| 1 | James Cook University | 501 | 26.5K | 9 | 12 |
| 2 | Australian Institute of Marine Science | 295 | 18.7K | 3 | 5 |
| 3 | Arc Centre of Excellence For Coral Reef Studies | 271 | 12.9K | 2 | 6 |
| 4 | Coral Reef Alliance | 285 | 12.6K | 13 | 15 |
| 5 | University of Miami | 135 | 9.4K | 115 | 83 |
| 6 | University of Queensland | 191 | 7.6K | 217 | 261 |
| 7 | Smithsonian Institution | 121 | 6.0K | 76 | 83 |
| 8 | University of Western Australia | 113 | 4.7K | 193 | 213 |
| 9 | University of Hawaiʻi at Mānoa | 97 | 3.9K | 77 | 85 |
| 10 | University of Newcastle | 27 | 3.4K | 419 | 169 |
| 11 | University of the Ryukyus | 89 | 3.3K | 27 | 20 |
| 12 | Newcastle University | 39 | 3.1K | 384 | 225 |
| 13 | University of Sydney | 63 | 2.5K | 534 | 525 |
| 14 | Woods Hole Oceanographic Institution | 46 | 2.4K | 64 | 73 |
| 15 | University of California, San Diego | 47 | 2.2K | 586 | 594 |
| 16 | University of Maine | 22 | 2.1K | 222 | 97 |
| 17 | Southern Cross University | 52 | 2.0K | 40 | 47 |
| 18 | Nova Southeastern University | 55 | 2.0K | 32 | 40 |
| 19 | King Abdullah University of Science and Technology | 62 | 2.0K | 94 | 137 |
| 20 | California State University | 57 | 1.9K | 477 | 445 |
| 21 | Scientific Centre of Monaco | 25 | 1.9K | 9 | 9 |
| 22 | Commonwealth Scientific and Industrial Research Organisation | 45 | 1.8K | 365 | 378 |
| 23 | University of California, Santa Barbara | 43 | 1.7K | 296 | 352 |
| 24 | University of Edinburgh | 24 | 1.7K | 803 | 531 |
| 25 | University of South Florida | 32 | 1.7K | 451 | 357 |
| 26 | University of New Hampshire | 14 | 1.7K | 349 | 158 |
| 27 | Tel Aviv University | 30 | 1.7K | 586 | 389 |
| 28 | University of Guam | 35 | 1.6K | 8 | 7 |
| 29 | Pennsylvania State University | 31 | 1.6K | 937 | 753 |
| 30 | University of Exeter | 31 | 1.5K | 417 | 354 |
| 31 | Stanford University | 23 | 1.5K | 1104 | 881 |
| 32 | National Autonomous University of Mexico | 41 | 1.5K | 453 | 337 |
| 33 | Florida Institute of Technology | 17 | 1.4K | 85 | 43 |
| 34 | University of Florida | 37 | 1.4K | 814 | 759 |
| 35 | Muséum national d'histoire naturelle | 25 | 1.4K | 155 | 136 |
| 36 | Caribbean Research and Management of Biodiversity Foundation | 25 | 1.4K | 1 | 3 |
| 37 | University of Auckland | 21 | 1.3K | 666 | 470 |
| 38 | Australian National University | 34 | 1.3K | 502 | 490 |
| 39 | University of Bremen | 33 | 1.3K | 199 | 186 |
| 40 | University of Tokyo | 32 | 1.3K | 788 | 667 |
| 41 | Curtin University | 44 | 1.3K | 279 | 330 |
| 42 | University of California, Los Angeles | 25 | 1.3K | 972 | 845 |
| 43 | Wildlife Conservation Society | 22 | 1.2K | 52 | 60 |
| 44 | University of Tasmania | 22 | 1.2K | 375 | 248 |
| 45 | National Center for Scientific Research | 17 | 1.2K | 744 | 436 |
| 46 | National University of Singapore | 48 | 1.2K | 539 | 658 |
| 47 | Victoria University of Wellington | 40 | 1.2K | 163 | 189 |
| 48 | Columbia University | 15 | 1.1K | 1172 | 858 |
| 49 | Florida International University | 25 | 1.1K | 340 | 277 |
| 50 | Louisiana State University | 22 | 1.1K | 623 | 502 |
| 51 | University of California, Santa Cruz | 19 | 1.1K | 322 | 292 |
| 52 | Academia Sinica | 36 | 1.1K | 341 | 324 |
| 53 | Naturalis Biodiversity Center | 44 | 1.1K | 32 | 49 |
| 54 | Manchester Metropolitan University | 14 | 1.0K | 295 | 140 |
| 55 | Georgia Tech | 23 | 1.0K | 532 | 526 |
| 56 | University of New South Wales | 23 | 1.0K | 912 | 760 |
| 57 | Bar-Ilan University | 30 | 1.0K | 264 | 270 |
| 57 | University of Technology Sydney | 43 | 1.0K | 251 | 340 |
| 59 | The Nature Conservancy | 20 | 992 | 58 | 62 |
| 60 | University of Amsterdam | 35 | 941 | 585 | 720 |
| 61 | University of the Virgin Islands | 21 | 930 | 4 | 8 |
| 62 | Oregon State University | 23 | 902 | 548 | 508 |
| 63 | University of California, Davis | 12 | 885 | 1163 | 856 |
| 64 | University of Melbourne | 24 | 869 | 968 | 884 |
| 65 | University of Delaware | 16 | 862 | 591 | 473 |
| 66 | Palau International Coral Reef Center | 16 | 806 | 1 | 1 |
| 67 | Dalhousie University | 17 | 804 | 611 | 503 |
| 68 | Royal Netherlands Institute of Sea Research | 21 | 800 | 54 | 71 |
| 69 | AIMS@JCU | 20 | 799 | 3 | 16 |
| 70 | University of Groningen | 15 | 783 | 873 | 722 |
| 71 | Spanish National Research Council | 28 | 761 | 562 | 601 |
| 72 | Macquarie University | 24 | 760 | 472 | 480 |
| 73 | Plymouth Marine Laboratory | 8 | 759 | 86 | 84 |
| 74 | Bermuda Institute of Ocean Sciences | 12 | 758 | 12 | 16 |
| 75 | University of Wollongong | 21 | 742 | 420 | 398 |
| 76 | University of Houston | 10 | 741 | 787 | 481 |
| 77 | New York University | 14 | 734 | 1009 | 804 |
| 78 | Boston University | 14 | 711 | 873 | 736 |
| 79 | University of Windsor | 6 | 690 | 561 | 203 |
| 80 | University of Montpellier | 5 | 689 | 780 | 326 |
| 81 | Bishop Museum | 11 | 688 | 21 | 11 |
| 82 | Princeton University | 8 | 684 | 868 | 612 |
| 83 | National Taiwan University | 26 | 682 | 594 | 611 |
| 84 | Lancaster Environment Centre | 14 | 674 | 111 | 127 |
| 85 | Institute for Biodiversity and Ecosystem Dynamics | 21 | 661 | 72 | 107 |
| 85 | University of South Alabama | 8 | 661 | 290 | 132 |
| 87 | University of São Paulo | 21 | 646 | 924 | 810 |
| 88 | University of Rhode Island | 21 | 641 | 295 | 334 |
| 89 | Federal University of Rio de Janeiro | 24 | 640 | 464 | 419 |
| 90 | University of California, Berkeley | 13 | 629 | 1207 | 1081 |
| 91 | University of Southampton | 14 | 626 | 858 | 741 |
| 92 | University of Panama | 2 | 610 | 119 | 13 |
| 93 | University of Hawaiʻi at Hilo | 13 | 605 | 38 | 33 |
| 94 | University of the West Indies | 15 | 595 | 207 | 127 |
| 95 | University of Cambridge | 15 | 590 | 1160 | 1058 |
| 96 | University of North Carolina at Chapel Hill | 12 | 589 | 860 | 767 |
| 97 | University of Adelaide | 10 | 588 | 950 | 714 |
| 98 | IFREMER | 23 | 576 | 143 | 180 |
| 99 | Ludwig-Maximilians-Universität München | 9 | 563 | 812 | 591 |
| 100 | University of North Carolina Wilmington | 22 | 553 | 83 | 102 |
| 101 | Stockholm University | 7 | 543 | 977 | 663 |
