | 748 | 205 | 158 |
| 436 | Environmental Science (University of Milan) | 10 | 748 | 44 | 20 |
| 438 | Ecology (Charles University) | 20 | 746 | 33 | 43 |
| 438 | Zoology (University of Wisconsin–Madison) | 6 | 746 | 137 | 89 |
| 438 | Conservation Biology (University of Lausanne) | 12 | 746 | 2 | 8 |
| 441 | Ecology (University of Lisbon) | 28 | 743 | 26 | 31 |
| 442 | Ecology (Victoria University of Wellington) | 7 | 740 | 12 | 3 |
| 443 | Genomics (Stellenbosch University) | 14 | 735 | 16 | 13 |
| 444 | Biology (Vassar College) | 3 | 725 | 68 | 9 |
| 444 | Environmental Engineering (École polytechnique) | 11 | 725 | 62 | 57 |
| 444 | Animal Science (Estonian University of Life Sciences) | 9 | 725 | 4 | 3 |
| 447 | Zoology (University of Gothenburg) | 12 | 717 | 56 | 55 |
| 448 | Immunology (Katholieke Universiteit Leuven) | 10 | 716 | 157 | 111 |
| 448 | Biology (University of North Carolina) | 4 | 716 | 108 | 37 |
| 450 | Marine Ecology (Aarhus University) | 5 | 714 | 32 | 17 |
| 451 | Genetics (University of California, Davis) | 17 | 711 | 40 | 46 |
| 452 | Marine Biology (University of Tromsø – The Arctic University of Norway) | 10 | 710 | 66 | 40 |
| 453 | Botany (University of Innsbruck) | 15 | 709 | 21 | 23 |
| 454 | Ecology (Stockholm University) | 24 | 708 | 37 | 69 |
| 455 | Plant Pathology (Kansas State University) | 5 | 704 | 68 | 46 |
| 456 | Evolutionary Biology (Michigan State University) | 6 | 703 | 32 | 15 |
| 457 | Ecology (Hebrew University of Jerusalem) | 16 | 701 | 29 | 40 |
| 458 | Biology (North Carolina State University) | 17 | 700 | 40 | 48 |
| 459 | Molecular Biology (University of Oregon) | 8 | 699 | 67 | 59 |
| 460 | Architecture (École polytechnique) | 11 | 690 | 61 | 52 |
| 461 | Biology (University of Missouri–St. Louis) | 16 | 688 | 38 | 57 |
| 462 | Zoology (Kyoto University) | 19 | 685 | 35 | 35 |
| 463 | Plant Pathology (ETH Zürich) | 12 | 683 | 17 | 21 |
| 464 | Genetics (Harvard University) | 10 | 679 | 245 | 221 |
| 464 | Biology (Columbus State University) | 5 | 679 | 5 | 3 |
| 464 | Genetics (Forest Research) | 15 | 679 | 4 | 9 |
| 467 | Biotechnology (BIOTEC) | 10 | 675 | 128 | 81 |
| 468 | Genetics (Iowa State University) | 13 | 673 | 62 | 56 |
| 469 | Genetics (University of Hull) | 7 | 672 | 17 | 5 |
| 469 | Ecology (University of Hull) | 7 | 672 | 7 | 8 |
| 471 | Biology (University of Iceland) | 10 | 671 | 27 | 25 |
| 472 | Biology (University of West Alabama) | 14 | 669 | 18 | 23 |
| 473 | Information Systems (École polytechnique) | 9 | 664 | 4 | 7 |
| 474 | Evolutionary Genetics (Pennsylvania State University) | 4 | 663 | 33 | 25 |
| 475 | Ecology (University at Buffalo) | 4 | 660 | 7 | 2 |
| 475 | Genomics (Katholieke Universiteit Leuven) | 19 | 660 | 20 | 21 |
| 477 | Biology (Harvard University) | 14 | 655 | 338 | 381 |
| 477 | Biology (University of Massachusetts Amherst) | 16 | 655 | 74 | 83 |
| 479 | Environmental Science (University College Dublin) | 19 | 653 | 33 | 48 |
| 480 | Ecology (Louisiana State University) | 2 | 652 | 98 | 29 |
| 480 | Evolutionary Biology (Louisiana State University) | 2 | 652 | 8 | 4 |
| 480 | Molecular Genetics (University of California, San Diego) | 4 | 652 | 86 | 54 |
| 483 | Geology (University at Buffalo) | 4 | 650 | 34 | 19 |
| 484 | Zoology (Tel Aviv University) | 16 | 649 | 96 | 90 |
| 484 | Marine Biology (James Cook University) | 9 | 649 | 28 | 30 |
| 484 | Biology (University of Alaska Southeast) | 6 | 649 | 6 | 12 |
| 487 | Biology (University of Zurich) | 9 | 646 | 86 | 101 |
| 487 | Statistics (University of California, Berkeley) | 10 | 646 | 63 | 71 |
| 489 | Zoology (University of Helsinki) | 9 | 645 | 47 | 29 |
| 490 | Evolutionary Biology (Université de Montréal) | 1 | 644 | 20 | 6 |
| 490 | Ecology (Université de Montréal) | 1 | 644 | 6 | 3 |
| 492 | Evolutionary Biology (University of Münster) | 1 | 641 | 10 | 2 |
| 492 | Biology (Harvey Mudd College) | 5 | 641 | 26 | 16 |
| 494 | Bioengineering (Central South University) | 3 | 638 | 171 | 91 |
| 495 | Biology (Leipzig University) | 12 | 635 | 48 | 49 |
| 496 | Zoology (Southern Illinois University Carbondale) | 7 | 633 | 81 | 42 |
| 497 | Biology (University of Duisburg-Essen) | 10 | 631 | 31 | 31 |
| 497 | Genetic Engineering (BIOTEC) | 9 | 631 | 129 | 78 |
| 499 | Ecology (Sorbonne University) | 15 | 629 | 18 | 17 |
| 499 | Ecology (Sorbonne Universités) | 7 | 629 | 11 | 7 |
| 501 | Pharmacy (James Cook University) | 3 | 628 | 49 | 20 |
| 502 | Ecology (University of Auckland) | 11 | 626 | 9 | 10 |
| 503 | Population Genetics (University of Cambridge) | 2 | 625 | 9 | 5 |
| 504 | Zoology (National Taiwan University) | 9 | 619 | 34 | 24 |
| 504 | Zoology (Masaryk University) | 23 | 619 | 44 | 52 |
| 504 | Oceanography (National Oceanography Centre) | 6 | 619 | 109 | 70 |
| 507 | Environmental Engineering (Dalian University of Technology) | 3 | 618 | 148 | 67 |