| 102 | Simon Fraser University | 9 | 536 | 803 | 551 |
| 103 | California Academy of Sciences | 18 | 533 | 41 | 41 |
| 104 | Simón Bolívar University | 10 | 528 | 112 | 46 |
| 105 | Hawaii Pacific University | 4 | 522 | 30 | 7 |
| 106 | Caron Treatment Centers | 21 | 520 | 10 | 24 |
| 107 | University of York | 9 | 517 | 754 | 575 |
| 108 | University of Texas at Austin | 15 | 515 | 1144 | 1095 |
| 108 | Griffith University | 19 | 515 | 569 | 652 |
| 110 | Tokyo Institute of Technology | 13 | 513 | 538 | 442 |
| 111 | University of Oxford | 19 | 504 | 1199 | 1247 |
| 112 | University of Essex | 11 | 499 | 424 | 348 |
| 113 | University of Hong Kong | 9 | 498 | 988 | 736 |
| 113 | McGill University | 13 | 498 | 1203 | 1076 |
| 113 | State University of New York | 16 | 498 | 1365 | 1224 |
| 116 | Duke University | 6 | 490 | 1468 | 1089 |
| 117 | Ohio State University | 11 | 479 | 1450 | 1208 |
| 118 | Wageningen University & Research | 12 | 469 | 614 | 636 |
| 119 | College of Charleston | 8 | 463 | 161 | 93 |
| 120 | Mote Marine Laboratory | 15 | 460 | 25 | 45 |
| 121 | Nanyang Technological University | 18 | 441 | 631 | 707 |
| 122 | Tohoku University | 8 | 440 | 952 | 680 |
| 123 | University of Vienna | 15 | 437 | 694 | 705 |
| 123 | Memorial University of Newfoundland | 8 | 437 | 617 | 416 |
| 125 | Hebrew University of Jerusalem | 13 | 431 | 936 | 864 |
| 126 | Helmholtz Association of German Research Centres | 12 | 429 | 882 | 822 |
| 127 | University of Konstanz | 11 | 415 | 380 | 402 |
| 127 | PSL Research University | 19 | 415 | 178 | 242 |
| 129 | Auburn University | 11 | 407 | 704 | 580 |
| 129 | University of Haifa | 13 | 407 | 510 | 450 |
| 131 | Florida Museum of Natural History | 9 | 400 | 84 | 75 |
| 132 | Silliman University | 10 | 395 | 3 | 11 |
| 133 | University of Toronto | 3 | 394 | 2183 | 1454 |
| 134 | Sea Education Association | 3 | 387 | 11 | 17 |
| 135 | National Sun Yat-sen University | 9 | 386 | 387 | 258 |
| 136 | Perry Institute for Marine Science | 2 | 383 | 6 | 5 |
| 137 | UCLouvain | 5 | 379 | 1053 | 742 |
| 137 | Japan Agency for Marine-Earth Science and Technology | 13 | 379 | 107 | 119 |
| 139 | University of British Columbia | 8 | 378 | 1543 | 1294 |
| 140 | Utrecht University | 8 | 375 | 1312 | 1146 |
| 141 | National Center for Ecological Analysis and Synthesis | 6 | 374 | 45 | 58 |
| 142 | University of St Andrews | 12 | 373 | 452 | 455 |
| 143 | Cornell University | 14 | 369 | 1327 | 1346 |
| 143 | University of Bologna | 13 | 369 | 941 | 910 |
| 145 | Western Washington University | 6 | 367 | 253 | 176 |
| 145 | University of Milan-Bicocca | 17 | 367 | 317 | 427 |
| 145 | University of Otago | 9 | 367 | 793 | 669 |
| 148 | Gordon and Betty Moore Foundation | 1 | 365 | 56 | 30 |
| 149 | Bangor University | 12 | 359 | 282 | 324 |
| 149 | Academy of Sciences of Uzbekistan | 13 | 359 | 399 | 399 |
| 151 | Leibniz Association | 16 | 358 | 624 | 717 |
| 151 | Florida State University | 10 | 358 | 805 | 728 |
| 153 | University of the South Pacific | 7 | 357 | 119 | 74 |
| 154 | Chinese Academy of Sciences | 19 | 356 | 1171 | 1215 |
| 155 | University of Mississippi | 13 | 350 | 463 | 476 |
| 155 | Nicholas School of the Environment | 4 | 350 | 194 | 161 |
| 157 | Lawrence Livermore National Laboratory | 4 | 348 | 470 | 328 |
| 158 | University of Victoria | 10 | 344 | 674 | 601 |
| 159 | Scottish Association for Marine Science | 4 | 338 | 90 | 76 |
| 160 | Universidade Federal de Santa Catarina | 13 | 333 | 469 | 398 |
| 161 | University of Iowa | 11 | 332 | 980 | 961 |
| 162 | Menlo College | 2 | 329 | 965 | 576 |
| 163 | Pfleger Institute of Environmental Research | 1 | 328 | 9 | 2 |
| 164 | University of Kansas | 5 | 322 | 1263 | 869 |
| 164 | Universidad de los Andes | 11 | 322 | 323 | 282 |
| 166 | University of South Carolina | 5 | 320 | 1152 | 823 |
| 167 | University of Barcelona | 13 | 310 | 803 | 843 |
| 168 | Leeward Community College | 3 | 301 | 3 | 5 |
| 169 | University of the Philippines | 7 | 295 | 193 | 113 |
| 169 | Council of Scientific and Industrial Research | 8 | 295 | 629 | 568 |
| 169 | Khaled bin Sultan Living Oceans Foundation | 6 | 295 | 4 | 7 |
| 172 | McMaster University | 4 | 294 | 1391 | 949 |
| 173 | University of Reading | 4 | 292 | 1049 | 674 |
| 173 | Universidade Federal de São Paulo | 5 | 292 | 632 | 371 |
| 175 | University of Aberdeen | 4 | 290 | 1101 | 739 |
| 176 | Kyushu University | 8 | 289 | 876 | 742 |
| 177 | Fluminense Federal University | 11 | 288 | 348 | 269 |
| 178 | University of Alabama at Birmingham | 5 | 285 | 894 | 671 |
| 179 | University of East Anglia | 6 | 284 | 842 | 676 |
| 179 | Japan Fisheries Research and Education Agency | 13 | 284 | 101 | 118 |
| 181 | Zayed University | 3 | 282 | 200 | 78 |
| 182 | Old Dominion University | 4 | 281 | 724 | 423 |
| 183 | University of Genoa | 11 | 276 | 605 | 663 |
| 184 | University of the Highlands and Islands | 6 | 273 | 49 | 46 |
| 185 | University of New Brunswick | 6 | 271 | 544 | 436 |
| 186 | University of Georgia | 12 | 269 | 167 | 247 |
| 187 | King Fahd University of Petroleum and Minerals | 3 | 268 | 515 | 292 |
| 188 | Kansas Geological Survey | 3 | 261 | 51 | 34 |
| 189 | La Trobe University | 4 | 256 | 1051 | 735 |
| 190 | National Aeronautics and Space Administration | 8 | 253 | 568 | 585 |
| 191 | Shizuoka University | 4 | 251 | 403 | 262 |
| 192 | Max Planck Society | 5 | 249 | 1040 | 899 |
| 193 | American University of Sharjah | 4 | 247 | 249 | 139 |
| 194 | CGIAR | 4 | 246 | 430 | 348 |
| 194 | Labex Corail | 14 | 246 | 5 | 17 |
| 196 | Deltares | 3 | 245 | 173 | 127 |
| 197 | Autonomous University of Barcelona | 5 | 244 | 1106 | 861 |
| 198 | Florida Atlantic University | 10 | 242 | 432 | 436 |
| 198 | National Dong Hwa University | 9 | 242 | 142 | 143 |
| 200 | Zoological Society of London | 7 | 241 | 92 | 114 |
| 201 | Federal University of Pernambuco | 10 | 239 | 469 | 375 |
| 202 | Michigan State University | 4 | 237 | 1737 | 1357 |
| 203 | Smith College | 5 | 233 | 235 | 172 |
| 203 | University of Plymouth | 7 | 233 | 529 | 531 |
| 205 | Vrije Universiteit Amsterdam | 3 | 231 | 1212 | 873 |
| 206 | Aix-Marseille University | 5 | 230 | 930 | 701 |
| 207 | University of Costa Rica | 7 | 229 | 240 | 228 |
| 208 | Oak Ridge National Laboratory | 2 | 227 | 778 | 531 |
| 209 | International Atomic Energy Agency | 5 | 224 | 159 | 150 |
| 210 | Pacific Biosciences | 6 | 222 | 68 | 78 |
| 211 | University of Colorado | 6 | 221 | 1529 | 1350 |
| 212 | University of Arkansas | 1 | 218 | 1560 | 708 |
| 212 | Hawai'i Department of Land and Natural Resources | 6 | 218 | 12 | 11 |
| 214 | Rice University | 6 | 217 | 712 | 649 |
| 215 | University of Copenhagen | 7 | 216 | 1331 | 1249 |
| 216 | Villanova University | 4 | 211 | 473 | 292 |
| 217 | University of Louisiana at Lafayette | 4 | 209 | 379 | 229 |
| 217 | Charles Darwin University | 6 | 209 | 271 | 236 |
| 217 | Autonomous University of Baja California Sur | 9 | 209 | 28 | 34 |
| 220 | Collège de France | 3 | 205 | 498 | 405 |
| 220 | University of New England | 8 | 205 | 461 | 496 |
| 220 | City University of New York | 4 | 205 | 1442 | 1064 |
| 220 | Northern Virginia Community College | 3 | 205 | 14 | 5 |
| 224 | Sorbonne University | 15 | 203 | 472 | 618 |
| 225 | Harvard University | 7 | 202 | 1711 | 1634 |
| 225 | Arizona State University | 5 | 202 | 1438 | 1257 |
| 225 | Shedd Aquarium | 9 | 202 | 4 | 11 |
| 228 | European Space Agency | 2 | 201 | 300 | 189 |
| 228 | University of Sheffield | 2 | 201 | 1779 | 1199 |
| 228 | Chinese University of Hong Kong | 10 | 201 | 931 | 995 |
| 231 | Rio de Janeiro State University | 4 | 200 | 576 | 305 |
| 232 | Pacific Science Center | 6 | 195 | 2 | 7 |
| 232 | Federal University of Espírito Santo | 9 | 195 | 284 | 240 |
| 234 | Universidade Federal da Paraíba | 9 | 193 | 332 | 258 |
| 235 | Alternative Energies and Atomic Energy Commission | 3 | 190 | 777 | 624 |
| 236 | Rio de Janeiro Botanical Garden | 5 | 188 | 58 | 49 |
| 236 | Hasanuddin University | 10 | 188 | 38 | 61 |
| 238 | University of Göttingen | 4 | 187 | 1011 | 859 |
| 238 | North Carolina State University | 5 | 187 | 1293 | 1173 |
| 240 | University of Warwick | 4 | 185 | 1231 | 1017 |
| 241 | University of Côte d'Azur | 1 | 184 | 677 | 304 |
| 242 | University College London | 5 | 180 | 1675 | 1485 |
| 242 | Waikiki Aquarium | 2 | 180 | 6 | 2 |
| 242 | Nature Conservation Foundation | 6 | 180 | 26 | 30 |
| 245 | Nature Seychelles | 1 | 179 | 37 | 17 |
| 246 | British Geological Survey | 2 | 178 | 237 | 174 |
| 246 | Laurentian University | 1 | 178 | 685 | 269 |
| 248 | El Colegio de la Frontera Sur | 8 | 176 | 100 | 118 |
| 249 | Arkansas State University | 7 | 175 | 103 | 121 |
| 249 | Syracuse University | 3 | 175 | 1426 | 1082 |
| 251 | Hokkaido University | 7 | 172 | 939 | 893 |
| 251 | De La Salle University | 4 | 172 | 165 | 98 |
| 251 | University of Derby | 6 | 172 | 200 | 201 |
| 254 | Okinawa Institute of Science and Technology | 8 | 170 | 133 | 159 |
| 255 | Texas A&M University | 6 | 168 | 1465 | 1405 |
| 255 | Coral Reef Research Foundation | 3 | 168 | 5 | 14 |
| 255 | Wageningen Marine Research | 2 | 168 | 67 | 48 |
| 258 | Nagoya University | 4 | 167 | 1062 | 874 |
| 258 | Fairleigh Dickinson University | 2 | 167 | 301 | 118 |
| 258 | Sultan Qaboos University | 6 | 167 | 418 | 403 |
| 261 | University of Zagreb | 5 | 166 | 860 | 710 |
| 262 | Ochanomizu University | 3 | 165 | 265 | 166 |
| 264 | University of Bristol | 4 | 163 | 1447 | 1215 |
| 264 | Carnegie Institution for Science | 6 | 163 | 214 | 273 |
| 264 | University of Mauritius | 2 | 163 | 228 | 126 |
| 267 | University of Manchester | 4 | 162 | 1658 | 1415 |
| 268 | Humboldt-Universität zu Berlin | 2 | 160 | 1284 | 863 |
| 268 | National Institute of Advanced Industrial Science and Technology | 6 | 160 | 147 | 137 |
| 268 | Sea Around Us | 1 | 160 | 39 | 31 |
| 271 | Guangxi University | 5 | 159 | 490 | 385 |
| 273 | Sorbonne Universités | 3 | 158 | 581 | 485 |
| 274 | Brown University | 3 | 156 | 1312 | 1026 |
| 274 | South Australian Research and Development Institute | 3 | 156 | 120 | 94 |
| 276 | University College Cork | 3 | 154 | 953 | 705 |
| 277 | Prince of Songkla University | 4 | 153 | 464 | 340 |
| 277 | Chesapeake Biological Laboratory | 3 | 153 | 91 | 104 |
| 277 | Universidad del Mar | 5 | 153 | 29 | 9 |
| 280 | University of Granada | 5 | 151 | 1067 | 918 |
| 280 | Academy of Sciences of the USSR | 2 | 151 | 448 | 242 |
| 282 | Clemson University | 9 | 149 | 689 | 843 |
| 282 | University of Miyazaki | 6 | 149 | 216 | 215 |
| 284 | National Geographic Society | 5 | 147 | 27 | 42 |
| 285 | University of Gothenburg | 5 | 146 | 1160 | 1099 |
| 286 | University of Algarve | 6 | 145 | 321 | 367 |
| 287 | Murdoch University | 5 | 144 | 595 | 592 |
| 288 | Statistics New Zealand | 3 | 140 | 1256 | 993 |
| 289 | Marine Conservation Institute | 1 | 139 | 14 | 17 |
| 289 | Senckenberg am Meer | 5 | 139 | 32 | 41 |
| 291 | Heriot-Watt University | 4 | 138 | 664 | 601 |
| 292 | Williams College | 4 | 137 | 249 | 253 |
| 292 | Kokushikan University | 2 | 137 | 76 | 20 |
| 292 | University of Girona | 3 | 137 | 566 | 448 |
| 292 | Diponegoro University | 5 | 137 | 136 | 85 |
| 296 | National Institute for Space Research | 2 | 134 | 182 | 116 |
| 297 | Yale University | 4 | 133 | 1585 | 1426 |
| 298 | Ewha Womans University | 4 | 132 | 649 | 535 |
| 298 | Russian Academy of Sciences | 3 | 132 | 667 | 470 |
| 300 | Ulster University | 1 | 131 | 1218 | 689 |
| 301 | Cornell College | 2 | 130 | 86 | 60 |
| 302 | University of Alabama | 10 | 128 | 770 | 1019 |
| 302 | University of Maryland | 1 | 128 | 1238 | 727 |
| 302 | Central Queensland University | 3 | 128 | 447 | 353 |
| 305 | University of Neuchâtel | 3 | 126 | 487 | 426 |
| 306 | Instituto de Estudos do Mar Almirante Paulo Moreira | 4 | 125 | 16 | 18 |
| 307 | University of Alberta | 4 | 123 | 1767 | 1588 |
| 307 | Bowdoin College | 3 | 123 | 210 | 180 |
| 309 | Georgia Southern University | 4 | 121 | 427 | 341 |
| 310 | University of La Laguna | 6 | 120 | 454 | 501 |
| 310 | Walla Walla University | 1 | 120 | 35 | 3 |
| 312 | Massachusetts Institute of Technology | 4 | 119 | 1321 | 1333 |