| 507 | Education (Dalian University of Technology) | 3 | 618 | 405 | 136 |
| 507 | Environmental Science (Dalian University of Technology) | 3 | 618 | 141 | 72 |
| 507 | Ecology (Dalian University of Technology) | 3 | 618 | 148 | 67 |
| 511 | Microbiology (Hebrew University of Jerusalem) | 5 | 616 | 141 | 76 |
| 511 | Plant Pathology (Hebrew University of Jerusalem) | 5 | 616 | 58 | 41 |
| 513 | Genomics (University of Edinburgh) | 13 | 611 | 26 | 27 |
| 514 | Biology (Utrecht University) | 7 | 610 | 207 | 168 |
| 514 | Biology (Universiti Malaysia Sabah) | 7 | 610 | 37 | 24 |
| 516 | Environmental Science (University of Maryland Center for Environmental Science) | 15 | 609 | 86 | 114 |
| 517 | Cell Biology (University of Connecticut) | 9 | 608 | 98 | 98 |
| 517 | Biology (Saint Mary's University) | 8 | 608 | 35 | 19 |
| 519 | Zoology (University of Potsdam) | 16 | 604 | 9 | 9 |
| 519 | Evolutionary Biology (University of Potsdam) | 16 | 604 | 10 | 9 |
| 521 | Microbiology (Duke University) | 7 | 603 | 135 | 111 |
| 522 | Ecology (University of Aberdeen) | 9 | 600 | 6 | 6 |
| 522 | Molecular Genetics (University of Aberdeen) | 9 | 600 | 5 | 3 |
| 524 | Biology (Trent University) | 25 | 599 | 23 | 47 |
| 525 | Biology (University of Innsbruck) | 2 | 592 | 57 | 18 |
| 526 | Biology (University of Bern) | 10 | 589 | 22 | 20 |
| 526 | Dynamics (ETH Zürich) | 9 | 589 | 56 | 62 |
| 528 | Botany (University of Georgia) | 10 | 586 | 27 | 37 |
| 529 | Molecular Biology (Marine Biological Laboratory) | 4 | 585 | 37 | 25 |
| 530 | Microbiology (University of Gothenburg) | 2 | 582 | 176 | 45 |
| 530 | Biology (University of Saskatchewan) | 10 | 582 | 106 | 82 |
| 530 | Plant Pathology (University of Nebraska–Lincoln) | 5 | 582 | 53 | 36 |
| 533 | Biochemistry (Uppsala University) | 15 | 579 | 100 | 124 |
| 533 | Microbiology (Uppsala University) | 15 | 579 | 68 | 87 |
| 533 | Biology (Swedish University of Agricultural Sciences) | 12 | 579 | 52 | 51 |
| 533 | Zoology (University of Hawaiʻi at Mānoa) | 8 | 579 | 22 | 12 |
| 537 | Biotechnology (Ankara University) | 2 | 577 | 107 | 15 |
| 538 | Microbiology (University of Copenhagen) | 8 | 576 | 139 | 111 |
| 539 | Ecology (University of Maryland) | 6 | 575 | 19 | 9 |
| 540 | Education (Xiamen University) | 7 | 574 | 212 | 80 |
| 541 | Biology (University of Salzburg) | 7 | 573 | 44 | 19 |
| 542 | Microbiology (Colorado State University) | 14 | 571 | 83 | 101 |
| 543 | Biology (Middle Tennessee State University) | 8 | 570 | 34 | 13 |
| 544 | Zoology (University of Hawaii) | 6 | 568 | 39 | 20 |
| 545 | Microbial Ecology (Lund University) | 5 | 567 | 47 | 26 |
| 545 | Pharmacy (Osaka Ohtani University) | 7 | 567 | 17 | 8 |
| 547 | Pathology (University of California, Davis) | 8 | 564 | 190 | 144 |
| 548 | Biology (Georgetown University) | 18 | 562 | 49 | 83 |
| 548 | Zoology (King Saud University) | 5 | 562 | 258 | 92 |
| 550 | Pathology (Colorado State University) | 13 | 560 | 88 | 94 |
| 550 | Immunology (Colorado State University) | 13 | 560 | 75 | 81 |
| 552 | Ecology (Curtin University) | 12 | 559 | 5 | 4 |
| 553 | Conservation Biology (Texas State University) | 7 | 558 | 5 | 4 |
| 554 | Biotechnology (University of Florida) | 5 | 555 | 48 | 33 |
| 554 | Biology (Arizona State University) | 7 | 555 | 79 | 80 |
| 554 | Mathematics (University of Porto) | 2 | 555 | 50 | 5 |
| 557 | Public Health (Harvard University) | 9 | 553 | 924 | 756 |
| 558 | Genomics (University of California, Berkeley) | 13 | 551 | 30 | 48 |
| 559 | Botany (Masaryk University) | 22 | 547 | 43 | 58 |
| 560 | Biology (Central Michigan University) | 4 | 546 | 85 | 19 |
| 560 | Genetics (University of Florida) | 7 | 546 | 139 | 105 |
| 562 | Mathematics (University of the Virgin Islands) | 1 | 545 | 24 | 2 |
| 563 | Biology (University of Iowa) | 9 | 543 | 93 | 94 |
| 564 | Genetics (University of Barcelona) | 10 | 541 | 40 | 43 |
| 564 | Statistics (University of California, Santa Cruz) | 6 | 541 | 31 | 18 |
| 564 | Molecular Biology (University of California, Berkeley) | 4 | 541 | 70 | 46 |
| 567 | Oceanography (University of Hawaiʻi at Mānoa) | 7 | 540 | 55 | 46 |
| 568 | Earth Science (Hokkaido University) | 14 | 537 | 101 | 104 |
| 569 | Biology (City College of New York) | 12 | 536 | 30 | 45 |
| 570 | Biology (University of Haifa) | 11 | 534 | 58 | 47 |
| 570 | Genetics (University of Zagreb) | 3 | 534 | 22 | 6 |
| 572 | Ecology (University of Colorado) | 5 | 533 | 31 | 23 |
| 572 | Evolutionary Biology (University of Colorado) | 5 | 533 | 31 | 23 |
| 574 | Ecology (Okinawa Institute of Science and Technology) | 4 | 532 | 10 | 5 |
| 575 | Geography (University of Marburg) | 4 | 530 | 65 | 31 |
| 576 | Immunology (University of Gothenburg) | 1 | 529 | 186 | 44 |
| 577 | Biology (Bucknell University) | 9 | 526 | 24 | 17 |
| 578 | Genetics (University of Copenhagen) | 4 | 522 | 117 | 59 |