| 315 | Weizmann Institute of Science | 3 | 117 | 683 | 660 |
| 315 | University of Geneva | 4 | 117 | 1112 | 1050 |
| 317 | Cheng Shiu University | 4 | 116 | 98 | 103 |
| 318 | University of Guadalajara | 5 | 115 | 402 | 349 |
| 318 | RMIT University | 1 | 115 | 1381 | 896 |
| 320 | Uppsala University | 1 | 111 | 1944 | 1360 |
| 320 | University of the Western Cape | 1 | 111 | 811 | 424 |
| 323 | University of Aveiro | 6 | 110 | 537 | 630 |
| 324 | University of Warsaw | 6 | 109 | 712 | 694 |
| 325 | Eckerd College | 4 | 108 | 71 | 98 |
| 326 | University of Liège | 4 | 107 | 1004 | 1007 |
| 326 | University of Sherbrooke | 2 | 107 | 1071 | 783 |
| 329 | Sarah Lawrence College | 3 | 106 | 40 | 30 |
| 329 | University of Valle | 8 | 106 | 168 | 281 |
| 331 | University of Winnipeg | 2 | 105 | 445 | 289 |
| 331 | Reef Environmental Education Foundation | 4 | 105 | 3 | 8 |
| 335 | Washington University in St. Louis | 1 | 104 | 1893 | 1334 |
| 336 | Papua New Guinea University of Technology | 1 | 103 | 97 | 26 |
| 336 | University of North Florida | 3 | 103 | 382 | 243 |
| 338 | University of Palermo | 5 | 102 | 805 | 891 |
| 339 | University of Central Florida | 4 | 101 | 1035 | 989 |
| 340 | University of Calgary | 2 | 100 | 1778 | 1419 |
| 340 | Mongolian Academy of Sciences | 1 | 100 | 253 | 126 |
| 340 | Natural History Museum, Vienna | 5 | 100 | 64 | 77 |
| 343 | University of Glasgow | 5 | 99 | 1351 | 1368 |
| 343 | Rhodes University | 3 | 99 | 475 | 407 |
| 343 | Komazawa University | 2 | 99 | 33 | 21 |
| 343 | Tokyo Metropolitan University | 2 | 99 | 663 | 477 |
| 343 | University of Virginia | 4 | 99 | 1362 | 1335 |
| 348 | Université de Montréal | 2 | 98 | 1855 | 1511 |
| 349 | University of Waikato | 7 | 97 | 452 | 630 |
| 349 | Oberlin College | 3 | 97 | 239 | 208 |
| 349 | Federal Rural University of Rio de Janeiro | 1 | 97 | 510 | 227 |
| 349 | Field Museum of Natural History | 5 | 97 | 128 | 154 |
| 353 | Harvey Mudd College | 3 | 96 | 130 | 136 |
| 354 | University of Southern California | 6 | 95 | 1349 | 1492 |
| 354 | Ghent University | 3 | 95 | 1754 | 1587 |
| 354 | Defenders of Wildlife | 2 | 95 | 13 | 20 |
| 357 | Flinders University | 4 | 94 | 990 | 985 |
| 358 | University of Namur | 1 | 93 | 631 | 402 |
| 359 | University of California, Merced | 2 | 92 | 612 | 499 |
| 360 | Autonomous University of Baja California | 2 | 91 | 347 | 196 |
| 360 | Universidade Federal de Alagoas | 5 | 91 | 298 | 310 |
| 362 | Austral University of Chile | 4 | 90 | 349 | 331 |
| 362 | Western Sydney University | 1 | 90 | 1425 | 980 |
| 364 | University of Bonn | 1 | 89 | 1560 | 1144 |
| 364 | Purdue University | 2 | 89 | 2032 | 1692 |
| 364 | University of the Sunshine Coast | 3 | 89 | 490 | 427 |
| 367 | Meio University | 1 | 88 | 15 | 9 |
| 368 | Imperial College London | 5 | 87 | 1399 | 1513 |
| 368 | Yarmouk University | 2 | 87 | 381 | 221 |
| 368 | Tokyo University of Science | 2 | 87 | 660 | 500 |
| 368 | Nagoya Institute of Technology | 1 | 87 | 502 | 303 |
| 368 | Okinawa Prefectural Institute of Health and Environment | 2 | 87 | 31 | 18 |
| 368 | Koninklijke Philips NV | 3 | 87 | 498 | 485 |
| 374 | Polish Academy of Sciences | 2 | 85 | 1424 | 1140 |
| 374 | Tetra Tech | 2 | 85 | 104 | 96 |
| 374 | Sydney Institute of Marine Science | 6 | 85 | 39 | 82 |
| 374 | Moreton Bay Research Station | 1 | 85 | 13 | 22 |
| 378 | University of Canterbury | 1 | 84 | 1324 | 905 |
| 378 | University of Dar es Salaam | 2 | 84 | 407 | 278 |
| 380 | Université du Québec | 1 | 83 | 1757 | 1271 |
| 380 | Ben-Gurion University of the Negev | 6 | 83 | 894 | 1033 |
| 380 | Colorado State University | 2 | 83 | 1516 | 1290 |
| 380 | Tennenbaum Marine Observatories Network | 2 | 83 | 18 | 24 |
| 386 | University of Cincinnati | 3 | 82 | 1426 | 1316 |
| 386 | California Institute of Technology | 4 | 82 | 795 | 899 |
| 386 | University of Puerto Rico at Mayaguez | 7 | 82 | 41 | 109 |
| 389 | University of Duisburg-Essen | 5 | 81 | 666 | 843 |
| 389 | University of the Azores | 3 | 81 | 105 | 117 |
| 389 | Qatar University | 3 | 81 | 675 | 658 |
| 389 | King's College | 1 | 81 | 1113 | 770 |
| 393 | University of New Orleans | 3 | 80 | 485 | 469 |
| 393 | North-West University | 5 | 80 | 532 | 619 |
| 393 | European Commission | 2 | 80 | 725 | 684 |
| 396 | Tufts University | 1 | 79 | 1592 | 1181 |
| 396 | Kyoto University | 2 | 79 | 1601 | 1369 |
| 396 | Queensland University of Technology | 7 | 79 | 846 | 1204 |
| 396 | Hong Kong University of Science and Technology | 3 | 79 | 830 | 841 |
| 400 | University of Massachusetts Boston | 2 | 78 | 606 | 507 |
| 400 | Johns Hopkins University | 1 | 78 | 2028 | 1617 |
| 400 | University of Hull | 1 | 78 | 1244 | 820 |
| 400 | University of Mons | 3 | 78 | 250 | 270 |
| 405 | University of Leeds | 7 | 76 | 1242 | 1548 |
| 405 | Polytechnic University of the Marches | 5 | 76 | 501 | 678 |
| 405 | Guangdong Ocean University | 2 | 76 | 338 | 239 |
| 405 | Geological Survey of Israel | 1 | 76 | 143 | 99 |
| 409 | University of Michigan | 5 | 75 | 1917 | 1986 |
| 409 | European Molecular Biology Laboratory | 1 | 75 | 385 | 354 |
| 409 | Cooperative Institute for Research in the Atmosphere | 1 | 75 | 73 | 60 |
| 409 | Emerging Pathogens Institute | 1 | 75 | 205 | 176 |
| 409 | University of California, San Francisco | 1 | 75 | 1228 | 1026 |
| 414 | United States Environmental Protection Agency | 3 | 74 | 480 | 558 |
| 414 | Consiglio Nazionale delle Ricerche | 3 | 74 | 1264 | 1253 |
| 414 | Pontifical Xavierian University | 3 | 74 | 384 | 343 |
| 414 | Bigelow Laboratory for Ocean Sciences | 3 | 74 | 79 | 110 |
| 414 | Instituto Nacional de Cardiología | 1 | 74 | 241 | 152 |
| 419 | Islamic Azad University | 3 | 73 | 1207 | 1078 |
| 419 | Universidade Estadual da Paraíba | 1 | 73 | 288 | 137 |
| 421 | National Taiwan Normal University | 3 | 72 | 573 | 571 |
| 421 | Instituto de Ciencias Biológicas | 5 | 72 | 362 | 437 |
| 423 | Friedrich-Alexander-Universität Erlangen-Nürnberg | 4 | 71 | 933 | 1007 |
| 423 | University of Cape Town | 2 | 71 | 1358 | 1164 |