| 579 | Ecology (Ulm University) | 15 | 521 | 24 | 40 |
| 579 | Ecology (Utah State University) | 20 | 521 | 46 | 77 |
| 579 | Evolutionary Biology (University of Copenhagen) | 7 | 521 | 10 | 16 |
| 579 | Genomics (Texas A&M University–Corpus Christi) | 6 | 521 | 4 | 3 |
| 583 | Biology (University of Tromsø – The Arctic University of Norway) | 9 | 519 | 132 | 129 |
| 584 | Landscape Ecology (ETH Zürich) | 5 | 518 | 12 | 11 |
| 585 | Botany (University of Western Australia) | 4 | 511 | 35 | 17 |
| 585 | Mathematics (University of Idaho) | 5 | 511 | 31 | 17 |
| 587 | Biology (Portland State University) | 13 | 510 | 30 | 35 |
| 588 | Biology (University of Colorado Denver) | 8 | 509 | 37 | 29 |
| 589 | Mathematics (University of Vienna) | 3 | 507 | 143 | 53 |
| 590 | Biology (Virginia Tech) | 5 | 506 | 165 | 87 |
| 590 | Biogeography (University of Vienna) | 7 | 506 | 12 | 14 |
| 590 | Biology (University of Ottawa) | 9 | 506 | 178 | 144 |
| 593 | Molecular Genetics (University of Bologna) | 2 | 503 | 29 | 8 |
| 593 | Ecology (Sciences Po) | 1 | 503 | 46 | 11 |
| 595 | Zoology (University of Cologne) | 11 | 502 | 36 | 27 |
| 595 | Biology (University of Marburg) | 8 | 502 | 72 | 72 |
| 597 | Biology (University of Geneva) | 6 | 500 | 83 | 75 |
| 598 | Ecology (Austral University of Chile) | 11 | 496 | 8 | 6 |
| 599 | Ecology (University of Sheffield) | 14 | 495 | 11 | 13 |
| 600 | Biology (Adam Mickiewicz University in Poznań) | 16 | 491 | 67 | 66 |
| 600 | Nuclear Energy (University of São Paulo) | 5 | 491 | 113 | 39 |
| 600 | Forest Ecology (Forest Research) | 15 | 491 | 4 | 11 |
| 603 | Microbial Ecology (Katholieke Universiteit Leuven) | 5 | 490 | 16 | 15 |
| 604 | Zoology (University of Cape Town) | 7 | 489 | 74 | 60 |
| 605 | Molecular Biology (Zhejiang University) | 10 | 488 | 62 | 59 |
| 606 | Immunology (Montana State University) | 5 | 487 | 44 | 29 |
| 606 | Microbiology (Montana State University) | 5 | 487 | 86 | 54 |
| 608 | Biology (Temple University) | 11 | 486 | 63 | 66 |
| 609 | Ecology (Beijing Normal University) | 6 | 485 | 158 | 102 |
| 609 | Botany (University of Hawaiʻi at Mānoa) | 9 | 485 | 40 | 44 |
| 609 | Global Health (University of Notre Dame) | 9 | 485 | 46 | 47 |
| 612 | Biology (University of Freiburg) | 11 | 484 | 131 | 141 |
| 612 | Evolutionary Biology (Austral University of Chile) | 9 | 484 | 5 | 3 |
| 614 | Ecology (Duke University) | 5 | 483 | 30 | 27 |
| 614 | Conservation Biology (University of Wollongong) | 6 | 483 | 36 | 22 |
| 616 | Zoology (University of Washington) | 9 | 480 | 66 | 83 |
| 616 | Ecology (University of Giessen) | 10 | 480 | 46 | 33 |
| 616 | Biology (University of Nottingham) | 10 | 480 | 60 | 71 |
| 619 | Plant Ecology (Katholieke Universiteit Leuven) | 6 | 479 | 9 | 12 |
| 620 | Biology (Goodwin University) | 6 | 477 | 44 | 35 |
| 620 | Earth Science (National Oceanography Centre) | 4 | 477 | 61 | 22 |
| 622 | Public Health (James Cook University) | 6 | 476 | 158 | 96 |
| 623 | Horticulture (Akdeniz University) | 1 | 475 | 25 | 7 |
| 623 | Horticulture (Ankara University) | 1 | 475 | 40 | 6 |
| 625 | Molecular Biology (University of São Paulo) | 4 | 474 | 84 | 36 |
| 626 | Biology (Memorial University of Newfoundland) | 9 | 473 | 92 | 78 |
| 627 | Developmental Biology (University of California, Los Angeles) | 5 | 472 | 125 | 97 |
| 628 | Biochemistry (University of California, Berkeley) | 3 | 471 | 102 | 62 |
| 629 | Genomics (University of Central Florida) | 6 | 469 | 8 | 2 |
| 630 | Ecology (Griffith University) | 7 | 468 | 5 | 5 |
| 631 | Biology (Osnabrück University) | 5 | 467 | 64 | 46 |
| 631 | Ecology (University of Miami) | 8 | 467 | 26 | 16 |
| 633 | Ecology (University of Freiburg) | 12 | 466 | 40 | 41 |
| 633 | Plant Pathology (University of Kentucky) | 2 | 466 | 81 | 38 |
| 635 | Psychology (Liverpool John Moores University) | 10 | 464 | 49 | 51 |
| 636 | Genetics (Northern Arizona University) | 6 | 463 | 3 | 3 |
| 637 | History (University of Kansas) | 10 | 462 | 60 | 57 |
| 638 | Biology (University of South Bohemia in České Budějovice) | 7 | 461 | 118 | 90 |
| 638 | Biology (Willamette University) | 5 | 461 | 9 | 7 |
| 640 | Biology (University of North Carolina at Charlotte) | 3 | 460 | 88 | 39 |
| 640 | Evolutionary Biology (Katholieke Universiteit Leuven) | 8 | 460 | 13 | 20 |
| 642 | Biochemistry (University of Cambridge) | 3 | 459 | 309 | 199 |
| 643 | Biology (West Virginia University) | 8 | 457 | 59 | 47 |
| 644 | Microbiology (Michigan State University) | 4 | 454 | 196 | 122 |
| 645 | Statistics (Oklahoma State University) | 4 | 451 | 60 | 24 |
| 645 | Education (Technical University of Denmark) | 1 | 451 | 23 | 3 |
| 645 | Biochemical Engineering (Technical University of Denmark) | 1 | 451 | 242 | 95 |
| 648 | Botany (University of Wisconsin–Madison) | 9 | 449 | 72 | 84 |
| 649 | Genetics (Stanford University) | 9 | 448 | 161 | 169 |
| 649 | Zoology (University of Johannesburg) | 13 | 448 | 26 | 25 |
| 651 | Biology (Laurentian University) | 10 | 446 | 42 | 26 |
| 652 | Applied Mathematics (University of California, Santa Cruz) | 4 | 445 | 39 | 18 |
| 653 | Zoology (Trinity College, Dublin) | 6 | 442 | 67 | 34 |
| 653 | Biology (Stellenbosch University) | 12 | 442 | 72 | 95 |
| 655 | History (University of Crete) | 8 | 441 | 23 | 20 |
| 656 | History (City University of New York) | 3 | 440 | 41 | 15 |
| 657 | Archaeology (University of Oxford) | 6 | 438 | 67 | 52 |
| 657 | Zoology (Swansea University) | 5 | 438 | 44 | 16 |
| 659 | Evolutionary Biology (Sorbonne Universités) | 6 | 436 | 8 | 4 |
| 660 | Zoology (Lomonosov Moscow State University) | 3 | 435 | 54 | 18 |
| 660 | Ecology (University of Wisconsin–Milwaukee) | 11 | 435 | 11 | 10 |
| 662 | Pathology (Vanderbilt University) | 3 | 434 | 364 | 210 |
| 662 | Immunology (Vanderbilt University) | 3 | 434 | 329 | 177 |
| 662 | Microbiology (Vanderbilt University) | 3 | 434 | 307 | 169 |
| 662 | Ecology (University of Hamburg) | 6 | 434 | 30 | 25 |
| 666 | Oceanography (University of the Azores) | 6 | 433 | 23 | 15 |
| 667 | Genetics (University of Canberra) | 2 | 432 | 6 | 2 |
| 667 | Cell Biology (University of California, Berkeley) | 5 | 432 | 214 | 208 |
| 669 | Microbiology (University of Pretoria) | 12 | 431 | 65 | 66 |
| 669 | Biology (Loyola Marymount University) | 3 | 431 | 20 | 6 |
| 669 | Immunology (University of California, Davis) | 8 | 431 | 144 | 138 |
| 672 | Ecology (ETH Zürich) | 11 | 430 | 27 | 60 |
| 672 | Biology (University of Minho) | 7 | 430 | 91 | 69 |
| 674 | Ecology (University of Hong Kong) | 13 | 429 | 38 | 55 |
| 674 | Genetics (University of Sydney) | 12 | 429 | 35 | 54 |
| 676 | Genetics (University of Toledo) | 7 | 428 | 2 | 4 |
| 677 | Ecology (University of Glasgow) | 6 | 425 | 34 | 30 |
| 677 | Evolutionary Biology (Sorbonne University) | 7 | 425 | 10 | 5 |
| 677 | Ecological Genetics (ETH Zürich) | 8 | 425 | 9 | 9 |
| 680 | Landscape Architecture (University of Maryland) | 4 | 424 | 66 | 47 |
| 681 | Genetics (University of Giessen) | 1 | 421 | 90 | 29 |
| 682 | Computational Biology (University of Southern California) | 12 | 420 | 45 | 75 |
| 683 | Biology (University of California, Santa Barbara) | 1 | 419 | 92 | 39 |
| 684 | Microbiology (University of California, Davis) | 8 | 418 | 120 | 122 |
| 685 | Zoology (University of Göttingen) | 5 | 417 | 86 | 56 |
| 686 | Statistics (University of Connecticut) | 1 | 416 | 216 | 37 |
| 687 | Conservation Biology (North Carolina State University) | 7 | 415 | 20 | 15 |
| 688 | Geography (University of Copenhagen) | 2 | 414 | 157 | 81 |
| 688 | Biology (University of Waterloo) | 3 | 414 | 314 | 156 |
| 690 | Biochemistry (Simon Fraser University) | 6 | 410 | 78 | 62 |
| 690 | Molecular Biology (Simon Fraser University) | 6 | 410 | 74 | 62 |
| 690 | Biology (Linköping University) | 13 | 410 | 109 | 138 |
| 690 | Botany (University of California, Riverside) | 10 | 410 | 76 | 104 |
| 694 | Biology (University of Regina) | 5 | 407 | 100 | 47 |
| 695 | Ecology (North Carolina State University) | 10 | 405 | 41 | 35 |
| 696 | History (University of Geneva) | 6 | 403 | 17 | 16 |
| 696 | Biochemistry (Wright State University) | 1 | 403 | 103 | 24 |
| 696 | Molecular Biology (Wright State University) | 1 | 403 | 92 | 22 |
| 699 | Marine Biology (Ghent University) | 10 | 402 | 35 | 35 |
| 700 | Microbiology (University of Tartu) | 4 | 401 | 43 | 24 |
| 700 | Pathology (Rutgers University) | 3 | 401 | 165 | 81 |
| 700 | Physiology (Nagoya University) | 10 | 401 | 23 | 21 |
| 700 | Forest Ecology (Nagoya University) | 10 | 401 | 9 | 5 |
| 700 | Evolutionary Biology (University of Konstanz) | 8 | 401 | 12 | 13 |
| 705 | Education (Kyoto University) | 6 | 400 | 127 | 59 |
| 705 | Ecology (Vilnius University) | 3 | 400 | 31 | 19 |
| 705 | Biochemistry (University of Otago) | 8 | 400 | 91 | 102 |
| 705 | Functional Genomics (Uppsala University) | 8 | 400 | 5 | 5 |
| 709 | Biology (University of Chicago) | 5 | 398 | 173 | 156 |
| 710 | Zoology (University of Geneva) | 4 | 397 | 43 | 39 |
| 711 | Genetics (Yale University) | 5 | 396 | 183 | 143 |
| 712 | Biology (University of Würzburg) | 10 | 394 | 72 | 88 |
| 712 | Environmental Science (University of Tokyo) | 10 | 394 | 53 | 48 |
| 712 | Ecology (University of Würzburg) | 10 | 394 | 43 | 46 |
| 715 | Ecology (University of Cologne) | 6 | 393 | 37 | 33 |
| 715 | Ecology (Tohoku University) | 14 | 393 | 18 | 27 |
| 715 | Environmental Science (Shimane University) | 2 | 393 | 143 | 37 |
| 715 | Genomics (Indiana University Bloomington) | 7 | 393 | 17 | 18 |
| 719 | Ecology (Eötvös Loránd University) | 8 | 392 | 52 | 48 |
| 719 | Zoology (Eötvös Loránd University) | 8 | 392 | 28 | 23 |
| 719 | Biology (University of Florence) | 8 | 392 | 91 | 63 |