| 425 | Monash University | 4 | 70 | 1716 | 1765 |
| 425 | Laval University | 1 | 70 | 1793 | 1375 |
| 425 | Pontifical Catholic University of Chile | 2 | 70 | 1105 | 914 |
| 425 | Universiti Malaysia Sabah | 4 | 70 | 213 | 262 |
| 425 | DHI | 2 | 70 | 57 | 81 |
| 430 | IBM | 2 | 69 | 446 | 408 |
| 430 | Lund University | 1 | 69 | 1983 | 1604 |
| 430 | Universidad Veracruzana | 5 | 69 | 257 | 274 |
| 430 | Federal University of Rio Grande | 4 | 69 | 188 | 216 |
| 434 | Howard University | 1 | 68 | 976 | 606 |
| 434 | Washington and Lee University | 3 | 68 | 142 | 159 |
| 434 | Florida Fish and Wildlife Conservation Commission | 7 | 68 | 67 | 112 |
| 437 | University of Frankfurt | 4 | 67 | 186 | 349 |
| 438 | Murray State University | 1 | 66 | 364 | 207 |
| 438 | University of Asmara | 1 | 66 | 43 | 40 |
| 440 | George Washington University | 2 | 65 | 1510 | 1328 |
| 440 | Lawrence Berkeley National Laboratory | 2 | 65 | 747 | 753 |
| 440 | Seto Marine Biological Laboratory | 1 | 65 | 38 | 29 |
| 440 | Fisheries and Oceans Canada | 2 | 65 | 273 | 283 |
| 444 | George Mason University | 6 | 64 | 805 | 1094 |
| 445 | Cape Breton University | 1 | 62 | 243 | 135 |
| 445 | University of Massachusetts Dartmouth | 1 | 62 | 475 | 352 |
| 445 | Columbus Zoo and Aquarium | 2 | 62 | 6 | 10 |
| 445 | University of Illinois Urbana–Champaign | 2 | 62 | 1924 | 1738 |
| 449 | Los Alamos National Laboratory | 1 | 61 | 866 | 745 |
| 449 | Deakin University | 3 | 61 | 1229 | 1264 |
| 449 | Florida Institute of Oceanography | 1 | 61 | 21 | 20 |
| 452 | Senckenberg Research Institute | 1 | 60 | 76 | 71 |
| 452 | Edith Cowan University | 3 | 60 | 659 | 766 |
| 454 | Hiroshima University | 2 | 59 | 1132 | 1008 |
| 454 | Kanazawa University | 1 | 59 | 1024 | 775 |
| 454 | University of Siena | 2 | 59 | 888 | 821 |
| 454 | Catalan Institution for Research and Advanced Studies | 1 | 59 | 753 | 661 |
| 458 | Rivier University | 1 | 58 | 32 | 9 |
| 459 | Australian Defence Force Academy | 2 | 57 | 209 | 216 |
| 459 | Chulalongkorn University | 3 | 57 | 898 | 908 |
| 459 | King's Buildings | 2 | 57 | 342 | 398 |
| 459 | Israel Oceanographic and Limnological Research | 1 | 57 | 193 | 170 |
| 463 | University of Tübingen | 1 | 56 | 1440 | 1156 |
| 463 | RWTH Aachen University | 1 | 56 | 1451 | 1187 |
| 463 | Natural History Museum of Los Angeles County | 2 | 56 | 101 | 93 |
| 463 | Nancy-Université | 1 | 56 | 375 | 329 |
| 467 | University of Connecticut | 3 | 55 | 1579 | 1586 |
| 467 | University of Stuttgart | 1 | 55 | 995 | 822 |
| 467 | University of Richmond | 3 | 55 | 324 | 357 |
| 467 | Wetlands International | 1 | 55 | 53 | 58 |
| 467 | Tokyo University of Fisheries | 1 | 55 | 130 | 127 |
| 473 | Climate Change Research Centre | 1 | 54 | 104 | 95 |
| 474 | University of Hamburg | 2 | 53 | 1516 | 1401 |
| 474 | Saudi Aramco | 2 | 53 | 178 | 199 |
| 474 | University of Wisconsin–Madison | 1 | 53 | 2305 | 1969 |
| 474 | Massey University | 5 | 53 | 853 | 1101 |
| 474 | Tabriz University of Medical Sciences | 3 | 53 | 687 | 729 |
| 479 | University of Porto | 3 | 52 | 1026 | 1109 |
| 479 | Tajen University | 1 | 52 | 168 | 122 |
| 481 | University of Nottingham | 2 | 51 | 1889 | 1768 |
| 481 | Rutgers University | 4 | 51 | 1617 | 1736 |
| 481 | University of Nairobi | 1 | 51 | 719 | 504 |
| 481 | IHE Delft Institute for Water Education | 1 | 51 | 207 | 199 |
| 481 | Edinburgh Napier University | 1 | 51 | 581 | 423 |
| 481 | University of Gdańsk | 1 | 51 | 765 | 528 |
| 487 | UNESCO | 1 | 50 | 167 | 114 |
| 487 | Stazione Zoologica Anton Dohrn | 6 | 50 | 109 | 184 |
| 487 | University of Minnesota | 2 | 50 | 2115 | 1971 |
| 487 | Ruhr University Bochum | 2 | 50 | 1010 | 951 |
| 487 | Monterey Bay Aquarium Research Institute | 1 | 50 | 131 | 134 |
| 487 | I-Shou University | 1 | 50 | 593 | 437 |
| 487 | Trent University | 1 | 50 | 699 | 501 |
| 494 | University of Texas at Arlington | 1 | 49 | 1157 | 888 |
| 496 | Hohai University | 1 | 47 | 776 | 625 |
| 496 | University College Dublin | 1 | 47 | 1778 | 1495 |
| 496 | University of Seville | 3 | 47 | 1156 | 1179 |
| 496 | Federal University of Bahia | 1 | 47 | 742 | 504 |
| 500 | Universidade Federal do Paraná | 3 | 46 | 892 | 886 |
| 500 | Blue Ventures | 2 | 46 | 20 | 37 |
| 502 | Moi University | 1 | 45 | 313 | 240 |
| 502 | Helmholtz Institute for Functional Marine Biodiversity | 2 | 45 | 59 | 57 |
| 504 | Utah Valley University | 1 | 44 | 276 | 162 |
| 504 | Xiamen University | 4 | 44 | 898 | 1032 |
| 506 | Northumbria University | 2 | 43 | 890 | 865 |
| 506 | Center for Microbial Oceanography: Research and Education | 1 | 43 | 37 | 48 |
| 506 | University of Maryland, Baltimore County | 1 | 43 | 808 | 715 |
| 510 | University of Havana | 3 | 42 | 313 | 381 |
| 510 | Santa Monica College | 1 | 42 | 29 | 25 |
| 510 | Ramkhamhaeng University | 1 | 42 | 150 | 121 |
| 510 | University of Santiago | 1 | 42 | 311 | 258 |
| 510 | Bethel University | 1 | 42 | 110 | 82 |
| 510 | Savannah State University | 3 | 42 | 55 | 104 |
| 510 | Central Caribbean Marine Institute | 4 | 42 | 1 | 6 |
| 517 | American Museum of Natural History | 2 | 40 | 245 | 275 |
| 517 | University of Oldenburg | 1 | 40 | 632 | 548 |
| 517 | Cardiff University | 3 | 40 | 1415 | 1580 |
| 517 | Northern Kentucky University | 1 | 40 | 349 | 242 |
| 517 | Seychelles Islands Foundation | 2 | 40 | 15 | 16 |
| 522 | Washington State University | 2 | 39 | 1553 | 1503 |
| 522 | Charles Darwin Foundation | 1 | 39 | 80 | 78 |
| 522 | North Carolina Central University | 1 | 39 | 470 | 361 |
| 522 | National Council for Scientific and Technological Development | 1 | 39 | 254 | 217 |
| 522 | Ocean Institute | 1 | 39 | 9 | 8 |
| 527 | University of Fribourg | 3 | 38 | 781 | 930 |
| 527 | Northern Arizona University | 1 | 38 | 925 | 742 |
| 527 | University of Rostock | 3 | 38 | 753 | 947 |
| 527 | Drexel University | 1 | 38 | 1433 | 1270 |
| 527 | University of Silesia in Katowice | 1 | 38 | 777 | 547 |
| 527 | National Chung Hsing University | 1 | 38 | 879 | 766 |
| 527 | University of Tampa | 1 | 38 | 321 | 226 |