| 722 | Cell Biology (University of Edinburgh) | 2 | 391 | 119 | 71 |
| 723 | Plant Pathology (Oklahoma State University) | 2 | 388 | 110 | 44 |
| 724 | Molecular Biology (London School of Hygiene & Tropical Medicine) | 5 | 387 | 34 | 36 |
| 725 | Biology (University of Southampton) | 3 | 385 | 71 | 25 |
| 725 | Ecology (University of São Paulo) | 6 | 385 | 61 | 44 |
| 725 | Mathematics (Saint Mary's University) | 1 | 385 | 61 | 3 |
| 728 | Microbiology (University of Bergen) | 3 | 384 | 103 | 56 |
| 728 | Biology (Vrije Universiteit Brussel) | 7 | 384 | 80 | 73 |
| 730 | History (University of Nevada, Las Vegas) | 7 | 383 | 10 | 11 |
| 731 | Landscape Ecology (University of Giessen) | 3 | 382 | 60 | 30 |
| 732 | Ecology (La Trobe University) | 15 | 381 | 26 | 34 |
| 732 | Agricultural Science (Tohoku University) | 13 | 381 | 101 | 115 |
| 734 | Ecology (University of Kent) | 11 | 379 | 36 | 41 |
| 735 | Genetics (Stockholm University) | 8 | 378 | 28 | 36 |
| 736 | Biology (Martin Luther University Halle-Wittenberg) | 11 | 377 | 67 | 81 |
| 736 | Environmental Science (Xiamen University) | 16 | 377 | 119 | 136 |
| 738 | Zoology (University of Zurich) | 8 | 376 | 25 | 40 |
| 738 | Evolutionary Biology (University of Hull) | 4 | 376 | 7 | 7 |
| 740 | Microbiology (ETH Zürich) | 3 | 375 | 93 | 76 |
| 740 | Statistics (University of Oxford) | 2 | 375 | 367 | 192 |
| 740 | Biology (University of Oradea) | 3 | 375 | 6 | 2 |
| 743 | Biology (Auburn University) | 7 | 373 | 12 | 13 |
| 743 | Biology (University of Windsor) | 6 | 373 | 40 | 19 |
| 743 | Ecology (City University of New York) | 5 | 373 | 8 | 6 |
| 746 | Biology (University of North Carolina at Greensboro) | 7 | 372 | 28 | 33 |
| 746 | Computer Science (Indiana University Bloomington) | 8 | 372 | 30 | 26 |
| 748 | Biology (Mahidol University) | 9 | 371 | 55 | 51 |
| 748 | Biology (Skidmore College) | 6 | 371 | 29 | 20 |
| 750 | Computer Science (University of Colorado Boulder) | 1 | 370 | 271 | 79 |
| 750 | Psychology (McMaster University) | 7 | 370 | 130 | 132 |
| 752 | Genetics (Lund University) | 3 | 368 | 52 | 27 |
| 752 | Biology (Syracuse University) | 13 | 368 | 46 | 91 |
| 752 | Population Genetics (Technical University of Munich) | 8 | 368 | 7 | 6 |
| 755 | Microbiology (University of Sydney) | 2 | 367 | 123 | 67 |
| 756 | Pathology (University of Münster) | 1 | 366 | 133 | 42 |
| 756 | Public Health (Johns Hopkins University) | 3 | 366 | 1,350 | 879 |
| 756 | Biology (University of Colorado Boulder) | 5 | 366 | 139 | 129 |
| 756 | Biology (Chinese University of Hong Kong) | 4 | 366 | 135 | 84 |
| 756 | Conservation Biology (University of California, Davis) | 10 | 366 | 51 | 72 |
| 761 | Zoology (University College London) | 3 | 364 | 59 | 38 |
| 761 | Statistics (University of Idaho) | 3 | 364 | 33 | 20 |
| 763 | Evolutionary Biology (University of Massachusetts Amherst) | 11 | 361 | 28 | 41 |
| 763 | Zoology (Estonian University of Life Sciences) | 2 | 361 | 31 | 13 |
| 763 | Botany (Estonian University of Life Sciences) | 2 | 361 | 32 | 12 |
| 763 | Ecology (Ben-Gurion University of the Negev) | 15 | 361 | 38 | 54 |
| 763 | Botany (University of Guelph) | 3 | 361 | 63 | 27 |
| 768 | Genetics (Aristotle University of Thessaloniki) | 5 | 359 | 78 | 42 |
| 769 | Evolutionary Biology (Tohoku University) | 13 | 358 | 11 | 12 |
| 769 | Biology (University of South Dakota) | 9 | 358 | 46 | 52 |
| 771 | History (University of Guelph) | 2 | 357 | 19 | 3 |
| 772 | Genomics (Ulm University) | 12 | 356 | 12 | 17 |
| 773 | Pathology (University of the Witwatersrand) | 2 | 355 | 302 | 100 |
| 773 | Population Genetics (University of Hohenheim) | 7 | 355 | 25 | 23 |
| 775 | Biology (Radboud University) | 3 | 354 | 125 | 75 |
| 776 | Molecular Biology (Brigham Young University) | 5 | 353 | 41 | 34 |
| 776 | Microbiology (Brigham Young University) | 5 | 353 | 45 | 44 |
| 778 | Computer Science (University of Cambridge) | 1 | 352 | 189 | 35 |
| 779 | Mathematics (Medical University of Vienna) | 1 | 351 | 2 | 2 |
| 780 | Archaeology (Norwegian University of Science and Technology) | 7 | 350 | 17 | 19 |
| 780 | Evolutionary Biology (University of Valencia) | 4 | 350 | 87 | 39 |
| 782 | Biology (Reed College) | 2 | 349 | 64 | 33 |
| 782 | Ecology (University of California, Riverside) | 10 | 349 | 24 | 21 |
| 784 | Biology (Tufts University) | 6 | 348 | 130 | 132 |
| 784 | Anthropology (University of Göttingen) | 5 | 348 | 84 | 65 |
| 784 | Environmental Science (University of Potsdam) | 6 | 348 | 66 | 55 |
| 784 | Marine Biology (Åbo Akademi University) | 3 | 348 | 59 | 29 |
| 788 | Plant Pathology (University of California, Davis) | 5 | 347 | 89 | 72 |
| 789 | Human Genetics (University of Oxford) | 6 | 346 | 200 | 194 |
| 789 | Pathology (Uppsala University) | 7 | 346 | 197 | 193 |
| 789 | Immunology (Uppsala University) | 7 | 346 | 165 | 145 |