| 527 | Australian Nuclear Science and Technology Organisation | 1 | 38 | 373 | 363 |
| 527 | National University of Córdoba | 1 | 38 | 1205 | 960 |
| 527 | King Abdulaziz University | 1 | 38 | 1295 | 1143 |
| 527 | Cawthron Institute | 3 | 38 | 100 | 149 |
| 527 | Universidad Laica Eloy Alfaro de Manabí | 1 | 38 | 40 | 31 |
| 539 | Brock University | 1 | 37 | 952 | 787 |
| 539 | University of Waterloo | 2 | 37 | 1557 | 1549 |
| 539 | Stellenbosch University | 1 | 37 | 1465 | 1221 |
| 539 | Hainan University | 3 | 37 | 535 | 589 |
| 543 | Goethe University Frankfurt | 2 | 36 | 784 | 915 |
| 543 | University of North Texas | 2 | 36 | 1165 | 1218 |
| 543 | Pacific Community | 2 | 36 | 52 | 73 |
| 543 | University of New Mexico | 2 | 36 | 1474 | 1511 |
| 543 | University of Natal | 1 | 36 | 546 | 481 |
| 543 | Chamberlain University | 2 | 36 | 7 | 3 |
| 543 | Indian Ocean Commission | 1 | 36 | 5 | 4 |
| 550 | University of Innsbruck | 1 | 35 | 1122 | 980 |
| 550 | University of Wales | 1 | 35 | 1654 | 1471 |
| 550 | Universidade Federal do Ceará | 4 | 35 | 669 | 838 |
| 554 | Leipzig University | 3 | 34 | 1048 | 1237 |
| 554 | Swiss Federal Institute of Technology in Lausanne | 1 | 34 | 1089 | 1045 |
| 554 | University of Jordan | 1 | 34 | 649 | 558 |
| 554 | City University of Hong Kong | 2 | 34 | 1200 | 1227 |
| 554 | University of Trento | 2 | 34 | 942 | 1080 |
| 554 | University of Papua | 2 | 34 | 12 | 32 |
| 554 | Cirsa | 2 | 34 | 52 | 94 |
| 554 | National Taiwan Ocean University | 1 | 34 | 599 | 534 |
| 563 | West Virginia University | 2 | 33 | 1387 | 1398 |
| 563 | University of Oslo | 2 | 33 | 1769 | 1783 |
| 563 | Tallinn University of Technology | 1 | 33 | 670 | 604 |
| 563 | University of Graz | 3 | 33 | 641 | 868 |
| 563 | Technical University of Dortmund | 1 | 33 | 915 | 810 |
| 563 | TU Braunschweig | 1 | 33 | 935 | 837 |
| 563 | University of Oregon | 1 | 33 | 1365 | 1185 |
| 563 | Suez Canal University | 1 | 33 | 699 | 608 |
| 563 | British Antarctic Survey | 1 | 33 | 285 | 304 |
| 573 | Long Island University | 1 | 32 | 470 | 382 |
| 573 | University of Alicante | 2 | 32 | 807 | 871 |
| 573 | Manta Trust | 2 | 32 | 10 | 33 |
| 576 | Federal Center of Technological Education of Minas Gerais | 1 | 31 | 171 | 107 |
| 576 | University of Lausanne | 2 | 31 | 1225 | 1302 |
| 576 | Fairfield University | 1 | 31 | 364 | 286 |
| 576 | University of Campinas | 5 | 31 | 1112 | 1373 |
| 576 | China Geological Survey | 1 | 31 | 250 | 198 |
| 576 | Otterbein University | 2 | 31 | 62 | 81 |
| 576 | Universidad de Oriente | 1 | 31 | 248 | 189 |
| 576 | École Normale Supérieure | 1 | 31 | 787 | 757 |
| 584 | Shahid Beheshti University | 2 | 30 | 1174 | 1169 |
| 584 | Emory University | 1 | 30 | 1655 | 1484 |
| 584 | Middlebury College | 1 | 30 | 419 | 372 |
| 584 | University of North Carolina at Charlotte | 1 | 30 | 1284 | 1125 |
| 584 | Ocean Conservancy | 1 | 30 | 34 | 44 |
| 584 | Antarctic Climate and Ecosystems Cooperative Research Centre | 1 | 30 | 82 | 100 |
| 590 | ETH Zürich | 2 | 29 | 1552 | 1651 |
| 590 | Pompeu Fabra University | 1 | 29 | 1063 | 1006 |
| 590 | Heidelberg University | 1 | 29 | 1500 | 1366 |
| 590 | University of Basel | 2 | 29 | 1185 | 1312 |
| 590 | University of Arizona | 2 | 29 | 1909 | 1957 |
| 590 | University of Lisbon | 3 | 29 | 1385 | 1584 |
| 590 | Universiti Malaysia Terengganu | 3 | 29 | 221 | 326 |
| 590 | Bio21 Institute | 1 | 29 | 237 | 266 |
| 590 | San Diego Supercomputer Center | 1 | 29 | 123 | 137 |
| 600 | University of Zurich | 1 | 28 | 1766 | 1667 |
| 600 | Kagoshima University | 2 | 28 | 726 | 744 |
| 600 | University of Papua New Guinea | 3 | 28 | 94 | 156 |
| 600 | University of South Wales | 1 | 28 | 427 | 353 |
| 600 | Operation Wallacea | 1 | 28 | 32 | 27 |
| 600 | University of California, Irvine | 1 | 28 | 1886 | 1722 |
| 607 | Nagasaki University | 2 | 27 | 715 | 775 |
| 607 | Marquette University | 1 | 27 | 1016 | 893 |
| 607 | Universiti Brunei Darussalam | 2 | 27 | 311 | 414 |
| 607 | Mahidol University | 1 | 27 | 1232 | 1096 |
| 607 | Tokyo University of Marine Science and Technology | 2 | 27 | 271 | 310 |
| 607 | Burke Museum of Natural History and Culture | 2 | 27 | 87 | 124 |
| 613 | National University of Malaysia | 2 | 26 | 946 | 989 |
| 613 | Indiana University | 2 | 26 | 1764 | 1813 |
| 613 | Japan Science and Technology Agency | 1 | 26 | 688 | 696 |
| 616 | University of Koblenz - Landau | 1 | 25 | 354 | 372 |
| 616 | Kōchi University | 2 | 25 | 473 | 550 |
| 616 | National Polytechnic Institute | 4 | 25 | 832 | 1070 |
| 616 | Hofstra University | 1 | 25 | 816 | 700 |
| 616 | University of Southern Denmark | 1 | 25 | 1479 | 1386 |
| 616 | Beltsville Agricultural Research Center | 1 | 25 | 308 | 340 |
| 616 | Alagappa University | 1 | 25 | 340 | 324 |
| 616 | East–West Center | 2 | 25 | 121 | 176 |
| 616 | National Institute of Agrobiological Sciences | 1 | 25 | 280 | 289 |
| 616 | VIT University | 1 | 25 | 513 | 501 |
| 627 | University of the Aegean | 1 | 24 | 542 | 516 |
| 627 | Western Oregon University | 1 | 24 | 168 | 164 |
| 627 | Montana State University | 3 | 24 | 815 | 1114 |
| 627 | Universidade do Vale do Itajaí | 1 | 24 | 190 | 174 |
| 627 | Federal Rural University of Pernambuco | 1 | 24 | 534 | 450 |
| 627 | University of Texas Rio Grande Valley | 1 | 24 | 583 | 474 |
| 627 | Physiology and Biotechnology of Algae laboratory | 1 | 24 | 5 | 10 |
| 635 | University of Southern Mississippi | 1 | 23 | 931 | 874 |
| 635 | University of Murcia | 1 | 23 | 1160 | 1106 |
| 635 | University of Antioquia | 1 | 23 | 347 | 385 |
| 635 | Sōka University | 1 | 23 | 250 | 219 |
| 635 | Universidade do Estado de Santa Catarina | 1 | 23 | 503 | 424 |
| 635 | Gloucester Marine Genomics Institute | 1 | 23 | 9 | 15 |
| 641 | Kiel University | 3 | 22 | 1003 | 1293 |
| 641 | Radboud University | 1 | 22 | 1690 | 1615 |
| 641 | Universidade Federal do Pará | 1 | 22 | 836 | 715 |
| 644 | Norwegian University of Science and Technology | 3 | 21 | 1342 | 1659 |
| 644 | Australian Antarctic Division | 1 | 21 | 163 | 194 |
| 644 | Ohio University | 1 | 21 | 1275 | 1216 |