| 792 | Genomics (Kansas State University) | 3 | 345 | 16 | 17 |
| 793 | Public Health (State University of New York at Albany) | 2 | 344 | 343 | 123 |
| 794 | Genomics (University of Nottingham) | 3 | 342 | 22 | 15 |
| 795 | Zoology (University of Vienna) | 6 | 341 | 53 | 47 |
| 795 | Biology (National Cheng Kung University) | 4 | 341 | 22 | 9 |
| 795 | Genetics (University of Leicester) | 12 | 341 | 58 | 92 |
| 795 | Biotechnology (Murdoch University) | 6 | 341 | 56 | 58 |
| 799 | Biology (University of Reading) | 2 | 339 | 17 | 9 |
| 799 | Plant Pathology (University of Pretoria) | 7 | 339 | 48 | 48 |
| 801 | Statistics (Pennsylvania State University) | 4 | 338 | 126 | 77 |
| 801 | Zoology (University of Western Australia) | 7 | 338 | 43 | 46 |
| 803 | Biology (University of Michigan) | 6 | 337 | 170 | 189 |
| 803 | Ecology (James Cook University) | 6 | 337 | 27 | 31 |
| 803 | Genetics (University of Groningen) | 8 | 337 | 101 | 125 |
| 803 | Systems Biology (Harvard University) | 5 | 337 | 162 | 161 |
| 807 | Geology (University of Copenhagen) | 1 | 336 | 154 | 55 |
| 807 | Ecology (University of Nevada, Reno) | 10 | 336 | 34 | 36 |
| 807 | Molecular Genetics (Michigan State University) | 3 | 336 | 173 | 121 |
| 810 | Biotechnology (Nanjing Normal University) | 10 | 335 | 89 | 80 |
| 811 | Environmental Engineering (Aalborg University) | 5 | 333 | 65 | 59 |
| 812 | Conservation Biology (University of Nevada, Reno) | 9 | 331 | 36 | 32 |
| 813 | Microbiology (Stockholm University) | 7 | 329 | 23 | 34 |
| 814 | Biology (University of São Paulo) | 4 | 328 | 209 | 84 |
| 815 | Biology (University of Bologna) | 5 | 327 | 93 | 79 |
| 815 | Computer Science (Aalto University) | 5 | 327 | 133 | 106 |
| 817 | Zoology (Aristotle University of Thessaloniki) | 4 | 326 | 91 | 46 |
| 818 | Molecular Biology (Aristotle University of Thessaloniki) | 4 | 325 | 56 | 31 |
| 818 | Forensic Science (Trent University) | 11 | 325 | 7 | 10 |
| 818 | Conservation Biology (National Oceanic and Atmospheric Administration) | 9 | 325 | 15 | 17 |
| 821 | Biology (University of Aveiro) | 8 | 323 | 134 | 133 |
| 821 | Biology (Newcastle University) | 7 | 323 | 70 | 107 |
| 821 | Biology (Coastal Carolina University) | 3 | 323 | 28 | 5 |
| 824 | Biotechnology (Graz University of Technology) | 4 | 322 | 89 | 76 |
| 825 | Ecology (Peking University) | 3 | 321 | 128 | 100 |
| 825 | Physiology (Radboud University) | 4 | 321 | 116 | 100 |
| 825 | Ecology (Pierre and Marie Curie University) | 3 | 321 | 14 | 9 |
| 825 | Evolutionary Biology (Pierre and Marie Curie University) | 3 | 321 | 9 | 4 |
| 829 | Biology (University of Tokyo) | 6 | 320 | 179 | 169 |
| 829 | Plant Pathology (University of Arkansas) | 3 | 320 | 66 | 27 |
| 829 | History (University of Oklahoma) | 4 | 320 | 42 | 31 |
| 829 | Molecular Genetics (Ohio State University) | 3 | 320 | 113 | 82 |
| 829 | Evolutionary Biology (University of Siena) | 3 | 320 | 65 | 44 |
| 829 | Environmental Science (University of New South Wales) | 4 | 320 | 45 | 30 |
| 835 | Biology (Amherst College) | 7 | 319 | 31 | 35 |
| 835 | Biology (University of Pretoria) | 2 | 319 | 93 | 38 |
| 837 | Biology (University of Houston) | 8 | 318 | 110 | 125 |
| 837 | Bioengineering (Dalian University of Technology) | 2 | 318 | 133 | 40 |
| 839 | Botany (Washington State University) | 3 | 317 | 38 | 35 |
| 839 | Biology (Kobe University) | 6 | 317 | 70 | 50 |
| 839 | Biotechnology (University of Bremen) | 4 | 317 | 15 | 19 |
| 839 | Molecular Genetics (University of Bremen) | 4 | 317 | 4 | 3 |
| 843 | Chemistry (University of Würzburg) | 7 | 316 | 47 | 71 |
| 843 | Ecology (University of Cambridge) | 5 | 316 | 32 | 42 |
| 845 | Genetics (University of the Free State) | 6 | 315 | 26 | 17 |
| 846 | Microbiology (University of Massachusetts Amherst) | 5 | 314 | 60 | 68 |
| 846 | Biology (Autonomous University of Barcelona) | 3 | 314 | 111 | 58 |
| 846 | Ecology (Autonomous University of Barcelona) | 3 | 314 | 105 | 56 |
| 849 | Ecology (National Taiwan University) | 7 | 313 | 29 | 24 |
| 849 | Biotechnology (National Fisheries Research and Development Institute) | 8 | 313 | 17 | 12 |
| 849 | Conservation Biology (Norwegian University of Science and Technology) | 5 | 313 | 13 | 13 |
| 852 | Zoology (Oklahoma State University) | 7 | 312 | 71 | 62 |
| 852 | Computational Biology (University of Lausanne) | 7 | 312 | 28 | 31 |
| 854 | Oceanography (University of Vienna) | 3 | 311 | 35 | 21 |
| 854 | Biology (Southern Illinois University Edwardsville) | 1 | 311 | 16 | 3 |
| 854 | Zoology (University of Konstanz) | 7 | 311 | 12 | 12 |
| 854 | Biology (Florida International University) | 2 | 311 | 87 | 39 |
| 854 | Dentistry (University of Aberdeen) | 2 | 311 | 149 | 59 |
| 859 | Evolutionary Biology (Adam Mickiewicz University in Poznań) | 10 | 309 | 7 | 6 |
| 859 | Ecology (University of Bayreuth) | 7 | 309 | 103 | 102 |