| 644 | Southern Connecticut State University | 1 | 21 | 317 | 288 |
| 644 | Udayana University | 1 | 21 | 218 | 206 |
| 644 | Teikyo University of Science | 1 | 21 | 148 | 150 |
| 644 | Pathwest Laboratory Medicine | 1 | 21 | 170 | 198 |
| 651 | Lomonosov Moscow State University | 2 | 20 | 892 | 977 |
| 651 | Wellesley College | 1 | 20 | 568 | 581 |
| 651 | Johannes Gutenberg University Mainz | 2 | 20 | 1106 | 1289 |
| 651 | University of Giessen | 2 | 20 | 711 | 929 |
| 651 | Bates College | 1 | 20 | 322 | 340 |
| 651 | University of North Dakota | 1 | 20 | 987 | 986 |
| 651 | Universidad Autónoma Metropolitana | 3 | 20 | 395 | 618 |
| 651 | Macalester College | 1 | 20 | 350 | 360 |
| 651 | Hatfield Marine Science Center | 2 | 20 | 108 | 169 |
| 651 | Roger Williams University | 3 | 20 | 64 | 174 |
| 651 | Center for NanoScience | 1 | 20 | 271 | 319 |
| 651 | Shimadzu Corp. | 1 | 20 | 186 | 215 |
| 663 | American Association for the Advancement of Science | 1 | 19 | 93 | 126 |
| 663 | Vrije Universiteit Brussel | 3 | 19 | 1070 | 1431 |
| 663 | University of Pisa | 2 | 19 | 1227 | 1424 |
| 663 | Tulane University of Louisiana | 1 | 19 | 1444 | 1360 |
| 663 | East Carolina University | 1 | 19 | 1344 | 1306 |
| 663 | Eastern Connecticut State University | 1 | 19 | 194 | 198 |
| 663 | Pontifical Catholic University of Valparaíso | 3 | 19 | 384 | 602 |
| 663 | University of Texas at El Paso | 1 | 19 | 1051 | 1014 |
| 663 | Adelphi University | 1 | 19 | 478 | 443 |
| 663 | Project Seahorse | 2 | 19 | 21 | 42 |
| 663 | National Institute of Fisheries Science | 1 | 19 | 182 | 191 |
| 675 | Zhejiang University | 1 | 18 | 1796 | 1720 |
| 675 | Université libre de Bruxelles | 3 | 18 | 1149 | 1526 |
| 675 | Tunghai University | 2 | 18 | 392 | 517 |
| 675 | Royal Roads University | 1 | 18 | 138 | 164 |
| 679 | ASICS | 3 | 17 | 1035 | 1307 |
| 679 | University of Alcalá | 1 | 17 | 1186 | 1149 |
| 679 | Temple University | 2 | 17 | 1364 | 1552 |
| 679 | King Juan Carlos University | 1 | 17 | 877 | 874 |
| 679 | California Institute for Telecommunications and Information Technology | 1 | 17 | 88 | 141 |
| 679 | Juniata College | 1 | 17 | 126 | 145 |
| 679 | Kuwait Institute for Scientific Research | 1 | 17 | 290 | 302 |
| 679 | Taronga Conservation Society | 1 | 17 | 52 | 56 |
| 679 | Iranian Fisheries Science Research Institute | 1 | 17 | 45 | 57 |
| 679 | Öko-Institut | 1 | 17 | 210 | 237 |
| 689 | Freie Universität Berlin | 1 | 16 | 1437 | 1469 |
| 689 | Technical University of Munich | 1 | 16 | 1668 | 1695 |
| 689 | Sapienza University of Rome | 2 | 16 | 1421 | 1640 |
| 689 | University of New Caledonia | 1 | 16 | 66 | 80 |
| 689 | University of Modena and Reggio Emilia | 1 | 16 | 1069 | 1146 |
| 689 | Universidad Michoacana de San Nicolás de Hidalgo | 1 | 16 | 493 | 478 |
| 689 | University of Crete | 1 | 16 | 959 | 1063 |
| 689 | Center for Genetic Engineering and Biotechnology | 1 | 16 | 352 | 398 |
| 689 | Natural History Museum of Denmark | 1 | 16 | 212 | 270 |
| 699 | Alten | 1 | 15 | 123 | 179 |
| 699 | University of Perugia | 1 | 15 | 1244 | 1296 |
| 699 | Masaryk University | 1 | 15 | 1249 | 1183 |
| 699 | Federal University of Rio Grande do Sul | 2 | 15 | 1355 | 1511 |
| 699 | Universidad de Málaga | 1 | 15 | 879 | 886 |
| 699 | Utah State University | 1 | 15 | 1348 | 1397 |
| 699 | Osaka City University | 1 | 15 | 847 | 870 |
| 699 | Roskilde University | 1 | 15 | 704 | 800 |
| 699 | Hainan Tropical Ocean University | 3 | 15 | 55 | 141 |
| 699 | Vietnam Academy of Science and Technology | 2 | 15 | 443 | 567 |
| 699 | Agropolis International | 1 | 15 | 286 | 361 |
| 699 | Future Earth | 1 | 15 | 88 | 105 |
| 711 | Boston College | 1 | 14 | 1166 | 1183 |
| 711 | Hungarian Academy of Sciences | 1 | 14 | 1105 | 1180 |
| 711 | Mount Allison University | 1 | 14 | 365 | 418 |
| 711 | Auckland University of Technology | 1 | 14 | 874 | 917 |
| 711 | Florida Southern College | 1 | 14 | 77 | 78 |
| 711 | Santa Barbara City College | 2 | 14 | 13 | 41 |
| 717 | Universidade Tecnológica Federal do Paraná | 1 | 13 | 591 | 577 |
| 717 | Addis Ababa University | 1 | 13 | 902 | 923 |
| 717 | University of Insubria | 1 | 13 | 806 | 889 |
| 717 | Facultad de Recursos Naturales | 1 | 13 | 121 | 165 |
| 717 | Universidade Federal do Vale do São Francisco | 1 | 13 | 276 | 262 |
| 722 | University of Bergen | 1 | 12 | 1597 | 1632 |
| 722 | Northwestern University | 1 | 12 | 1970 | 1978 |
| 722 | University of Düsseldorf | 1 | 12 | 1152 | 1224 |
| 722 | Burapha University | 1 | 12 | 328 | 408 |
| 722 | Xi'an Jiaotong University | 2 | 12 | 1267 | 1479 |
| 722 | Carleton University | 1 | 12 | 1538 | 1547 |
| 722 | University of Kentucky | 1 | 12 | 1917 | 1911 |
| 722 | College of the Atlantic | 1 | 12 | 55 | 86 |
| 722 | Wenzhou University | 2 | 12 | 472 | 623 |
| 731 | University of Galway | 1 | 11 | 671 | 708 |
| 731 | College of William & Mary | 1 | 11 | 939 | 1037 |
| 731 | São Paulo State University | 1 | 11 | 1421 | 1410 |
| 731 | Istituto superiore per la protezione e la ricerca ambientale | 1 | 11 | 179 | 228 |
| 731 | National Chiayi University | 1 | 11 | 483 | 580 |
| 731 | Virginia Institute of Marine Science | 1 | 11 | 209 | 256 |
| 731 | MingDao University | 1 | 11 | 122 | 158 |
| 731 | Istituto Zooprofilattico Sperimentale del Mezzogiorno | 1 | 11 | 57 | 89 |
| 739 | University of Cologne | 1 | 10 | 1519 | 1597 |
| 739 | Virginia Tech | 1 | 10 | 1905 | 1963 |
| 739 | Leibniz University Hannover | 1 | 10 | 965 | 1086 |
| 739 | University of Guelph | 2 | 10 | 1338 | 1660 |
| 739 | Kindai University | 1 | 10 | 845 | 913 |
| 739 | University of Cyprus | 1 | 10 | 831 | 966 |
| 739 | Montclair State University | 1 | 10 | 756 | 823 |
| 739 | Sam Ratulangi University | 1 | 10 | 104 | 148 |
| 739 | St. George's University | 2 | 10 | 228 | 411 |
| 739 | The Cyprus Institute | 1 | 10 | 143 | 208 |
| 739 | Woosuk University | 1 | 10 | 196 | 227 |
| 739 | ELETTRA | 1 | 10 | 209 | 280 |
| 739 | Austrian Federal Railways | 1 | 10 | 993 | 1164 |
| 754 | Anna University | 2 | 9 | 663 | 900 |
| 754 | Mokpo National University | 1 | 9 | 300 | 389 |
| 754 | Shanghai Jiao Tong University | 1 | 9 | 1768 | 1819 |
| 754 | University of Oklahoma | 2 | 9 | 1441 | 1760 |
| 754 | Ocean University of China | 2 | 9 | 749 | 981 |
| 754 | North Carolina Agricultural and Technical State University | 1 | 9 | 519 | 664 |
| 754 | Indian Institute of Tropical Meteorology | 2 | 9 | 93 | 147 |
| 754 | WISERD | 1 | 9 | 201 | 295 |
| 763 | Tokai University | 1 | 8 | 959 | 1061 |
| 763 | Ryukoku University | 1 | 8 | 333 | 401 |
| 763 | University of Nebraska–Lincoln | 1 | 8 | 1852 | 1967 |
| 763 | Alexandria University | 1 | 8 | 1043 | 1158 |
| 763 | Graduate University for Advanced Studies | 1 | 8 | 378 | 479 |
| 763 | International Council for the Exploration of the Sea | 2 | 8 | 17 | 42 |
| 769 | University of Bordeaux | 1 | 7 | 1155 | 1323 |
| 769 | Rensselaer Polytechnic Institute | 1 | 7 | 995 | 1152 |
| 769 | Sun Yat-sen University | 1 | 7 | 1679 | 1760 |
| 769 | Çanakkale Onsekiz Mart University | 1 | 7 | 476 | 600 |
| 769 | Arcadis | 1 | 7 | 125 | 192 |
| 769 | University of Tehran | 1 | 7 | 1727 | 1801 |
| 769 | University of Chester | 1 | 7 | 427 | 583 |
| 769 | Autonomous University of Madrid | 1 | 7 | 1525 | 1661 |
| 769 | University of Las Palmas de Gran Canaria | 1 | 7 | 458 | 620 |
| 769 | University of Salford | 1 | 7 | 1029 | 1180 |
| 769 | Australian Institute of International Affairs | 1 | 7 | 790 | 949 |
| 769 | Bicol University | 1 | 7 | 17 | 26 |
| 781 | Leiden University | 2 | 6 | 1544 | 1897 |
| 781 | Adam Mickiewicz University in Poznań | 1 | 6 | 574 | 661 |
| 781 | Haverford College | 1 | 6 | 280 | 408 |
| 781 | University of Tsukuba | 2 | 6 | 1306 | 1639 |
| 781 | Oklahoma State University | 1 | 6 | 1500 | 1686 |
| 781 | Case Western Reserve University | 1 | 6 | 1608 | 1726 |
| 781 | Alexandru Ioan Cuza University | 1 | 6 | 501 | 646 |
| 781 | Middle Tennessee State University | 1 | 6 | 576 | 736 |
| 781 | pavillon Charles-Eugène-Marchand | 1 | 6 | 52 | 114 |
| 781 | Centre for Alternative Technology | 1 | 6 | 9 | 24 |
| 781 | Inbios | 1 | 6 | 72 | 127 |
| 781 | Graduate University of Chinese Academy of Sciences | 1 | 6 | 538 | 702 |
| 781 | Centro de Investigaciones de Ecosistemas Costeros | 1 | 6 | 5 | 12 |
| 781 | Indian Institute of Science Education and Research, Bhopal | 1 | 6 | 150 | 211 |
| 795 | University of Strasbourg | 1 | 5 | 1207 | 1408 |
| 795 | Niigata University | 1 | 5 | 884 | 1066 |
| 795 | Concordia University | 1 | 5 | 1414 | 1622 |
| 795 | Ehime University | 1 | 5 | 821 | 976 |
| 795 | Western University | 1 | 5 | 1852 | 2023 |
| 795 | University of Saskatchewan | 1 | 5 | 1668 | 1838 |
| 795 | National Penghu University of Science and Technology | 1 | 5 | 110 | 175 |
| 795 | Institut Louis-Malardé | 1 | 5 | 59 | 95 |
| 795 | Bloomsbury Publishing | 1 | 5 | 243 | 437 |
| 795 | Microbial Ecology | 2 | 5 | 320 | 543 |
| 795 | Harte Research Institute for Gulf of Mexico Studies | 1 | 5 | 42 | 74 |
| 807 | Tsinghua University | 1 | 4 | 1663 | 1811 |
| 807 | Azerbaijan National Academy of Sciences | 1 | 4 | 469 | 579 |
| 807 | University of Huelva | 1 | 4 | 561 | 743 |
| 807 | Hong Kong Baptist University | 1 | 4 | 1096 | 1321 |
| 807 | Vigo University | 1 | 4 | 978 | 1193 |
| 807 | Hong Kong Polytechnic University | 1 | 4 | 1491 | 1679 |
| 807 | Catholic University of the Most Holy Conception | 1 | 4 | 292 | 408 |
| 807 | University of Trieste | 1 | 4 | 1226 | 1456 |
| 807 | Federal University of the State of Rio de Janeiro | 1 | 4 | 371 | 485 |
| 807 | Federal University of ABC | 1 | 4 | 702 | 873 |
| 807 | Linfield University | 1 | 4 | 115 | 238 |
| 807 | Research Institute for Humanity and Nature | 1 | 4 | 180 | 312 |
| 807 | Skidaway Institute of Oceanography | 1 | 4 | 101 | 143 |
| 807 | Fundación La Salle de Ciencias Naturales | 1 | 4 | 33 | 61 |
| 822 | Tata Institute of Fundamental Research | 1 | 3 | 464 | 623 |
| 822 | Ateneo de Manila University | 1 | 3 | 301 | 474 |
| 822 | University of Trinidad and Tobago | 1 | 3 | 52 | 143 |
| 822 | Halliburton | 1 | 3 | 102 | 183 |
| 822 | Kaohsiung Medical University | 1 | 3 | 768 | 926 |
| 822 | Cabrillo College | 1 | 3 | 23 | 57 |
| 822 | Environmental Defense Fund | 1 | 3 | 123 | 180 |
| 822 | University of Ostrava | 1 | 3 | 328 | 496 |
| 822 | Joint Genome Institute | 1 | 3 | 155 | 219 |
| 822 | Sokoine University of Agriculture | 1 | 3 | 322 | 443 |
| 822 | Federal University of Viçosa | 1 | 3 | 775 | 924 |
| 822 | Sathyabama University | 1 | 3 | 164 | 273 |
| 822 | Tanjungpura University | 1 | 3 | 100 | 151 |
| 822 | Catholic University of Salvador | 1 | 3 | 59 | 78 |
| 822 | Universidade Federal de Goiás | 1 | 3 | 894 | 1084 |
| 822 | Western Australian Herbarium | 1 | 3 | 35 | 61 |
| 822 | University of Ruhuna | 1 | 3 | 279 | 447 |
| 822 | ARC Centre of Excellence in Synthetic Biology | 1 | 3 | 54 | 100 |
| 822 | Museum of Vertebrate Zoology | 1 | 3 | 133 | 176 |
| 841 | University of San Diego | 1 | 2 | 553 | 783 |
| 841 | Biocon | 1 | 2 | 96 | 193 |
| 841 | United States Department of Agriculture | 1 | 2 | 1028 | 1217 |
| 841 | Aarhus University | 1 | 2 | 1806 | 2036 |
| 841 | Okayama University | 1 | 2 | 1045 | 1293 |
| 841 | Golder Associates | 1 | 2 | 143 | 261 |
| 841 | Kean University | 1 | 2 | 290 | 456 |
| 841 | Japan International Research Center for Agricultural Sciences | 1 | 2 | 212 | 309 |
| 850 | Tongji University | 1 | 1 | 1387 | 1664 |
| 850 | University of Naples Federico II | 1 | 1 | 1378 | 1694 |
| 850 | Nihon University | 1 | 1 | 1000 | 1278 |
| 850 | United States Naval Research Laboratory | 1 | 1 | 633 | 808 |
| 850 | Kirkwood Community College | 1 | 1 | 10 | 26 |
| 850 | Murdoch Children's Research Institute | 1 | 1 | 532 | 695 |
| 850 | Royal Holloway, University of London | 1 | 1 | 968 | 1264 |
| 857 | University of Antwerp | 1 | 0 | 1619 | 1959 |
| 857 | Jagiellonian University | 1 | 0 | 1392 | 1698 |
| 857 | Ibaraki University | 1 | 0 | 587 | 787 |
| 857 | National Academy of Sciences of Armenia | 1 | 0 | 405 | 556 |
| 857 | Central Research Institute of Electric Power Industry | 1 | 0 | 363 | 533 |