| 859 | Biotechnology (University of Milan-Bicocca) | 9 | 309 | 89 | 114 |
| 862 | Biology (Kangwon National University) | 1 | 308 | 80 | 23 |
| 862 | Biology (University of Nevada, Las Vegas) | 1 | 308 | 18 | 4 |
| 864 | Zoology (Johannes Gutenberg University Mainz) | 9 | 307 | 33 | 54 |
| 864 | Biology (University of Wisconsin–Madison) | 11 | 307 | 70 | 134 |
| 866 | Anatomy (University of Chicago) | 3 | 306 | 119 | 93 |
| 867 | Clinical Genetics (Leiden University) | 2 | 305 | 172 | 88 |
| 868 | Botany (Duke University) | 4 | 304 | 76 | 82 |
| 868 | Statistics (University of Auckland) | 3 | 304 | 158 | 84 |
| 870 | Biology (University of Hawaii) | 4 | 303 | 74 | 70 |
| 870 | Global Health (NOVA University Lisbon) | 3 | 303 | 96 | 37 |
| 872 | Genetics (Southern Cross University) | 5 | 302 | 26 | 18 |
| 872 | Evolutionary Biology (University of Sheffield) | 10 | 302 | 12 | 9 |
| 874 | Biology (Saint Louis University) | 7 | 301 | 53 | 48 |
| 875 | Biochemistry (University of Cologne) | 2 | 300 | 183 | 100 |
| 876 | Biology (Addis Ababa University) | 5 | 299 | 57 | 48 |
| 876 | Biology (University of North Carolina Wilmington) | 6 | 299 | 66 | 44 |
| 876 | Human Genetics (University of California, Los Angeles) | 8 | 299 | 131 | 174 |
| 876 | History (Hunter College) | 1 | 299 | 31 | 5 |
| 876 | Applied Mathematics (Icahn School of Medicine at Mount Sinai) | 1 | 299 | 10 | 2 |
| 881 | Biology (Sapienza University of Rome) | 12 | 298 | 132 | 186 |
| 881 | Ecology (University of Oklahoma) | 7 | 298 | 19 | 29 |
| 881 | Biology (University of Hawaiʻi at Hilo) | 5 | 298 | 19 | 12 |
| 881 | Biotechnology (Chinese Culture University) | 4 | 298 | 26 | 17 |
| 885 | Molecular Biology (University of Gothenburg) | 2 | 296 | 275 | 135 |
| 885 | Zoology (University of Basel) | 12 | 296 | 18 | 35 |
| 885 | Biology (University of Arizona) | 7 | 296 | 95 | 136 |
| 885 | Genetics (University of Montana) | 4 | 296 | 6 | 4 |
| 889 | Biology (Joseph Fourier University) | 1 | 295 | 23 | 17 |
| 889 | Molecular Genetics (University of Sheffield) | 10 | 295 | 4 | 5 |
| 889 | Genomics (University of Notre Dame) | 1 | 295 | 11 | 3 |
| 892 | Marine Biology (University of Algarve) | 1 | 294 | 1 | 1 |
| 892 | Ecology (University of Montana) | 6 | 294 | 11 | 16 |
| 892 | Marine Biology (University of Alaska Fairbanks) | 1 | 294 | 11 | 4 |
| 895 | Biochemistry (University of Houston) | 7 | 293 | 99 | 105 |
| 895 | Evolutionary Biology (City University of New York) | 4 | 293 | 8 | 8 |
| 897 | Biology (CUNY Graduate School and University Center) | 6 | 292 | 34 | 36 |
| 897 | Evolutionary Biology (National Taiwan University) | 6 | 292 | 37 | 24 |
| 899 | Physiology (Linköping University) | 7 | 291 | 36 | 33 |
| 899 | Genomics (Linköping University) | 7 | 291 | 10 | 11 |
| 901 | Environmental Science (Swedish University of Agricultural Sciences) | 8 | 290 | 6 | 22 |
| 901 | Freshwater Ecology (Griffith University) | 5 | 290 | 8 | 13 |
| 903 | Microbiology (University of Lausanne) | 3 | 288 | 131 | 89 |
| 903 | Ecology (Pontifical Catholic University of Chile) | 10 | 288 | 55 | 75 |
| 903 | Agroforestry (United Nations) | 5 | 288 | 28 | 36 |
| 906 | Oceanography (Marine Biological Laboratory) | 4 | 287 | 62 | 57 |
| 906 | Agronomy (Hunan Agricultural University) | 2 | 287 | 87 | 48 |
| 908 | Ecology (University of Marburg) | 5 | 286 | 41 | 29 |
| 909 | Evolutionary Biology (Bielefeld University) | 8 | 285 | 14 | 11 |
| 910 | Education (Virginia Commonwealth University) | 2 | 284 | 112 | 31 |
| 910 | Biomedical Sciences (University of New Hampshire) | 10 | 284 | 23 | 31 |
| 912 | Ecology (University of Vienna) | 8 | 283 | 59 | 81 |
| 912 | Genomics (James Cook University) | 5 | 283 | 25 | 26 |
| 914 | Evolutionary Biology (University of Oklahoma) | 6 | 282 | 17 | 25 |
| 914 | Computer Science (Florida State University) | 1 | 282 | 115 | 35 |
| 914 | Information Technology (Florida State University) | 1 | 282 | 65 | 23 |
| 917 | Genetics (University of Adelaide) | 2 | 279 | 89 | 50 |
| 917 | Biology (Technical University of Munich) | 3 | 279 | 41 | 42 |
| 917 | Education (Southwest University) | 8 | 279 | 234 | 200 |
| 917 | Animal Physiology (University of Tübingen) | 3 | 279 | 37 | 26 |
| 917 | Epidemiology (Université libre de Bruxelles) | 4 | 279 | 43 | 34 |
| 922 | Zoology (La Trobe University) | 5 | 278 | 61 | 51 |
| 923 | Molecular Biology (University of Texas at Austin) | 4 | 277 | 169 | 169 |
| 923 | Education (Marine Biological Laboratory) | 2 | 277 | 21 | 20 |
| 925 | Evolutionary Biology (University of Southampton) | 2 | 276 | 10 | 5 |
| 926 | Biotechnology (University of Wisconsin–Madison) | 3 | 275 | 43 | 45 |
| 927 | Biology (RWTH Aachen University) | 4 | 274 | 126 | 102 |
| 927 | Marine Biology (University of North Carolina Wilmington) | 5 | 274 | 73 | 47 |
| 927 | Zoology (University of Peshawar) | 1 | 274 | 52 | 5 |
| 927 | Biology (California State University, Northridge) | 10 | 274 | 29 | 48 |
| 931 | Oceanography (Massachusetts Institute of Technology) | 2 | 273 | 49 | 28 |
| 931 | Botany (University of Cologne) | 3 | 273 | 24 | 20 |
| 931 | Genomics (Princeton University) | 8 | 273 | 62 | 89 |
| 934 | Molecular Biology (Kiel University) | 8 | 272 | 59 | 88 |
| 935 | Anthropology (University of Auckland) | 3 | 271 | 38 | 11 |
| 935 | Biology (University of Minnesota Duluth) | 6 | 271 | 43 | 44 |
| 935 | Toxicology (Stockholm University) | 6 | 271 | 34 | 44 |
| 938 | Biology (National and Kapodistrian University of Athens) | 7 | 270 | 178 | 161 |
| 939 | Genetics (University of Helsinki) | 5 | 269 | 109 | 110 |
| 939 | Molecular Biology (Brown University) | 3 | 269 | 145 | 113 |
| 939 | Marine Biology (University of Vienna) | 1 | 269 | 50 | 18 |
| 939 | Biology (Providence College) | 3 | 269 | 27 | 19 |
| 943 | Biology (California State Polytechnic University, Humboldt) | 8 | 268 | 8 | 8 |
| 943 | Biology (College of William & Mary) | 5 | 268 | 72 | 63 |
| 945 | Microbiology (University of North Carolina at Chapel Hill) | 1 | 267 | 351 | 179 |
| 945 | Physiology (University of North Carolina at Chapel Hill) | 1 | 267 | 367 | 182 |
| 945 | Immunology (University of North Carolina at Chapel Hill) | 1 | 267 | 384 | 197 |
| 945 | Environmental Health (University of Guelph) | 1 | 267 | 2 | 2 |
| 945 | Evolutionary Genetics (University of Neuchâtel) | 9 | 267 | 6 | 14 |
| 945 | Environmental Health (Utah State University) | 1 | 267 | 2 | 2 |
| 951 | Conservation Biology (University of British Columbia) | 1 | 266 | 20 | 7 |
| 951 | Chemistry (Aalborg University) | 5 | 266 | 159 | 113 |
| 953 | Biology (Yamagata University) | 3 | 265 | 48 | 18 |
| 953 | Developmental Biology (University of Vienna) | 5 | 265 | 11 | 11 |
| 955 | Anatomy (University of Sydney) | 2 | 264 | 200 | 108 |
| 955 | Genetics (Vigo University) | 9 | 264 | 18 | 36 |
| 955 | History (University of Cologne) | 3 | 264 | 8 | 6 |
| 955 | Biochemistry (Vigo University) | 9 | 264 | 22 | 36 |
| 955 | Immunology (Vigo University) | 9 | 264 | 21 | 43 |
| 960 | Geology (McMaster University) | 1 | 262 | 166 | 74 |
| 961 | Human Genetics (University of Tübingen) | 5 | 261 | 27 | 21 |
| 961 | Genetics (Hebrew University of Jerusalem) | 7 | 261 | 124 | 157 |
| 961 | Marine Biology (University of Oslo) | 2 | 261 | 29 | 15 |
| 961 | Genetics (Washington State University) | 3 | 261 | 69 | 57 |
| 961 | Biology (University of Toledo) | 1 | 261 | 126 | 42 |
| 966 | Biology (Sonoma State University) | 4 | 260 | 49 | 39 |
| 966 | Ecology (Linnaeus University) | 7 | 260 | 28 | 39 |
| 968 | Evolutionary Biology (University of Freiburg) | 7 | 259 | 18 | 21 |
| 968 | Molecular Genetics (Duke University) | 5 | 259 | 122 | 142 |
| 968 | Education (Shanghai Ocean University) | 9 | 259 | 75 | 58 |
| 971 | Zoology (Washington State University) | 3 | 258 | 63 | 41 |
| 971 | Biology (University of Colorado) | 3 | 258 | 137 | 97 |
| 973 | Ecology (Marine Biological Laboratory) | 1 | 257 | 26 | 12 |
| 973 | Evolutionary Biology (Marine Biological Laboratory) | 1 | 257 | 10 | 8 |
| 975 | Ecology (University of Geneva) | 4 | 256 | 14 | 20 |
| 975 | Agricultural Biotechnology (Seoul National University) | 4 | 256 | 282 | 202 |
| 975 | Genetics (James Madison University) | 2 | 256 | 4 | 7 |
| 978 | Biotechnology (Swinburne University of Technology) | 1 | 255 | 187 | 58 |
| 978 | Chemistry (Swinburne University of Technology) | 1 | 255 | 153 | 51 |
| 980 | Botany (Stockholm University) | 2 | 253 | 114 | 91 |
| 981 | Biology (University of Alabama at Birmingham) | 6 | 252 | 213 | 221 |
| 982 | Ecology (Martin Luther University Halle-Wittenberg) | 7 | 251 | 12 | 27 |
| 982 | Evolutionary Biology (University of Vienna) | 6 | 251 | 26 | 32 |
| 984 | Biology (Vanderbilt University) | 3 | 250 | 318 | 243 |
| 984 | Ecology (Sun Yat-sen University) | 16 | 250 | 40 | 57 |
| 984 | Biology (Jagiellonian University) | 12 | 250 | 37 | 50 |
| 984 | Geology (Lund University) | 3 | 250 | 102 | 79 |
| 984 | Physiology (University of Kansas) | 2 | 250 | 264 | 151 |
| 989 | Immunology (University of Edinburgh) | 4 | 249 | 66 | 72 |
| 989 | Zoology (University of Copenhagen) | 2 | 249 | 58 | 27 |
| 989 | Biology (University of Paris) | 1 | 249 | 99 | 28 |
| 992 | Ecology (Stellenbosch University) | 5 | 248 | 100 | 82 |
| 993 | Anthropology (University of California, Los Angeles) | 2 | 247 | 124 | 82 |
| 994 | Oceanography (National Taiwan University) | 8 | 246 | 85 | 94 |
| 995 | Ecology (Kyoto University) | 7 | 245 | 40 | 51 |
| 995 | Pathology (University of Zaragoza) | 1 | 245 | 110 | 29 |
| 997 | Physiology (University of Würzburg) | 6 | 244 | 48 | 61 |
| 997 | Biology (University of Alberta) | 2 | 244 | 115 | 64 |
| 997 | Genomics (Harvard University) | 4 | 244 | 138 | 150 |
| 997 | Conservation Biology (University of Bern) | 4 | 244 | 37 | 36 |